Chapter 17

How beautifully simple then, by the aid of these discoveries, appears the process of nutrition in animals, the formation of their organs, in which vitality chiefly resides. Those vegetable constituents which are used by animals to form blood contain the essential ingredients of blood ready formed. In point of fact, vegetables produce in their organism the blood of all animals; for the carnivora, in consuming the blood and flesh of the graminivora, consume, strictly speaking, the vegetable principles which have served for the nourishment of the latter. In this sense we may say the animal organism gives to blood only its form; and, further, that it is incapable of forming blood out of other compounds which do not contain the chief ingredients of that fluid.

Animal and vegetable life are, therefore, closely related, for the first substance capable of affording nutriment to animals is the last product of the creative energy of vegetables. The seemingly miraculous in the nutritive power of vegetables disappears in a great degree, for the production of the constituents of blood cannot appear more surprising than the occurrence of the principal ingredient of butter in palm-oil and of horse-fat and train-oil in certain of the oily seeds.

_IV.--Food the Fuel of Life_

We have still to account for the use in food of substances which are destitute of nitrogen but are known to be necessary to animal life. Such substances are starch, sugar, gum, and pectine. In all of these we find a great excess of carbon, with oxygen and hydrogen in the same proportion as water. They therefore add an excess of carbon to the nitrogenised constituents of food, and they cannot possibly be employed in the production of blood, because the nitrogenised compounds contained in the food already contain exactly the amount of carbon which is required for the production of fibrine and albumen. Now, it can be shown that very little of the excess of this carbon is ever expelled in the form either of solid or liquid compounds; it must be expelled, therefore, in the gaseous state. In short, these compounds are solely expended in the production of animal heat, being converted by the oxygen of the air into carbonic acid and water. The food of carnivorous animals does not contain non-nitrogenised matters, so that the carbon and hydrogen necessary for the production of animal heat are furnished in them from the waste of their tissues.

The transformed matters of the organs are obviously unfit for the further nourishment of the body--that is, for the increase or reproduction of the mass. They pass through the absorbent and lymphatic vessels into the veins, and their accumulation in these would soon put a stop to the nutritive process were it not that the blood has to pass through a filtering apparatus, as it were, before reaching the heart. The venous blood, before returning to the heart, is made to pass through the liver and the kidneys, which separate from it all substances incapable of contributing to nutrition. The new compounds containing the nitrogen of the transformed organs, being utterly incapable of further application in the system, are expelled from the body. Those which contain the carbon of the transformed tissues are collected in the gall-bladder as bile, a compound of soda which, being mixed with water, passes through the duodenum and mixes with chyme. All the soda of the bile, and ninety-nine-hundredths of the carbonaceous matter which it contains, retain the capacity of re-absorption by the absorbents of the small and large intestines--a capacity which has been proved by direct experiment.

The globules of the blood, which in themselves can be shown to take no share in the nutritive process, serve to transport the oxygen which they give up in their passage through the capillary vessels. Here the current of oxygen meets with the carbonaceous substances of the transformed tissues, and converts their carbon into carbonic acid, their hydrogen into water. Every portion of these substances which escapes this process of oxidation is sent back into the circulation in the form of bile, which by degrees completely disappears.

It is obvious that in the system of the graminivora, whose food contains relatively so small a proportion of the constituents of blood, the process of metamorphosis in existing tissues, and consequently their restoration or reproduction, must go on far less rapidly than in the carnivora. Otherwise, a vegetation a thousand times as luxuriant would not suffice for their sustenance. Sugar, gum, and starch, which form so large a proportion of their food, would then be no longer necessary to support life in these animals, because in that case the products of waste, or metamorphosis of organised tissues, would contain enough carbon to support the respiratory process.

When exercise is denied to graminivorous and omnivorous animals this is tantamount to a deficient supply of oxygen. The carbon of the food, not meeting with a sufficient supply of oxygen to consume it, passes into other compounds containing a large excess of carbon--or, in other words, fat is produced. Fat is thus an abnormal production, resulting from a disproportion of carbon in the food to that of the oxygen respired by the lungs or absorbed by the skin. Wild animals in a state of nature do not contain fat. The production of fat is always a consequence of a deficient supply of oxygen, for oxygen is absolutely indispensable for the dissipation of excess of carbon in the food.

_V.--Animal Life-Chemistry_

The substances of which the food of man is composed may be divided into two classes--into nitrogenised and non-nitrogenised. The former are capable of conversion into blood, the latter incapable of this transformation. Out of those substances which are adapted to the formation of blood are formed all the organised tissues. The other class of substances in the normal state of health serve to support the process of respiration. The former may be called the plastic elements of nutrition; the latter, elements of respiration.

