The Manual of the Botany of the Northern United States Including the District East of the Mississippi and North of North Carolina and Tennessee

Part 33

Chapter 333,244 wordsPublic domain

Heads small, mostly clustered in the axils of feather-veined leaves 3--7

Heads mostly large, in a terminal thyrse; leaves feather-veined.

Western species 8, 9

Northern or mountain species 10--12

Heads mostly small or middle-sized; inflorescence paniculate (sometimes thyrsoidal).

Leaves 3-ribbed; heads in 1-sided spreading panicled racemes.

Stem and leaves smooth and glabrous 29--32

Pubescent or scabrous 33--36

Leaves not 3-ribbed, or only obscurely triple-nerved.

Heads large; leaves thickish, very smooth, entire. Seashore 13

Panicle virgate or thyrsoid; leaves nearly entire 14--17

Heads very small in a short broad panicle; leaves nearly entire 18--20

Heads racemosely paniculate; leaves ample, the lower serrate 21--28

Sec. 1. VIRGAUREA. _Rays mostly fewer than the disk-flowers; heads all more or less pedicelled._

[*] _Scales of the much imbricated and rigid involucre with abruptly spreading herbaceous tips; heads in clusters or glomerate racemes, disposed in a dense somewhat leafy and interrupted wand-like compound spike._

1. S. squarrosa, Muhl. Stem stout (2--5 deg. high), hairy above; leaves large, oblong, or the lower spatulate-oval and tapering into a margined petiole, serrate, veiny; heads numerous; scales obtuse or acute; disk-flowers 16--24, the rays 12--16.--Rocky and wooded hills, Maine and W. Vt. to Penn., Ohio, and the mountains of Va.; rather rare.

2. S. petiolaris, Ait. Minutely hoary or downy; stem strict, simple (1--3 deg. high); leaves small (1/2--2' long), oval or oblong, mucronate, veiny, rough-ciliolate; the upper entire and abruptly very short-petioled, the lower often serrate and tapering to the base; heads few, in a wand-like raceme or panicle, on slender bracted pedicels; rays about 10, elongated; scales of the pubescent involucre lanceolate or linear-awl-shaped, the outer loose and spreading, more or less foliaceous.--S. W. Ill. to Kan. and southward.--The name is misleading, as the leaves are hardly petioled.

[*][*] _Involucral scales without green tips and wholly appressed._

[+] _Heads small (3'' long), clustered along the stem in the axils of the feather-veined leaves, or the upper forming a thyrse._

[++] _Achenes pubescent._

3. S. caesia, L. Smooth; _stem terete, mostly glaucous_, at length much branched and diffuse; _leaves lanceolate_ or oblong-lanceolate, serrate, pointed, sessile; heads in very short clusters, or somewhat racemose-panicled on the branches.--Rich woodlands, common; west to S. E. Minn., Ill., and Ky.

4. S. latifolia, L. Smooth or nearly so; _stem angled, zigzag_, simple or paniculate-branched (1--3 deg. high); _leaves broadly ovate or oval, very strongly and sharply serrate, conspicuously pointed at both ends_ (thin, 3--6' long); heads in very short axillary clusters, or the clusters somewhat prolonged at the end of the branches; rays 3--4.--Moist shaded banks; common northward, and south along the mountains.

5. S. Curtisii, Torr. & Gray. Smooth or nearly so; stem angled, usually branched; leaves oblong to long-lanceolate with narrowed entire base, serrate above with subulate teeth; heads in small, loose clusters; rays 4--7.--Open woods at low elevations in the mountains of Va. and southward.

[++][++] _Achenes glabrous; inflorescence more thyrsoid._

6. S. bicolor, L. _Hoary or grayish with soft hairs_; stem mostly simple; leaves oblong or elliptical-lanceolate, acute at both ends, or the lower oval and tapering into a petiole, slightly serrate; _clusters or short racemes from the axils of the upper leaves_, forming an interrupted spike or crowded panicle; scales very obtuse; _rays (5--14) small, cream-color or nearly white_.--Var. CONCOLOR, Torr. & Gray, has the _rays yellow_.--Dry copses, west to Minn. and Mo.

7. S. monticola, Torr. & Gray. Nearly glabrous; stem slender, 1--2 deg. high; leaves oblong-ovate to lanceolate, acute or tapering at both ends, the lower sparingly serrate; heads small, the scales acutish; rays 5--6.--Alleghany Mts., from Md. southward.

