Part 13

The feet of the Camels and of the Llamas are very different in form from those of all the other Ruminants. They are, it is true, deeply divided, like those of the latter, into two apparent toes; but cannot be said, like them, to part the hoof, for they have no real hoof, and the extremities of their protruded toes are armed only with short, thick, and crooked claws. These toes are in the Camels united posteriorly by a horny process, which is wanting in the Llamas. The teeth of both are nearly similar: they consist of six incisors in the lower jaw and two in the upper; of two canines in each; and of six molars in the upper, and five in the lower, on each side. None of the other Ruminants exhibit the least appearance of cutting teeth in the upper jaw. The nostrils of both consist externally of mere fissures in the skin, which may be opened and closed at pleasure, and which are surrounded by a naked muzzle; and their upper lip is divided into two distinct portions, which are very extensible, and capable of much separate motion.

The species of the group, of which the Llama forms the type, have been involved by the imperfect descriptions of naturalists in almost inextricable confusion. No less than five have been admitted; but the variations of colour and of size, and the degree of length and fineness of the wool, differences rather commercial than natural, afford almost the only positive distinctions that have yet been laid down between them; and when we consider that some of them have been for ages in a state of domestication, it will readily be allowed that such characters as these are, to say the least, trivial and uncertain. Our animals, which are nearly four feet in height at the shoulder, and somewhat more than five feet to the top of the head, have the neck, the back, the sides, and the tail, which is rather short, covered with a beautiful coat of long, bright brown, woolly hair. The long and pointed ears, and the small and attenuated head, on which the hair is short, close, and even, are of a grayish mouse-colour; the outside of the legs is of the same colour with the sides of the body; and their inside, as also the under part of the body and the throat, pure white. The hair on the limbs is short and smooth. In these respects they offer but little to distinguish them from any of the animals which have been exhibited in this country under the various names of Llamas, Pacos, and Guanacos. There is, however, at present in the Garden of the Zoological Society, an animal, which besides being of larger size, covered with longer and coarser wool, and entirely white (which latter circumstance may be purely accidental), differs remarkably in the form of the forehead, which in it is perfectly flat, while in our animals it rises in a strong curve. This character, it is probable, affords a permanent ground of distinction, although we venture not at present to speak decidedly respecting it.

The Llamas congregate together in considerable herds on the sides of the Andes, and generally in the colder and more elevated regions. When the Spaniards first arrived in Peru they were the only beasts of burden employed by the natives; and even at the present day, when horses have become so excessively common, they are usually preferred for passing the mountains, on which their sureness of footing, exceeding even that of the mule, gives them a manifest superiority. Generally speaking they are quiet, docile and timid; but they occasionally exhibit much spitefulness, especially if teased or ill treated. Their mode of evincing this is very peculiar, and consists in darting their saliva through their nostrils with considerable force. Like all the other Ruminants they subsist entirely on vegetables. Those in the Tower Menagerie have a particular fondness for carrots; and if one of these is abstracted from them while they are eating, their anger is immediately roused, and they spit, as it is termed, with the greatest vehemence, covering with their saliva a surface of three or four yards in extent. One of the animals in the cut is represented in the act.

THE MALAYAN RUSA-DEER.

_CERVUS EQUINUS._ CUV.

The Deer constitute a numerous and beautiful group of Ruminants, which are readily distinguished by the graceful symmetry of their form, by their long and slender, but firm and sinewy, legs, by their broad and pointed ears, and by the comparative shortness of their tails; but more especially by the generally large and branching horns which ornament the heads of the males. Like all the ruminating animals, with the exception of those mentioned in the preceding article, they are furnished with eight cutting-teeth in the lower jaw, opposed to a callous and toothless surface in the upper; and with expanded, flat, and deeply bifurcated hoofs, constituting two distinct and apparent toes, above which they have also the rudiments of two others. Some of the species have canine teeth in the upper jaw, generally in the males alone; and they have all six molars on each side. In the greater number of them the nostrils are surrounded by a naked muzzle; and most of them are also provided with a sinus or sac, of greater or less extent, immediately beneath the inner angle of the eye, called the sub-orbital sinus, the _larmier_ of the French zoologists.