Among the former we may reckon--vegetable fibrine, vegetable albumen, vegetable casein, animal flesh, animal blood.

Among the elements of respiration in our food are--fat, starch, gum, cane sugar, grape-sugar, sugar of milk, pectine, bassorine, wine, beer, spirits.

The nitrogenised constituents of vegetable food have a composition identical with that of the constituents of the blood.

No nitrogenised compound the composition of which differs from that of fibrine, albumen, and casein, is capable of supporting the vital process in animals.

The animal organism undoubtedly possesses the power of forming from the constituents of its blood the substance of its membranes and cellular tissue, of the nerves and brain, of the organic part of cartilages and bones. But the blood must be supplied to it ready in everything but its form--that is, in its chemical composition. If this is not done, a period is put to the formation of blood, and, consequently, to life.

The whole life of animals consists of a conflict between chemical forces and the vital power. In the normal state of the body of an adult these stand in equilibrium: that is, there is equilibrium between the manifestations of the causes of waste and the causes of supply. Every mechanical or chemical agency which disturbs the restoration of this equilibrium is a cause of disease.

Death is that condition in which chemical or mechanical powers gain the ascendancy, and all resistance on the part of the vital force ceases. This resistance never entirely departs from living tissues during life. Such deficiency in resistance is, in fact, a deficiency in resistance to the action of the oxygen of the atmosphere.

Disease occurs when the sum of vital force, which tends to neutralise all causes of disturbance, is weaker than the acting cause of disturbance.

Should there be formed in the diseased parts, in consequence of the change of matter, from the elements of the blood or of the tissue, new products which the neighbouring parts cannot employ for their own vital functions; should the surrounding parts, moreover, be unable to convey these products to other parts where they may undergo transformation, then these new products will suffer, at the place where they have been formed, a process of decomposition analogous to putrefaction.

In certain cases, medicine removes these diseased conditions by exciting in the vicinity of the diseased part, or in any convenient situation, an artificial diseased state (as by blisters), thus diminishing by means of artificial disturbance the resistance offered to the external causes of change in these parts by the vital force. The physician succeeds in putting an end to the original diseased condition when the disturbance artificially excited (or the diminution of resistance in another part) exceeds in amount the diseased state to be overcome.

The accelerated change of matter and the elevated temperature in the diseased part show that the resistance offered by the vital force to the action of oxygen is feebler than in the healthy state. But this resistance only ceases entirely when death takes place. By the artificial diminution of resistance in another part, the resistance in the diseased organ is not, indeed, directly strengthened; but the chemical action, the cause of the change of matter, is diminished in the diseased part, being directed to another part, where the physician has succeeded in producing a still more feeble resistance to the change of matter, to the action of oxygen.

SIR CHARLES LYELL

The Principles of Geology

Sir Charles Lyell, the distinguished geologist, was born at Kinnordy, Forfarshire, Scotland, Nov. 14, 1797. It was at Oxford that his scientific interest was first aroused, and after taking an M.A. degree in 1821 he continued his scientific studies, becoming an active member of the Geological and Linnæan Societies of London. In 1826 he was elected a fellow of the Royal Society, and two years later went with Sir Roderick Murchison on a tour of Europe, and gathered evidence for the theory of geological uniformity which he afterwards promulgated. In 1830 he published his great work, "Principles of Geology: Being an Attempt to Explain the Former Changes of the Earth's Surface by References to Causes now in Action," which converted almost the whole geological world to the doctrine of uniformitarianism, and may be considered the foundation of modern geology. Lyell died in London on February 22, 1875. Besides his great work, he also published "The Elements of Geology," "The Antiquity of Man," "Travels in North America," and "The Student's Elements of Geology."

_I.--Uniformity in Geological Development_

According to the speculations of some writers, there have been in the past history of the planet alternate periods of tranquillity and convulsion, the former enduring for ages, and resembling the state of things now experienced by man; the other brief, transient, and paroxysmal, giving rise to new mountains, seas, and valleys, annihilating one set of organic beings, and ushering in the creation of another. These theories, however, are not borne out by a fair interpretation of geological monuments; but, on the contrary, nature indicates no such cataclysms, but rather progressive uniformity.

Igneous rocks have been supposed to afford evidence of ancient paroxysms of nature, but we cannot consider igneous rocks proof of any exceptional paroxysms. Rather, we find ourselves compelled to regard igneous rocks as an aggregate effect of innumerable eruptions, of various degrees of violence, at various times, and to consider mountain chains as the accumulative results of these eruptions. The incumbent crust of the earth is never allowed to attain that strength and coherence which would be necessary in order to allow the volcanic force to accumulate and form an explosive charge capable of producing a grand paroxysmal eruption. The subterranean power, on the contrary, displays, even in its most energetic efforts, an intermittent and mitigated intensity. There are no proofs that the igneous rocks were produced more abundantly at remote periods.