[+][+] _Heads mostly large (smaller in n. 12), many-flowered, forming an erect terminal thyrse; leaves feather-veined._

[++] _Leaves numerous, short, sessile, entire, uniform in size and shape; western._

8. S. Bigelovii, Gray. Cinereous-puberulent, 2 deg. high; leaves oval and oblong, mostly obtuse at both ends; thyrse rather loose; involucre broad.--S. Kan. and southward. Probably running into the next.

9. S. Lindheimeriana, Scheele. Less puberulent; leaves lanceolate or oblong, more acute; heads narrower and more densely clustered; achenes glabrous.--S. Kan. and southward.

[++][++] _Northern or mountain species, bright green._

10. S. macrophylla, Pursh. _Stem stout_ (1--4 deg. high), _wand-like_, pubescent near the summit, simple; _leaves thin, ovate, irregularly and coarsely serrate with sharp salient teeth, large_ (lower 3--4' long), all but the uppermost abruptly contracted into _long and margined petioles; heads large_ (5--6'' long), many-flowered, crowded in an oblong or wand-like raceme or contracted panicle (2--18' long); scales loose and thin, long, lanceolate, taper-pointed; rays 8--10, elongated; achenes smooth. (S. thyrsoidea, _E. Mey_.)--Wooded sides of mountains, N. Maine to N. Y. (south to the Catskills), shore of L. Superior and northward.--Very near a European form of S. Virgaurea.

11. S. Virgaurea, Linn. An extremely variable and confused species in the Old World, represented in North America by

Var. alpina, Bigel. Dwarf (1--8' high), with few (1--12) pretty large heads (3--4'' long, becoming smaller as they increase in number); leaves thickish, mostly smooth, spatulate or obovate, mostly obtuse, finely serrate or nearly entire, the uppermost lanceolate; heads few in a terminal cluster or subsolitary in the upper axils; _scales_ lanceolate, _acute or acutish_; rays about 12.--Alpine summits of Maine, N. H., and N. Y., and shore of L. Superior.

12. S. humilis, Pursh. Low (6--12' high) and smooth, bearing several or numerous loosely thyrsoid smaller heads, which, with the peduncles, etc., are mostly somewhat glutinous; _scales obtuse_; rays 6--8, short; upper leaves lanceolate to linear, entire, the lower becoming spatulate and sparingly serrate. (S. Virgaurea, var. humilis, _Gray_.)--Rocky banks, W. Vt., along the Great Lakes, and northward; also on islands in the Susquehanna, near Lancaster, and at the Falls of the Potomac.--At the base of the White Mountains, on gravelly banks, occurs a form with the minutely pubescent stout stem 1--2 deg. high, the leaves larger, broader, and coarsely toothed, and the heads very numerous in an ample compound raceme; rays occasionally almost white.

Var. Gillmani, Gray. Larger (2 deg. high), rigid, with compound ample panicle and laciniately toothed leaves.--Sand-hills of the lake-shores, N. Mich.

[+][+][+] _Heads small or middle-sized (large in n. 13 and 17), panicled or sometimes thyrsoidal, not in a terminal corymbiform cyme; not alpine._

[++] _Leaves veiny, not 3-ribbed, but sometimes obscurely triple-nerved._

[=] 1. _Heads commonly large; leaves thickish, very smooth, entire, elongated._

13. S. sempervirens, L. Smooth and stout (1--8 deg. high); leaves lanceolate, slightly clasping, or the lower ones lanceolate-oblong, obscurely triple-nerved; racemes short, in an open or contracted panicle.--Salt marshes, or rocks on the shore, Maine to Va.--Heads showy; the golden rays 7--10. Varies, in less brackish swamps, with thinner elongated linear-lanceolate leaves, tapering to each end, and more erect racemes in a narrower panicle.

[=] 2. _Heads small, in a narrow virgate or thyrsoid panicle; scales thin, acute; leaves nearly entire._

14. S. stricta, Ait. _Very smooth_ throughout; _stem strict and simple, wand-like_ (2--4 deg. high), slender, beset with small and entire appressed lanceolate-oblong thickish leaves, gradually reduced upward to mere bracts, the lowest oblong-spatulate; _heads crowded in a very narrow compound spicate raceme_; rays 5--7. (S. virgata, _Michx._)--Damp pine barrens, N. J. and southward.