The horns, which form the most distinguishing character of the genus, are perfectly solid throughout their whole extent. Their form varies very considerably in the different races; but they are constantly uniform in the same species, unless accidentally or artificially perverted from their natural growth. In some they are simple at the base and terminate in a broad and palmate expansion, which is variously lobed and divided; in others they are more or less branched, giving off antlers in different directions; and in some few they are short and nearly simple. They fall off and are renewed annually in all the species which inhabit the northern and temperate regions of the earth, and in those in which they attain any considerable size; but Sir T. Stamford Raffles was of opinion, and his opinion has been in some measure confirmed by the observations of Major C. Hamilton Smith, that several of the tropical species with small and nearly simple horns are exempted from this general law. The horns are smaller and less developed in the young than in the full grown and adult animal, and diminish again in size, and frequently become irregular, as he advances in age. In one species alone, the Rein-Deer of the North, the female wears the same palmy honours with the male; but they do not in her reach the same enormous extent.

The high degree of domestication to which this latter species has been brought, and the invaluable services which it renders to the Laplander, added to the tranquil content which most of the deer manifest in a state of captivity, afford sufficient proofs that there is nothing in the constitution of the group repugnant to their being tamed and familiarized with man; but from none of the other races have any real or essential advantages been as yet derived. The quiet confidence, mixed with a certain air of cautious timidity, which they exhibit in their half-restricted state, in the park or the chase, where they are kept more for ornament than use, is perfectly indicative of their general character. But the very mildness of their disposition has been turned to their disadvantage, and one of the gentlest of animals, because endowed by nature with a high degree of fleetness, with some sagacity, and with a certain share of timidity, has been marked out by man as the chosen victim of his cruelty, disguised under the captivating name of sport.

The Samboo Deer, as the present species is called by his keepers, belongs to the Rusa group, which are distinguished from the rest of the genus by their horns being provided with a single antler at the base, and with a lateral snag which forms a kind of bifurcation towards the extremity. They are usually of large stature and nearly uniform colours, and are, for the most part, furnished with a rough and shaggy mane, a broad and expanded muzzle, and sub-orbital openings of considerable size. The handsome Stag now before us is dark cinereous brown above, nearly black on the throat and breast, and light fawn, intermixed with dirty white, on the inside of the limbs. His eyes are surrounded by a fawn-coloured disc, and patches of the same colour occupy the fore knees, and a space above each of the hoofs in front. His nose, which is black, is enveloped in an extensive muzzle; his ears are nearly naked on the inside, and marked by a patch of dirty white at the base externally; and his mane, which spreads downwards over the neck and throat, is remarkably thick and heavy. His tail is black above, and light fawn beneath; and a disc of the latter colour occupies the posterior part of the buttocks, having on each side a blackish line which separates it from the lighter tinge of the inside of the thighs. His horns, when properly grown, consist of a broad burr, from which the pointed basal antler rises almost perpendicularly to the extent of nine or ten inches; of a stem, which is first directed outwards, and then forms a bold curve inwards; and of a snag, or second antler of smaller size, arising from the stem near its extremity on the posterior and internal side, and forming with it a terminal fork, the branch however being shorter than the stem, and not exceeding five or six inches in length. The entire length of the horns is about two feet; they are of a dark colour, very strong, and deeply furrowed throughout.

The foregoing description of the horns, it should be observed, is taken from those of the year before last, which were of the genuine or normal form. Those of the last year, which are represented in the cut prefixed, were from some cause or other remarkably different, that of the right side especially exhibiting a singular monstrosity in the production of additional branches of irregular form. Whether this was the effect of disease or of advancing age, or whether it arose solely from some temporary and accidental cause, will probably be determined by the growth of the present year, which is not yet sufficiently advanced to enable us to ascertain its probable form.

With regard to the sub-orbital sinus, which in this and all the neighbouring species is of very considerable size, its uses are evidently connected with the function of respiration, and probably also with the sense of smell. It is denoted externally by a longitudinal fissure, placed beneath the inner angle of each of the eyes, and leading into a sac or cavity, which in some cases communicates internally with the nose; and its inner surface is lined by a membrane abundantly supplied with follicles for the secretion of mucus, which is sometimes produced in very large quantities. This latter circumstance has induced some naturalists to regard these openings as mere cuticular appendages. That they really, in some species at least, communicate with the nostrils, is proved by the observations of Mr. White of Selbourne, who states that in consequence of this communication the Fallow-Deer are enabled to take long-continued draughts with their noses deeply immersed in the water, the air in the mean time passing through the sub-orbital slits. So singular a statement was naturally enough doubted and called in question; but it has never, so far as we know, been impugned on ocular testimony; while it has received the fullest confirmation from other observations made upon the very species now under consideration, in which the air passing from the sub-orbital sinus, while the animal drinks, may be felt by the hand, and even affects the flame of a candle. Another proof of the connexion of these cavities with the nose is derived from the fact that the animals which are provided with them frequently apply their orifices, equally with those of the nostrils, to the food which they are about to take, opening and shutting them with great rapidity.

