Chapter 41

During the formation or improvement of a breed, its members will always be found to vary much in those characters to which especial attention is directed, and of which each slight improvement is eagerly sought and selected. Thus, with short-faced tumbler-pigeons, the shortness of the beak, shape of head and plumage,--with carriers, the length of the beak and wattle,--with fantails, the tail and carriage,--with Spanish fowls, the white face and comb,--with long-eared rabbits, the length of ear, are all points which are eminently variable. So it is in every case; and the large price paid for first-rate animals proves the difficulty of breeding them up to the highest standard of excellence. This subject has been discussed by fanciers (21/55. 'Cottage Gardener' 1855 December page 171; 1856 January pages 248, 323.), and the greater prizes given for highly improved breeds, in comparison with those given for old breeds which are not now undergoing rapid improvement, have been fully justified. Nathusius makes (21/56. 'Ueber Shorthorn Rindvieh' 1857 s. 51.) a similar remark when discussing the less uniform character of improved Shorthorn cattle and of the English horse, in comparison, for example, with the unennobled cattle of Hungary, or with the horses of the Asiatic steppes. This want of uniformity in the parts which at the time are undergoing selection chiefly depends on the strength of the principle of reversion; but it likewise depends to a certain extent on the continued variability of the parts which have recently varied. That the same parts do continue varying in the same manner we must admit, for if it were not so, there could be no improvement beyond an early standard of excellence, and we know that such improvement is not only possible, but is of general occurrence.

As a consequence of continued variability, and more especially of reversion, all highly improved races, if neglected or not subjected to incessant selection, soon degenerate. Youatt gives a curious instance of this in some cattle formerly kept in Glamorganshire; but in this case the cattle were not fed with sufficient care. Mr. Baker, in his memoir on the Horse, sums up: "It must have been observed in the preceding pages that, whenever there has been neglect, the breed has proportionally deteriorated." (21/57. 'The Veterinary'