Part 10
The rain falling through the air carries with it a certain amount of carbonic acid and ammonia, which the air always contains, and this is the whole source of the nitrogen which forms a very important part of the bodies of plants and animals. When the rain arrives at the surface of the earth, it sinks down into it and carries with it all soluble vegetable or animal matter which it meets with, together with any soluble earthy matter which may exist in the soil; this forms the sap of the tree. When it arrives at the surface of the leaf, the watery part of it combines with the carbonic acid of the air (through the influence of light) and appropriating its carbon, gives out the oxygen; this is the true respiration of plants, and is exactly the reverse of what takes place during the respiration of animals, in which case oxygen is absorbed and carbonic acid given off. The carbon thus retained by the plant combines with the elements of the water to form the solid green substance called chlorophyll, which is the basis of all the tissues of the plant, the ammonia is also decomposed, and its nitrogen combining with the oxygen and hydrogen of the water, and the carbon of the carbonic acid forms those compounds which constitute the most nourishing parts of vegetables, such as albumen, gluten, &c., and of which all the animal tissues are built up, for the production of these organic substances takes place in the vegetable only, animals simply appropriating them from their food. The sap which reaches the leaf is not all converted into chlorophyll, but also into those peculiar juices which are found in plants, some of which contain sugar, some gum, others (as the pine tribe) turpentine, and in the laurel tribe camphor, all of which are substances containing much carbon; moreover the solid wood and bark are deposited from these juices as they descend from the leaf after having been acted on by light (or the actinic power associated with it). Now, as the water, ammonia, and carbonic acid which descend with the rain are from the air, and as the vegetable is formed wholly by their absorption, it may be fairly said that the vegetable kingdom (and therefore the animal) feeds upon the air, and that the trees do not grow out of the earth but into it.
With respect to the position which the vegetable kingdom occupies in creation, there can be no doubt that it is subordinate to the animal kingdom, and takes a place between it and the mineral world, inasmuch as it prepares food from the one kingdom and transfers it to the higher. It has been supposed that the vegetable and animal kingdoms aid and support each other equally and mutually; this is true with respect to respiration, but not as regards nutrition, for although in the decomposition of animal matters, food is given off for the vegetable, yet they are quite independent of this source of nourishment. A forest of trees would be quite as well nourished if there were no animals, but on the other hand the animal kingdom would shortly cease to exist if there were no vegetables.
It was formerly supposed that the lowest grades of the animal kingdom were higher than the highest of the vegetable kingdom; this is not strictly true, the best way of viewing the connection between the two kingdoms is to approximate the lowest of each, when it will be found that our most acute physiologists are only just beginning to determine their distinctive characters.
Dr. Carpenter says:--"In the present state of science it would be very difficult, and is perhaps impossible, to lay down any definite line of demarcation between the two kingdoms, since there is no single character by which the animal or vegetable nature of any organism can be tested.
"Probably the one which is most generally applicable, among those lowest organisms which most closely approximate to one another, is not, as formerly supposed, the presence or absence of spontaneous motion, but the dependence of the being for nourishment upon organic compounds already formed, which it takes (in some way or other) into the interior of its body, or its possession of the power of obtaining its own alimentary matter by absorption from the inorganic elements on its exterior. The former is the characteristic of the animal kingdom as a whole, the latter is the attribute of the vegetable."
Both vegetables and animals begin with a simple nucleated cell, having certain vital properties, a double chain ascends from this simple type, one branch of which is developed more and more till it arrives at a perfect vegetable, and the other branch is developed till it arrives at a perfect animal. Thus, by the addition of distinctive and characteristic appendages, one being acquires properties regarded as vegetable, while by the addition of other appendages equally characteristic, the other being obtains those properties which cause it to rank as an animal; but it must not be inferred from this that all organic forms have been a simple cell at some former period, but that there are two classes of organic beings, the vegetable and the animal, and that each embraces forms, ranging from a simple cell to the highest, and that each of these forms (from the lowest to the highest) is and always was, from their first creation, the same as they now are, in individual shape and size. Some of the lower members of the animal kingdom resemble the higher members of the vegetable kingdom, both in outward appearance and in intimate structure; Dr. Darwin, who wrote scarce half-a-century ago, says that a tree should no more be considered as one plant than a branch of coral as one animal, for as it is found that in the coral hundreds of separate beings exist associated in one habitation, so (he says) should every bud on a tree be considered as a separate being. Even Dr. Carpenter, one of our most eminent physiologists, seems in a great measure to favour this idea. He says: "The radiata possess many points of affinity with the vegetable kingdom, and of these the circular arrangement of their parts is one of the most evident. Many species of sea-anemones, for instance, present an appearance so much resembling that of various composite flowers, as to have been commonly termed animal-flowers, a designation to which they seem further entitled from the small amount of sensibility they manifest, and the evident influence of light upon their opening and closing.