The subject of the present article, which, like all the rest of the minor group of which he forms a part, is a native of India and of the Indian Islands, was a present to his Majesty, who kept him for some time, in company with another of the same species, at large in the great park at Windsor. As both, however, happened to be males, they disagreed so violently, and their quarrels at length rose to such a pitch, that in order to preserve peace it was found absolutely necessary to separate them; and our animal, as the most outrageous of the two, was dismissed the royal service, and condemned to the captivity of the Tower. Since this period he has become exceedingly tame, the cause of his former ill temper being removed, and demeans himself as quietly as the most harmless and gentlest of his tribe.

THE INDIAN ANTELOPE.

_ANTILOPE CERVICAPRA._ PALL.

In the elegant symmetry of their form and the light and graceful agility of their motions, the Antelopes are superior even to the Deer, whom, however, they closely resemble, not merely in outward shape, but also in internal structure. Like them, in addition to the coincidence of a slightly made and beautifully proportioned figure, they are frequently furnished with a naked muzzle, and with the same remarkable sinus beneath the inner angle of the eye; and their ears are generally of considerable size, erect, and pointed. But they are strikingly distinguished from them and from all the other animals of the order by the peculiar character of their horns, which are formed of an elastic sheath enclosing a solid nucleus, and are for the most part common to the females as well as to the males. They have no canine teeth, and exhibit no appearance of a beard such as is seen in the Goats. The horns vary greatly in the different races; they are sometimes straight and upright, at other times slightly curved, and frequently spirally twisted with the most beautiful regularity: they are usually surrounded by elevated rings or by a spiral ridge, are constantly of the same form in the same species, and are not subject to an annual falling off and renewal, as in the Deer, from which they differ also in their mode of growth, the horns of the latter group lengthening at their apices, while those of the former receive their increase at the base.

In their natural habits the numerous species of which this group is composed approach very closely to the Deer; there is, however, considerable variety in their mode of life. They inhabit almost every description of country; the sandy desert, the open plain, the thicket, the forest, the mountain, and the precipice, being, each in its turn, the favourite haunt of the different races; but, with the exception of a few species, they do not advance much beyond the limits of the tropics. The smaller ones usually prefer a solitary life, but the larger, for the most part, congregate together in herds, which are generally few in number. In their manners they exhibit much of that cautious vigilance and easily startled timidity, combined with a certain degree of occasional boldness and not a little curiosity, which are the natural consequences of their wild and unrestricted habits, of their trivial means of defence against the numerous enemies to whose attacks they are exposed, and of the unequalled fleetness of their speed. In some this latter quality consists of a continued and uniform gallop, which in others is interrupted at every third or fourth stroke by a long and generally a lofty bound, producing a beautiful effect by its constant and rapid recurrence.

The Indian Antelope, of which the specimen in the Tower constitutes a remarkable and highly interesting variety, is not only one of the most beautiful, but also the most celebrated species of the group. It occupies the place of Capricorn in the Indian Zodiac, and is consecrated to the service of Chandra or the Moon. In size and form it closely resembles the Gazelle of the Arabs, the well known emblem of maiden beauty, typified, according to the poets, in the elastic lightness of its bound, the graceful symmetry of its figure, and the soft lustre of its full and hazel eye. From this truly elegant creature our Antelope is, however, essentially distinguished by several striking characters. Its horns, which are peculiar to the male, are spirally twisted, and form, when fully grown, three complete turns; they are closely approximated to each other at the base, but diverge considerably as they proceed upwards. They occasionally attain a length of nearly two feet, and are surrounded throughout by elevated and close-set rings. The two horns taken together have frequently been compared to the branches of a double lyre. The extremity of the nose is bare, forming a small and moist muzzle; the sub-orbital openings are larger and more distinct than in almost any other species; and the ears are pointed and of moderate size. The natural colours vary with the age of the animal, but correspond in general pretty closely with those of the common deer. They may be shortly described as fawn above and whitish beneath, becoming deeper with age, and lighter in the females than in the males. The occasional stripes of a lighter or darker colour, which are generally visible on various parts of the body, can scarcely be considered as occurring with sufficient regularity to allow of their being described as characteristic of the species.

