CHAPTER XV.

_THE ANTELOPES._

BY F. C. SELOUS.

The TRUE ANTELOPES (including the Gazelles) are strictly confined to the Old World, the Prongbuck of North America differing so much from all other living ruminants, in its horn growth and other particulars, that it is considered to be the sole representative of a distinct family.

THE HARTEBEESTS.

With the exception of one species--the Bubal--which is found both in North Africa and Arabia, the HARTEBEESTS are entirely confined to the African Continent. They are animals of large size, standing from 43 to 48 inches at the shoulder, and are characterised by their long, narrow faces, high withers, and doubly curved horns, which are present in both sexes. Nine different species of this group are known to exist.

Although the ranges of these various species of hartebeest cover the greater part of the African Continent, it is noteworthy that each species keeps to its own ground, their several ranges but rarely overlapping.

All the hartebeests have a strong family resemblance, and are very similar in their habits. They are never found either in dense forests or in swampy or mountainous country, but are inhabitants of the arid deserts of Northern and South-western Africa, and of the open grassy plains and thinly forested regions of the high plateaux of the interior of that continent. They are extraordinarily fleet and enduring, and in my own experience I have never heard of one of these animals, of whatever species, having been overtaken or ridden to a standstill by a man on horseback. They are very inquisitive, and where they have not been molested will allow any unaccustomed object--such as a European in clothes--to walk to within easy shot of them before running off. They soon gain experience, however; and in countries where they have been most persecuted hartebeests are the keenest-sighted and the most wary of all African game. They are very fond of climbing to the top of the large ant-heaps with which the plains of Africa are profusely studded, and from this point of vantage surveying the surrounding country. They live, I believe, entirely upon grass, and in the desert areas of their range seem able to subsist for long periods without drinking water. Their meat I have always thought very palatable. They are generally in fairly good condition, though they seldom carry much fat. Their fat, after being melted, becomes solid again immediately on cooling, and clogs on the teeth whilst being eaten. But very few African species, except the eland, ever become really fat; their life is too active, and the food-supply too uncertain, for them to put on flesh like European deer.

BONTEBOK AND BLESBOK GROUP.

Nearly allied to the hartebeests are certain other antelopes, of which it will be sufficient to mention but two species--_viz._ the BONTEBOK and the BLESBOK. These two antelopes, though doubtless distinct, since their points of difference are constant and unvarying, are nevertheless so much alike, and evidently so closely allied, that I look upon the former as a highly coloured and specialised race of the latter. The blesbok once had a far wider range than the bontebok, and ran in countless herds on the plains of the northern districts of the Cape Colony, the Orange River Colony, the Transvaal, Griqualand West, and British Bechuanaland, whilst the latter animal has always been confined to the sandy wastes in the neighbourhood of Cape Agulhas, the extreme southern point of Africa.

I think it, however, not improbable that ages ago the blesbok ranged right through Cape Colony to the sea-shore, and that subsequently the gradual desiccation of the south-western portions of the country--which is still continuing--or several years of continuous drought, caused the withdrawal of the species northwards from the waterless parts of the country. Those, however, which had reached the neighbourhood of Cape Agulhas, where there is plenty of water, would have remained behind and formed an isolated race, which, being influenced by local conditions, would naturally in course of time have become differentiated from the parent stock. Be this as it may, the bontebok of to-day is nothing but a glorified blesbok, being slightly larger and more richly coloured than the latter animal. Its horns, too, are always black, whilst those of the blesbok are of a greenish hue. When they are in good condition, the coats of both these species of antelope, as well as of the Sassaby, another member of this group, show a beautiful satiny sheen, which plays over their purple-brown hides like shadows on sunlit water.

The few bonteboks which still survive are now all preserved on large enclosed farms; but their numbers are very small--less than 300, it is believed. The farmers of Dutch descent now do their best to preserve rare species on their land.

THE GNUS.

These remarkable animals were once distributed throughout the greater part of Africa from the Cape to Abyssinia, and their range is even now very extensive, though what was once the most numerous and the most eccentric-looking species of the group has almost ceased to exist.

The gnus are of large size, and at first sight appear to have the head of a buffalo, the tail of a horse, and the limbs and hoofs of an antelope. Their heads are very massive, with broad muzzles and widely separated, hairy nostrils; their necks are maned, tails long and bushy, and both sexes carry horns. They are known as "wilde beeste," or "wild cattle," to the Dutch colonists of South Africa.

The WHITE-TAILED GNU, or BLACK WILDEBEEST, as it is more commonly called, was once found in great numbers on the karroos of Northern Cape Colony, and throughout the vast plains of the Orange River Colony, Transvaal, Griqualand West, and British Bechuanaland. Its range, in fact, was coequal with that of the blesbok. Even as lately as in 1875 and 1876 I personally saw very considerable herds of these quaint animals in the Orange River Colony and the Western Transvaal. When the present war broke out in 1899, there were only two herds of black wildebeest left alive. These animals numbered some 500 head altogether, and were protected by Dutch farmers. There are probably very few of them left to-day, and it is scarcely possible that this most interesting animal will long escape complete extinction.