"But it is in the tendency to the production of compound fabrics, each containing a number of individuals, which have the power of existing independently, but which are to a certain degree connected with one another, that we recognise the greatest affinity in structure between this group and the vegetable kingdom. Every tree is made up of a large number of buds composed of leaves arranged round a common axis; each bud has the power of preserving its own life and reproducing the original structure when removed from the parent stem, if placed in circumstances favourable to its growth, and yet all are connected in the growing tree by a system of vessels which form a communication between them. This is precisely the nature of those structures which are formed by the animals of the class that may be regarded as the most characteristic of the group. Every mass of coral is the skeleton of a compound animal, consisting of a number of polypes, connected together by a soft flesh in which vessels are channelled out; the polypes are capable of existing separately, since each one, when removed from the rest, can in time produce a massive compound fabric like that of its parent, but they all contribute to the maintenance of the composite structure so long as they are in connection with it. In some instances the skeleton is stony, and is formed by the deposition of calcareous matter either in the centre of each fleshy column, so as to form a solid stem, or on its exterior so as to form a tube. In other cases it is horny, and then it may be a flexible axis in a delicate tube. Both the stony and horny corals often possess the form of plants or trees, and as their skeletons are often found with no obvious traces of the animals to which they belong, they have been accounted vegetable growths."
DESCRIPTION OF A VEGETABLE AND ITS PARTS.
As has been said before, it is extremely difficult to make any distinction between the lower tribes of the vegetable and animal kingdoms, and physiologists are not yet agreed, with respect to some members, as to which kingdom they belong. Their whole substance is made up of cellular tissue, and there are but few distinctions of parts, forming generally a broad foliaceous expansion called the "thallus," as in lichens and sea-weeds, or a sort of root composed of fibres and called the "mycelium," as in the fungi. But, a very few steps higher, the distinctive characters become so evident that they are impossible to be mistaken, the following description will therefore apply to vegetables of the more elevated character, such as trees or flowering plants.
Plants are fixed to the earth, and receive nourishment from it by imbibing its liquids, which circulating upwards through the porous structure of the plant itself, and becoming exposed to the air in the leaves, attract to themselves nourishment from that source also. The part of a plant which grows into the earth is called the root; this has a variety of forms, in some it is branched like the upper part, and these branches divide into rootlets or fibres penetrating deep into the ground, and absorbing nourishment in all directions, but this absorption does not take place from the whole surface of the root but from spots covering it, and from the slightly expanded ends of the fibres, these portions are formed of new and porous cellular tissue, and are called "spongioles." The part of plants which springs upwards from the earth is called the stem, if large the trunk, this divides into branches and twigs; stems in some cases continue for a distance more or less underground. The part of a potato plant, usually called the root, and from which the tubers or potatoes grow, is in reality an underground stem, and the fibres which spring from it are the roots; the underground suckers of mint are also portions of stem, and in some cases these are greatly expanded, they then obtain the names of "tubers" (as the potato), or "corm," as in the crocus and meadow saffron; when the stem is thin and runs along the ground, sending in roots at intervals (as in the strawberry), it is called a "runner," when thicker and running horizontally under the ground, it is called a "rhizome."