But for these shades of colour, or for any other, we should look in vain in the animal of the Tower Menagerie, which, in consequence of a particular conformation, not unfrequent in some species of animals, and occasionally met with even in the human race, is perfectly and purely white. In order to explain this phenomenon, which is one of the most curious, but at the same time one of the most simple in physiology, it is necessary to observe that there exists beneath the epidermis, or outer covering of the skin, both in man and animals, a peculiar membrane of very fine and delicate texture, which is scarcely visible in the European but sufficiently obvious in the Negro, termed by anatomists the rete mucosum. In this net-work is secreted, from the extremities of the minute vessels which terminate upon its surface, a mucous substance which varies in colour according to the complexion of the individual, of the varieties in which it is the immediate cause; and from the substance thus secreted the colouring matter of the hairs and of the iris is derived. The pure whiteness then of the covering of the animal in question, and of all those which exhibit a similar variation from their natural tinge, is attributable solely to the absence of this secretion from whatever cause. It is always accompanied, as in the present instance, by a redness of the eyes, arising from the blood-vessels of the iris being exposed to view in consequence of the want of the usual coating formed by this secretion, by which they are naturally protected from the too great influence of the light. In the human race the individuals who are thus afflicted, characterized by the dull whiteness of their skins, the deep redness of their eyes, and their colourless, or, as it is generally termed, flaxen, hair, are called Albinos. They are generally timid in disposition, languid in character, and weak both in mind and body. The same original conformation, for it is always born with the individual and never acquired in after life, although sometimes prolonged beyond its limits in the shape of an hereditary legacy, is common to many animals. Perhaps the most familiar instances among these are the white mice, the white rabbits, and the white pigeons, which are known to every one. But it has also been occasionally seen in many other species, as monkeys, squirrels, moles, pigs, and even cows and horses, and, to come a little closer to our present subject, in goats and deer. Not even that massive and stupendous beast the Elephant is exempted from its influence. It can hardly be necessary to recall to the reader the title on which the ruler of millions of not uncivilized Asiatics, the Burmese monarch, prides himself more than on any other, inasmuch as it is the emblem of power and prosperity, that of Lord of the White Elephant; a title, which, while it demonstrates the fact of the existence of this deviation in the Elephant as well as in other animals, proves also the extreme rarity of its occurrence. It has moreover been noticed in many species of birds.

The present species of Antelope is spread over the whole of the Peninsula of Hindoostan and a part of Persia; but it is questionable whether it has been found in Africa, as is commonly asserted. They are said to bound with apparent ease over a distance of from twenty-five to thirty feet, and mounting to the height of ten or twelve. It is consequently useless to attempt to chase them in the common mode with hounds; and their pursuit is restricted to the higher nobility, who employ for the purpose either hawks, who pounce upon their quarry and detain it until the dogs can come up, or Chetahs, who attack them by surprise in the manner before described.

The elegant Albino now in the Tower was brought from Bombay by Captain Dalrymple of the Vansittart, and remained for a considerable time at Sand Pit Gate, where it was an especial favourite with his Majesty, as well on account of the gentleness of its disposition, as for its rarity and beauty. It bears its confinement in the Menagerie with perfect resignation, and is remarkable for the mildness and tranquillity of its deportment.

THE AFRICAN SHEEP.

_OVIS ARIES._ LINN.--Var. _GUINEENSIS._

In characterizing the present genus, were we to look solely at the animal such as we have it daily before our eyes, the distinction between it and all the other Ruminants is too striking to be for a moment mistaken. But the insensible gradations which connect this familiar denizen of our downs and pastures with the untamed native of the desert and the precipice, and the close affinity which subsists between the latter and the goats, render it almost impossible to isolate them by any satisfactory characters. On the present occasion we shall content ourselves with observing that the sheep may generally be distinguished by the direction of their horns, by the elevation of their profile, and by their want of beard: characters neither essential nor infallible, but the best that can be offered.

The variety figured over leaf is in one of the many intermediate stages between unreclaimed barbarism and complete domestication. It is an awkward looking creature, high on the legs, narrow in the loins, and covered with a rough and shaggy coat. The back and sides are nearly black; the shoulders reddish brown; and the posterior part of the body, the haunches, the hind legs, and the tail, white; as are also the ears, which are rather large, the nose, and a spot over each eye. The horns, although the specimen is a male, are remarkably small, and enclose the ears within their curve. If the ears are freed from their confinement, the animal becomes very uneasy, and never rests until he has succeeded in replacing them, which he cannot accomplish without considerable difficulty. He was presented to the Menagerie by Lord Liverpool about six years ago, and is extremely mild in his temper.

THE GREAT SEA-EAGLE.

_HALIAETOS OSSIFRAGUS._ SAV.

THE GOLDEN EAGLE.

_AQUILA CHRYSAETOS._ SAV.