15. S. puberula, Nutt. _Stem_ (1--3 deg. high, simple or branched) _and panicle minutely hoary; stem-leaves lanceolate, acute, tapering to the base_, smoothish; the lower wedge-lanceolate and _sparingly toothed, heads_ very numerous and _crowded in compact short racemes forming a prolonged and dense_ narrow or pyramidal _panicle; scales linear-awl-shaped_, appressed; rays about 10.--Sandy soil, Maine to Va. and southward, mostly near the coast.

[=] 3. _Heads middle-sized, in a thyrsoid panicle; involucral scales rather firm, obtuse; leaves entire or little serrate, smooth._

16. S. uliginosa, Nutt. _Smooth_ nearly throughout; stem simple, strict (2--3 deg. high); _leaves lanceolate_, pointed, the lower tapering into winged petioles, partly sheathing at the base, sparsely serrulate or entire; _racemes much crowded and appressed in a dense wand-like panicle_; scales linear-oblong; rays 5--6, small. (S. stricta, _Ait._)--Peat-bogs, Maine to Penn., Minn., and northward. Root-leaves 6--10' long. Flowers earlier than most species, beginning in July.

17. S. speciosa, Nutt. Stem stout (3--6 deg. high), smooth; _leaves thickish, smooth_ with rough margins, _oval or ovate_, slightly serrate, the uppermost oblong-lanceolate, the lower contracted into a margined petiole; _heads_ somewhat crowded _in numerous erect racemes, forming an ample pyramidal or thyrsiform panicle_; peduncles and pedicels rough-hairy; scales of the cylindrical involucre oblong; rays about 5, large.--Var. ANGUSTATA, Torr. & Gray, is a dwarf form, with the racemes short and clustered, forming a dense interrupted or compound spike.--Copses, Maine to Minn., and southward.--A very handsome species; the lower leaves 4--6' long and 2--4' wide in the larger forms.

[=] 4. _Heads very small in slender spreading secund clusters forming a mostly short and broad panicle; leaves entire or nearly so._

18. S. odora, Ait. (SWEET GOLDEN-ROD.) _Smooth_ or nearly so throughout; _stem slender_ (2--3 deg. high), _often reclined; leaves linear-lanceolate, entire_, shining, _pellucid-dotted_; racemes spreading in a small one-sided panicle; rays 3--4, rather large.--Border of thickets in dry or sandy soil, Maine and Vt. to Ky., and southward.--The crushed leaves yield a pleasant anisate odor; but an occasional form is nearly scentless.

19. S. tortifolia, Ell. Stem scabrous-puberulent, 2--3 deg. high; leaves linear, short, commonly twisted, roughish-puberulent or glabrate; rays very short.--Dry soil, coast of Va. and southward.

20. S. pilosa, Walt. _Stem stout, upright_ (3--7 deg. high), _clothed with spreading hairs; leaves oblong-lanceolate, roughish, hairy beneath_, at least on the midrib, serrulate, the upper ovate-lanceolate or oblong and entire, closely sessile; racemes many, recurved, in a dense pyramidal panicle; rays 7--10, very short.--Low grounds, pine barrens of N. J. to Va. and southward.

[=] 5. _Heads small or middle-sized, racemosely paniculate; leaves broad or ample, veiny, at least the lower serrate (or entire in n. 28); involucral scales obtuse._

21. S. patula, Muhl. _Stem strongly angled, smooth_ (2--4 deg. high); _leaves_ (4--8' long) _ovate_, acute, serrate, pale, _very smooth and veiny underneath_, but the _upper surface very rough_, like shagreen; racemes rather short and numerous on the spreading branches; heads rather large.--Swamps; common.

22. S. rugosa, Mill. _Rough-hairy, especially the very leafy stem_ (1--6 deg. high); _leaves ovate-lanceolate, elliptical or oblong, often thickish and very rugose_; racemes spreading; involucral scales linear; rays 6--9; the disk-flowers 4--7. (S. altissima, _Torr. & Gray_, not _L._)--Borders of fields and copses; very common, presenting a great variety of forms; usually one of the lowest of common _Golden-rods_. It flowers early. Aug.--Sept.