Nor can we find proof of catastrophic discontinuity when we examine fossil plants and fossil animals. On the contrary, we find a progressive development of organic life at successive geological periods.

In Palæozoic strata the entire want of plants of the most complex organisation is very striking, for not a single dicotyledonous angiosperm has yet been found, and only one undoubted monocotyledon. In Secondary, or Mesozoic, times, palms and some other monocotyledons appeared; but not till the Upper Cretaceous era do we meet with the principal classes and orders of the vegetable kingdom as now known. Through the Tertiary ages the forms were perpetually changing, but always becoming more and more like, generically and specifically, to those now in being. On the whole, therefore, we find progressive development of plant life in the course of the ages.

In the case of animal life, progression is equally evident. Palæontological research leads to the conclusion that the invertebrate animals flourished before the vertebrate, and that in the latter class fish, reptiles, birds, and mammalia made their appearance in a chronological order analogous to that in which they would be arranged zoologically according to an advancing scale of perfection in their organisation. In regard to the mammalia themselves, they have been divided by Professor Owen into four sub-classes by reference to modifications of their brain. The two lowest are met with in the Secondary strata. The next in grade is found in Tertiary strata. And the highest of all, of which man is the sole representative, has not yet been detected in deposits older than the Post-Tertiary.

It is true that in passing from the older to the newer members of the Tertiary system we meet with many chasms, but none which separate entirely, by a broad line of demarcation, one state of the organic world from another. There are no signs of an abrupt termination of one fauna and flora, and the starting into life of new and wholly distinct forms. Although we are far from being able to demonstrate geologically an insensible transition from the Eocene to the Miocene, or even from the latter to the recent fauna, yet the more we enlarge and perfect our general survey the more nearly do we approximate to such a continuous series, and the more gradually are we conducted from times when many of the genera and nearly all the species were extinct to those in which scarcely a single species flourished which we do not know to exist at present. We must remember, too, that many gaps in animal and floral life were due to ordinary climatic and geological factors. We could, under no circumstances, expect to meet with a complete ascending series.

The great vicissitudes in climate which the earth undoubtedly experienced, as shown by geological records, have been held to be themselves proof of sudden violent revolutions in the life-history of the world. But all the great climatic vicissitudes can be accounted for by the action of factors still, in operation--subsidences and elevations of land, alterations in the relative proportions and position of land and water, variations in the relative position of our planet to the sun and other heavenly bodies.

Altogether, the conclusion is inevitable that from the remotest period there has been one uniform and continuous system of change in the animate and inanimate world, and accordingly every fact collected respecting the factors at present at work in forming and changing the world, affords a key to the interpretation of its part. And thus, although we are mere sojourners on the surface of the planet, chained to a mere point in space, enduring but for a moment of time, the human mind is enabled not only to number worlds beyond the unassisted ken of mortal eye, but to trace the events of indefinite ages before the creation of our race, and to penetrate into the dark secrets of the ocean and the heart of the solid globe.

_II.--Changes in the Inorganic World now in Progress_

The great agents of change in the inorganic world may be divided into two principal classes--the aqueous and the igneous. To the aqueous belong rain, rivers, springs, currents, and tides, and the action of frost and snow; to the igneous, volcanoes and earthquakes. Both these classes are instruments of degradation as well as of reproduction. But they may also be regarded as antagonist forces, since the aqueous agents are incessantly labouring to reduce the inequalities of the earth's surface to a level; while the igneous are equally active in restoring the unevenness of the external crust, partly by heaping up new matter in certain localities, and partly by depressing one portion of the earth's envelope and forcing out another.

We will treat in the first place of the aqueous agents.

RAIN AND RIVERS. When one considers that in some parts of the world as much as 500 or 600 inches of rain may fall annually, it is easy to believe that rain _qua_ rain may be a denuding and plastic agent, and in some parts of the world we find evidence of its action in earth pillars or pyramids. The best example of earth pillars is seen near Botzen, in the Tyrol, where there are hundreds of columns of indurated mud, varying in height from 20 feet to 100 feet. These columns are usually capped by a single stone, and have been separated by rain from the terrace of which they once formed a part.