The stem consists of a central portion, either made up of long bundles of woody fibre running side by side, as in the endogenæ, or deposited in rings and having a central cylinder of pith, as in the exogenæ. In these the wood at the central part (or the oldest) is called "duramen," or "heart-wood," while that at the outer part or nearest to the bark (the newest), is called "alburnum" or "sap-wood." The former is the harder and the latter the softer portion. From the pith in the centre, through the woody part, rays or laminæ of cellular tissue similar to that which constitutes the pith itself, are sent outwards through the woody rings to the inner part of the bark, which they form; these are called medullary rays, and may be seen in wood which has been cut across the grain, they are often called the "silver grain," and are very evident in oak, beech, and elm, the inner part of the bark is called the "liber." The bark itself is made of cellular tissue dried and hardened by age; the outer portion (called the "epidermis") is in many cases shed, and may be constantly seen hanging loosely from the bark of the birch and other trees in loose white silvery portions. The outer part of the bark of some trees is so largely developed as to be of considerable thickness, this is especially the case in the Cork oak (_Quercus suber_), which is the cork of commerce. The bark cannot stretch as the circumference of the tree increases, it is therefore split up and cracked, which accounts for the rough state of it on those trees which do not shed the outer part.
The chief appendage of the branches, is called a leaf, it grows from a small projection called a leaf-bud, which contains the leaf rolled up. The method in which this occurs is different in different plants, in some it is folded backwards and forwards, in others doubled up with the opposite leaf alternately, and in various forms in other plants. The leaf consists of two parts, the stalk (petiole) and the blade (lamina); the blade is of different forms, and has ribs and veins covering it, in some of a reticulated or network pattern, these belong to the exogenæ, and in others running parallel, which is the kind of venation found in the leaves of the endogenæ. When leaves are placed on a stalk, they are said to be "petiolate," when without one, "sessile."
When leaves are not separated into different portions (although they may be much notched) they are said to be "simple," as in the oak or willow, but when divided into separate portions, as in the ash, they are said to be "compound."
The following are the chief forms of leaves, named according to variations of their several parts:
At the base of many leaves are a pair of scales called stipules; the petiole or leaf-stalk is generally cylindrical, but frequently triangular, and in grasses it is flat and surrounds the stem, this is called a sheath; when leaves are narrow and not expanded into a lamina, as in the pine tribe, they are said to be "acicular."
The stalks which bear the flowers are called "pedicels," at the base of which are a pair of scales called "bracts;" when these are large and expanded, so as to enclose the flowers, they are called "spathes" (this is seen in the arum), and when there are a number of flower-stalks arising from one point the bracts there collected are called an "involucre." A flower consists of several parts, the outermost green scales, composing a set, are called the calyx, and each part of it, is called a "sepal," within this is the "corolla" or that coloured part which forms the most characteristic feature of the flower, each part of the corolla is called a "petal;" when the corolla consists of but one piece, it is called "monopetalous," and when of many, "polypetalous."
The forms of corolla vary according to the form and the mode in which the petals are placed, whether united or separated, and to what extent, whether regular or irregular; the most usual forms are the following:--
Within the corolla are placed the "stamens" (male reproductive organs) these consist generally of two parts, the head and stalk, the former called the "anther" and the latter the "filament," which last is sometimes absent, and the anther is said to be "sessile;" on the surface of the anther is the "pollen" or fertilising dust.
Within the centre of the flower is the "pistil" (female reproductive organ), this consists of one or several cells called "ovaries," from the pistil a tube rises, having an expanded end called the "stigma," it is by the application of the pollen dust to this stigma that the ovaries are fertilised, and the various insects, especially bees, who seek for honey, shake off by their movements the pollen from the anthers and cause it to be applied to the stigma, thus unconsciously performing a necessary office for the plant while they rob it of that only which is not required.
The stamens are sometimes separate, sometimes bound up into one or more bundles, and are placed in various situations, names are given to describe such arrangements as follows:--
Stamens in one bundle, Monadelphous. Stamens in two bundles, Diadelphous. Stamens in more than two bundles, Polyadelphous. Filaments placed directly below the pistil, Hypogynous. Placed upon the sides of the calyx, Perigynous. On the sides of the corolla, Epipetalous. On the top of the ovary, Epigynous.
When the ovaries are fertilised the flower dies and they begin to enlarge and ripen to form the fruit, which is the pistil enlarged, and contains the ovules ripened into seeds.