23. S. ulmifolia, Muhl. _Stem smooth_, the branches hairy; _leaves thin, elliptical-ovate or oblong-lanceolate_, pointed, _tapering to the base_, loosely veined, beset with soft hairs beneath; racemes panicled, recurved-spreading; involucral scales lanceolate-oblong; rays about 4.--Low copses; common.--Too near the last; distinguished only by its smooth stem and thin larger leaves.

24. S. Elliottii, Torr. & Gray. _Smooth_; stem stout (1--3 deg. high), _very leafy; leaves elliptical or oblong-lanceolate_, acute (2--3' long), closely sessile, slightly serrate, strongly veined, _thick, smooth both sides, shining above; heads in dense spreading racemes which are crowded in a close pyramidal panicle_; peduncles and achenes strigose-pubescent. (S. elliptica, _Torr. & Gray_, not _Ait._)--Swamps (fresh or brackish) near the coast, Mass. to N. J. and southward.--Heads showy, 3'' long; the rays 8--12.

25. S. neglecta, Torr. & Gray. _Smooth_; stem stout (2--4 deg. high), less leafy; _leaves thickish, smooth both sides, opaque; the upper oblong-lanceolate_, mostly acute and nearly entire; the _lower ovate-lanceolate or oblong_, sharply serrate, tapering into a petiole; _racemes short and dense, at length spreading_, disposed in an elongated or pyramidal close panicle; peduncles and achenes nearly glabrous.--Swamps, Maine to Md., Wisc., and Minn.--Heads rather large, crowded; the racemes at first erect and scarcely one-sided. Very variable, the forms approaching n. 16 and 27.

Var. linoides, Gray. The most slender form; radical leaves 4--8' long and 4--6'' wide, the upper very small, erect, branches of panicle rather few, one-sided; rays 2--5. (S. linoides, _Torr. & Gray._)--Mass. to N. J.

26. S. Boottii, Hook. Smooth, or scabrous-pubescent or below hirsute, slender, often branched, 2--5 deg. high; leaves rather finely serrate, ovate to oblong-lanceolate, pointed; the upper small, oblong to narrowly lanceolate, often entire; heads loosely racemose; rays 1--5 or none; achenes pubescent.--Dry grounds, Va. and southward.

27. S. arguta, Ait. _Smooth; stem angled; leaves_ (large and thin) _ovate_, and the upper elliptical-lanceolate, _very sharply and strongly serrate_ (entire only on the branches), _pointed at both ends_, the lowest on margined petioles; _racemes pubescent, spreading, disposed in an elongated open panicle; rays 6--7, large_; achenes usually glabrous. (S. Muhlenbergii, _Torr. & Gray._)--Copses and moist woods, N. H. to Penn., Ont., and N. E. Minn.--Racemes much shorter and looser than in the next; the involucral scales thin and more slender; the heads somewhat larger, fully 3'' long.

28. S. juncea, Ait. _Smooth throughout_ (1--3 deg. high); _radical and lower stem-leaves elliptical or lanceolate-oval, sharply serrate_ with spreading teeth, _pointed_, tapering into winged and ciliate petioles; _the others lanceolate or narrowly oblong_, slightly triple-nerved, tapering to each end, the _uppermost entire; racemes dense, naked, at length elongated and recurved, forming a crowded and flat corymb-like panicle; rays 8--12, small_. (S. arguta, _Torr. & Gray._)--Var. SCABRELLA, Gray, is somewhat roughish-pubescent (Wisc. to Ky.).--Copses and banks; common. Well distinguished by its long or drooping racemes, and the closely appressed rigid scales of the involucre, small rays, etc. Heads seldom over 2'' long, the scales small and pale.

[++][++] _Leaves more or less plainly 3-ribbed; heads in one-sided spreading or recurved racemes, forming an ample panicle. Not maritime._

[=] _Smooth and glabrous, at least the stem and bright green leaves._

[a.] _Leaves firm and rather rigid; involucral scales thickish, obtuse, quite unequal._

29. S. Missouriensis, Nutt. Smooth throughout (1--3 deg. high); _leaves linear-lanceolate_, or the lower broadly lanceolate, tapering to both ends, with very rough margins; teeth, if any, sharp and rigid; heads and dense crowded racemes nearly as in n. 28; _achenes nearly glabrous_.--Dry prairies, from Wisc. and Ind. south and westward.--Heads 1{1/2}--2'' long.