As a rule, however, rain acts through rivers. The power of rivers to denude and transport is exemplified daily. Even a comparatively small stream when swollen by rain may move rocks tons in weight, and may transport thousands of tons of gravel. The greatest damage is done when rivers are dammed by landslips or by ice. In 1818 the River Dranse was blocked by ice, and its upper part became a lake. In the hot season the barrier of ice gave way, and the torrent swept before it rocks, forests, houses, bridges, and cultivated land. For the greater part of its course the flood resembled a moving mass of rock and mud rather than of water. Some fragments of granite rock of enormous size, which might be compared to houses, were torn out and borne down for a quarter of a mile.

The rivers of unmelted ice called the glaciers act more slowly, but they also have the power of transporting gravel, sand, and boulders to great distances, and of polishing and scoring their rocky channels. Icebergs, too, are potent geological agents. Many of them are loaded with 50,000 to 100,000 tons of rock and earth, which they may carry great distances. Also in their course they must break, and polish, and scratch the peaks and points of submarine mountains.

Coast ice, likewise, may transport rocks and earth. Springs also must be considered as geological agents affecting the face of the globe.

But running water not only denudes it, but also creates land, for lakes, seas, rivers are seen to form deltas. That Egypt was the gift of the Nile was the opinion of the Egyptian priests, and there can be no doubt that the fertility of the alluvial plain above Cairo, and the very existence of the delta below that city, are due to the action of that great river, and to its power of transporting mud from the interior of Africa and depositing it on its inundated plains as well as on that space which has been reclaimed from the Mediterranean and converted into land. The delta of the Ganges and Brahmapootra is more than double that of the Nile. Even larger is the delta of the Mississippi, which has been calculated to be 12,300 square miles in area.

TIDES AND CURRENTS. The transporting and destroying and constructive power of tides and currents is, in many respects, analogous to that of rivers, but extends to wider areas, and is, therefore, of more geological importance. The chief influence of the ocean is exerted at moderate depths below the surface on all areas which are slowly rising, or attempting, as it were, to rise above the sea; but its influence is also seen round the coast of every continent and island.

* * * * *

We shall now consider the igneous agents that act on the earth's surface. These agents are chiefly volcanoes and earthquakes, and we find that both usually occur in particular parts of the world. At various times and at various places within historical times volcanic eruptions and earthquakes have both proved their potency to alter the face of the earth.

The principal geological facts and theories with regard to volcanoes and earthquakes are as follows.

The primary causes of the volcano and the earthquake are to a great extent the same, and connected with the development of heat and chemical action at various depths in the interior of the globe.

Volcanic heat has been supposed to be the result of the original high temperature of the molten planet, and the planet has been supposed to lose heat by radiation. Recent inquiries, however, suggest that the apparent loss of heat may arise from the excessive local development of volcanic action.

Whatever the original shape of our planet, it must in time have become spheroidal by the gradual operation of centrifugal force acting on yielding materials brought successively within its action by aqueous and igneous causes.

The heat in mines and artesian wells increases as we descend, but not in uniform ratio in different regions. Increase at a uniform ratio would imply such heat in the central nucleus as must instantly fuse the crust.

Assuming that there are good astronomical grounds for inferring the original fluidity of the planet, yet such pristine fluidity need not affect the question of volcanic heat, for the volcanic action of successive periods belongs to a much more modern state of the globe, and implies the melting of different parts of the solid crust one after the other.

The supposed great energy of the volcanic forces in the remoter periods is by no means borne out by geological observations on the quantity of lava produced by single eruptions in those several periods.

The old notion that the crystalline rocks, whether stratified or unstratified, such as granite and gneiss, were produced in the lower parts of the earth's crust at the expense of a central nucleus slowly cooling from a state of fusion by heat has now had to be given up, now that granite is found to be of all ages, and now that we know the metamorphic rocks to be altered sedimentary strata, implying the denudation of a previously solidified crust.

The powerful agency of steam or aqueous vapour in volcanic eruptions leads us to compare its power of propelling lava to the surface with that which it exerts in driving water up the pipe of an Icelandic geyser. Various gases also, rendered liquid by pressure at great depths, may aid in causing volcanic outbursts, and in fissuring and convulsing the rocks during earthquakes.

The chemical character of the products of recent eruptions suggests that large bodies of salt water gain access to the volcanic foci. Although this may not be the primary cause of volcanic eruptions, which are probably due to the aqueous vapour intimately mixed with molten rock, yet once the crust is shattered through, the force and frequency of eruptions may depend in some measure on the proximity of large bodies of water.

The permanent elevation and subsidence of land now observed, and which may have been going on through past ages, may be connected with the expansion and contraction of parts of the solid crust, some of which have been cooling from time to time, while others have been gaining heat.