The fruits of different plants are known by various names according to the state of development of the various parts composing them. If the ripe fruit split open so as to let out the seeds (as in the common pea) it is called "dehiscent," if it do not so split (as in the apple) it is said to be "indehiscent;" the outer part of fruit is called the "pericarp," and this may be soft and fleshy as in the apple or cherry, or hard as in the filbert. The following are the names of the principal varieties of fruit:--
The seed is the ovule ripened, it contains the germ of the future plant, called the "embryo," the outer part of the seed is called the "testa," and the space between this and the embryo is generally filled with starchy matter called the "albumen." The embryo consists of the plumule or stem, the "radicle" or root, and the cotyledons or leaves of the future plant; when the seed has but one cotyledon it is called "monocotyledonous" and when it has two "dicotyledonous."
Flowers are arranged in various ways upon the plants which produce them, and receive names accordingly; the whole arrangement of flowers is called the "inflorescence."
The following are the principal forms of inflorescence:--
The UMBEL, in which all the flower-stalks (pedicels) radiate from one point, as in the carrot (daucus carota). Umbels are sometimes compound, that is, the flowers are placed in umbels at the end of stalks themselves radiating from a point and so forming an umbel, as in the Hemlock (Conium maculatum).
The SPIKE, is that kind of infloresence in which all the flowers are seated without stalks upon a general peduncle or axis, as in the Plantain (Plantago media), in which the spike is entire, or as the Lavender (Lavandula Vera), in which the spike is interrupted, that is, the inflorescence is not continuous.
CATKIN, or AMENTUM, is the same as a spike, but in which the flowers are imperfectly developed, as in the Hazel (Corylus), Willow (Salix), White Poplar (Populus Alba), &c.
The RACEME has the inflorescence placed along a common axis or centre, the same as a spike, but with the flowers placed upon stalks instead of being sessile, as in Water-cress (Nasturtium officinale) and Red Currant (Ribes rubrum).]
The CORYMB is a form of inflorescence pretty much the same as raceme, but the flowers of which proceed upwards till they are all on a level, as in Candy-tuft (Iberis).
The PANICLE, the same as the raceme, but having the flower stalks themselves divided into branches, as in Creeping Soft Grass (Holcus mollis). This and the spike are the most usual form of inflorescence found among grasses, in many of which the panicle, however, is often contracted almost to a spike.
The CYME, this resembles the panicle shortened in such a manner as to become flattened or almost corymbose, as in the Elder Tree (Sambucus nigra), in which there are five principal stalks of inflorescence.
CAPITULUM, in which the flowers arise from a broad round head or receptacle as in the composite flowers, such as Chamomile (Anthemis nobilis); in such flowers the star-like ray of florets are called the florets of the ray, and those composing the centre the florets of the disc.
The SPADIX is that form of inflorescence in which the expanded bract, called a spathe, forms a sort of sheath inclosing the flowers. This spathe is white in the example given, and is often mistaken for part of the flower itself, as in the Wake-robin or Arum (Arum maculatum).
The vegetable kingdom is divided into three great natural families, the Acrogenæ, the Endogenæ, and the Exogenæ.
The acrogenous plants are those which as a general rule have neither branches, leaves, nor flowers; they are almost wholly made up of cellular tissue, and are many of them so minute that they are quite invisible individually to the unassisted eye, and are among the most wonderful works of the Creator, having an amount of beauty in form and elaboration of ornament bestowed on them, quite equal to anything among the higher and larger creations, and yet some of these are so small that tens of millions may be placed in the space of a cubic inch, of such are the Diatomaceæ and Desmidiaceæ.
The acrogens take an immense range in the scale of organisation, from the ferns (which appear but little inferior to the exogenæ or endogenæ, have stems and leaves, and in some cases, as in hot and moist climates, assume the size and form of a tree), to the very lowest state of vegetable existence, consisting of simple cells uniting into strings or forming simple threads, such as the green algæ which form on stagnant waters and damp ground or wood-work, and the mould or mildew which forms on all decomposing substances. The general name for these acrogens, "cryptogamia," which has been in use for a long time and is commonly still used, indicates that the reproductive organs are invisible, hence the expression used by one of Shakespeare's characters, "We have the secret of fern seed, we walk invisible;" but this is not really the case, for the "sorri" at the back of fern-leaves, are vessels filled with spore cases each having a number of angular seeds within it; the lower tribes of the acrogens do not commonly grow from seed but by an extension of their several parts by the development of the cells of which they are composed, and by their separation into portions.