30. S. Shortii, Torr. & Gray. Stem slender, simple (2--4 deg. high), minutely roughish-pubescent above; _leaves_ (the larger 2--3' long) _oblong-lanceolate_, acute, the lower mostly serrate with a few fine teeth; racemes mostly short in a crowded panicle; _achenes silky-pubescent_.--Rocks at the Falls of the Ohio; Ark.--A handsome species; heads 3'' long, narrow.

[b.] _Leaves thinner; involucral scales thin, chiefly linear, obtuse._

31. S. serotina, Ait. _Stem_ stout (2--7 deg. high), _smooth, often glaucous; leaves quite smooth both sides_, lanceolate, taper-pointed, very sharply serrate, except the narrowed base, rough-ciliate; the ample panicle pubescent; _rays 7--14, rather long_. (S. gigantea, of previous ed.)--Copses and fence-rows; common, and presenting many varieties. Seldom very tall.

Var. gigantea, Gray. Commonly tall, 5--8 deg. high; leaves more or less pubescent or hispidulous beneath. (S. gigantea, _Ait._; S. serotina of previous ed.)--Thickets and low grounds, Can. to Tex.

32. S. rupestris, Raf. _Stem smooth, slender_, 2--3 deg. high; _leaves linear-lanceolate, tapering both ways, entire_ or nearly so; panicle narrow; _heads very small; rays 4--6, very short_.--Rocky river-banks, W. Va. to Ky. and Ind.

[=][=] _Pubescent (at least the stem) or hispidulous-scabrous._

33. S. Canadensis, L. _Stem rough-hairy_, tall and stout (3--6 deg. high); _leaves lanceolate_, pointed, sharply serrate (sometimes almost entire), _more or less pubescent beneath and rough above; heads small; rays very short_.--Borders of thickets and fields; very common.--Varies greatly in the roughness and hairiness of the stem and leaves, the latter oblong-lanceolate or elongated linear-lanceolate;--in var. PROCERA, whitish-woolly underneath; and in var. SCABRA also very rough above, often entire, and rugose-veined.

34. S. nemoralis, Ait. _Clothed with a minute and close grayish-hoary_ (soft or roughish) _pubescence_; stem simple or corymbed at the summit ({1/2}--2{1/2} deg. high); leaves _oblanceolate or spatulate-oblong_, the lower somewhat crenate-toothed and tapering into a petiole; racemes numerous, dense, at length recurved, forming a large and crowded compound raceme or panicle which is usually turned to one side; scales of the involucre linear-oblong, appressed; rays 5--9.--Dry sterile fields; very common. Flowers very bright yellow, beginning early in Aug.--Var. INCANA, Gray, of Minn., and westward, is a dwarf form, with rigid oval or oblong leaves, rather strongly serrate or entire, and the clusters of heads in a dense oblong or conical thyrse.

35. S. radula, Nutt. Stem and _oblong or obovate-spatulate leaves rigid and very rough, not hoary_, the upper sessile; scales oblong, rigid; rays 3--6; otherwise nearly as in n. 34.--Dry hills, W. Ill., Minn. Kan., and southward.

36. S. Drummondii, Torr. & Gray. _Stem_ (1--3 deg. high) _and lower surface of the broadly ovate or oval somewhat triple-ribbed leaves minutely velvety-pubescent_, some of the leaves almost entire; racemes panicled, short; scales of the involucre oblong, obtuse; rays 4 or 5.--S. W. Ill., Mo., and southward.

[+][+][+][+] _Heads in a compound corymb terminating the simple stem, not at all racemose; leaves mostly with a strong midrib._

[++] Leaves flat, not 3-nerved.

37. S. rigida, L. _Rough and somewhat hoary_ with a minute pubescence; stem stout (2--5 deg. high), very leafy; corymb dense; _leaves oval or oblong_, copiously feather-veined, thick and rigid; the upper closely sessile by a broad base, slightly serrate, the uppermost entire; heads large, over 30-flowered; the rays 7--10.--Dry soil, N. Eng. to Minn., and southward.

38. S. Ohioensis, Riddell. _Very smooth_ throughout; stem wand-like, slender, leafy (2--3 deg. high); _stem-leaves oblong-lanceolate, flat_, entire, obscurely feather-veined, closely sessile; the lower and radical ones elongated, slightly serrate toward the apex, tapering into long margined petioles; head numerous, on smooth pedicels, small, 16--20-flowered; the rays 6 or 7.--Moist meadows or prairies, W. New York to Ind. and Wisc.--Root-leaves 1 deg. long; the upper reduced to 1--2', with rough margins, like the rest.

[++][++] _Leaves somewhat folded, entire, the lower slightly 3-nerved._

39. S. Riddellii, Frank. _Smooth and stout_ (2--4 deg. high), _very leafy_, the branches of the dense corymb and pedicels rough-pubescent; _leaves linear-lanceolate, elongated_ (4--6' long), acute, partly clasping or sheathing, _mostly recurved_, the lowest elongated-lanceolate and tapering into a long keeled petiole; _heads very numerous_, clustered, 20--30-flowered; the rays 7--9.--Wet grassy prairies, Ohio to Minn. and Mo.; Ft. Monroe, Va.--Heads larger than in the last, 2--3'' long. Stem-leaves upright and partly sheathing at the base, then gradually recurved-spreading.

40. S. Houghtonii, Torr. & Gray. _Smooth; stem rather low and slender_ (1--2 deg. high); _leaves scattered, linear-lanceolate, acutish_, tapering into a narrowed slightly clasping base, or the lower into margined petioles; _heads few or several_, 20--30-flowered; the rays 7--9.--Swamps, north shore of Lake Michigan; Genesee Co., N. Y. July, Aug.--Leaves rough-margined, 2--5' long, 2--4'' wide, 1-nerved, or the lower obscurely 3-nerved above; veins obscure. Heads large, nearly 1/2' long. Scales of the involucre obtuse.

Sec. 2. EUTHAMIA. _Corymbosely much branched; heads small, sessile, in little clusters crowded in flat-topped corymbs; the closely appressed involucral scales somewhat glutinous; receptacle fimbrillate; rays 6--20, short, more numerous than the disk-flowers; leaves narrow, entire, sessile._

41. S. lanceolata, L. _Leaves lanceolate-linear, 3--5-nerved_; the nerves, margins, and angles of the branches minutely rough-pubescent; heads obovoid-cylindrical, in dense corymbed clusters; _rays 15--20_.--River-banks, etc., in moist soil; common.--Stem 2--3 deg. high; leaves 3--5' long.

42. S. tenuifolia, Pursh. Smooth, slender; _leaves very narrowly linear, mostly 1-nerved, dotted_; heads obovoid-club-shaped, in numerous clusters of 2 or 3, disposed in a loose corymb; _rays 6--12_.--Sandy fields, Mass. to Ill., and southward; common near the coast.

18. BRACHYCHAETA, Torr. & Gray. FALSE GOLDEN-ROD.

Heads and flowers nearly as in Solidago, except the pappus, which is a row of minute rather scale-like bristles, shorter than the achene.--A perennial herb, with rounded or ovate serrate leaves, all the _lower ones heart-shaped_; the small yellow heads in sessile clusters racemed or spiked on the branches. (Name composed of [Greek: brachy/s], _short_, and [Greek: chi/te], _bristle_, from the pappus.)

1. B. cordata, Torr. & Gray. Wooded hills, S. Ind. and E. Ky. to N. Ga. Oct.--Plant 2--4 deg. high, slender, more or less pubescent.

19. BELLIS, Tourn. DAISY.

Heads many-flowered, radiate; the rays numerous, pistillate. Scales of the involucre herbaceous, equal, in about 2 rows. Receptacle conical, naked. Achenes obovate, flattened, wingless, and without any pappus.--Low herbs (all but our single species natives of the Old World), either stemless, like the true _Daisy_, B. PERENNIS (which is found as an occasional escape from cultivation), or leafy-stemmed, as is the following. (The Latin name, from _bellus_, pretty.)

1. B. integrifolia, Michx. (WESTERN DAISY.) Annual or biennial, diffusely branched (4'--1 deg. high), smoothish; leaves lanceolate or oblong, the lower spatulate-obovate; heads on slender peduncles; rays pale violet-purple.--Prairies and banks, Ky. and southwestward. March--June.

20. APHANOSTEPHUS, DC.