Black wildebeests, before they had been much persecuted, were so inquisitive that, in the words of Gordon Cumming, they would "caper and gambol" round a hunter's waggon or any other unusual object, and sometimes approach to within a couple of hundred yards, when, whisking their long white tails, they would gallop off with loud snorts. They were always, however, very keen-sighted, and soon became extremely wary and almost impossible to approach on foot in the open plains they frequented, whilst their powers of endurance and fleetness of foot were such that they could only be overtaken by a well-mounted hunter. In spite of these advantages, however, the value of their skins, and the ever-increasing number of hunters, armed with long-range rifles, practically brought about the extermination of this species of gnu in a few decades.

The BRINDLED GNU is a larger animal than the last-named species, standing 4½ feet and upwards at the shoulder. This animal once ranged from the Vaal River northwards, throughout Eastern and Central Africa, to the north of Kilimanjaro, where its range overlaps that of a closely allied form, the WHITE-BEARDED GNU, which is only found in certain districts of Eastern Africa. In general habits these two varieties seem to be identical.

In the interior of Southern Africa, both north and south of the Zambesi, I have met with very large numbers of BLUE WILDEBEESTS. They usually run in herds of from ten to twenty individuals, but towards the end of the dry season collect in droves of 200 or 300. They are often found in company with zebras and sassaby antelopes. Their flesh resembles coarse beef, and, to my thinking, is not ill-flavoured.

THE SMALLER BUCKS.

In addition to the great number of antelopes of large size which inhabit the African Continent, there are also very many small species, the life history and habits of some of which are as yet but imperfectly known, since they are denizens of dense forests, and feed principally at night.

All these small African antelopes are divided into two sub-families. The first comprises the African DUIKERS and the Indian FOUR-HORNED ANTELOPE, and the second the DIK-DIKS, ORIBIS, KLIPSPRINGER, and certain other small bush-antelopes.

The African duikers are distributed throughout Africa south of the Sahara, and are represented by some twenty different species, the largest of which approaches a small donkey in size, whilst the smallest is not much larger than a hare.

The majority of these dainty little antelopes are inhabitants of the dense tangled forests of the coast-belts of Africa, and are therefore but seldom seen by travellers and sportsmen. One species of the group, however, the COMMON DUIKER of South Africa, is a very well-known animal. This little antelope inhabits much more open country than most of its congeners, and has an enormous range, extending from Cape Agulhas to Somaliland, whilst two very nearly allied forms are found in Senegal and Abyssinia respectively.

In most species of duikers both sexes are horned, but in the case of the common duiker it is very exceptional to find a female with horns, and in all my experience I have only known of three such cases.

The FOUR-HORNED ANTELOPE is the Indian representative of the African duikers, and is found along the foot of the Himalaya from the Punjab to Nepal, and in suitable localities throughout the peninsula of India. It frequents wooded hills, but avoids dense jungle. Like its nearest allies, the duikers, it is solitary in its habits, more than two of these antelopes seldom being seen together. The growth of four horns on the skull of this antelope and on certain breeds of domesticated sheep is a curious fact which has not roused as much comment as it deserves.

THE KLIPSPRINGER.

Turning to the second sub-family, we may select the KLIPSPRINGER as the most characteristic species to describe. This beautiful little animal, which is often called the African Chamois, is found in suitable localities from the Cape to Abyssinia. In the southern and northern portions of its range the klipspringer is an inhabitant only of rugged mountain-ranges, and ascends to a height of 9,000 or 10,000 feet above sea-level. In the more central regions of its habitat, however, although it always lives amongst rocks, and thoroughly justifies its name of "rock-jumper," it is often found in regions where there are no high mountain-ranges. It used to be very common in Matabililand, both in the Matopo Hills and on the isolated granite kopjes which are so numerous in that country, and usually are not more than 200 or 300 feet in height. In Mashonaland I have found it living amongst granite rocks in the beds of the larger rivers, and actually on the same level as the surrounding country; whereas on Wedza, a great mountain-mass of slate and ironstone, which rises to a height of about 2,000 feet above the surrounding country, and to the top of which I once climbed, I did not see any klipspringers. The hoofs of this little animal are curiously different from those of any other African antelope, being remarkably short and small, with very deep hollows. This adaptation to its requirements enables the klipspringer to obtain a foothold on any small projecting piece of rock, and to climb in a series of little jumps up the faces of cliffs which seem almost perpendicular.

In height the klipspringer stands about 1 foot 9 inches at the shoulder. The males alone carry horns, which are straight and ringed at the base, and vary from 3 to 5 inches in length. The coat is of a greeny yellow-brown colour, with the hairs hollow and brittle. These little animals are usually met with singly, or in twos and threes together. When caught young, they become wonderfully tame, and make the most charming pets, being very playful and fond of jumping, with surprising ease and grace, from the floor of a room on to any elevated position, such as a table, mantelpiece, or window-sill.

THE WATERBUCKS.

The largest animals in the first of three groups now to be considered are the WATERBUCKS, antelopes of stout and sturdy build, standing from 45 to 50 inches at the shoulder, and covered with long, coarse hair, especially on the neck, in both sexes. The males alone carry horns, which vary from 20 to 36 inches in length, and are strongly ringed in front for three-fourths of their length. They are sublyrate in shape, being first inclined backwards and then forwards at the tips. There are three well-marked species of waterbuck--_viz._ the COMMON WATERBUCK of South Africa, whose range extends from the Limpopo northwards, through Nyasaland to German and British East Africa, and to the Shebeyli River, in Somaliland; the SING-SING of Senegal and Gambia; and the DEFASSA WATERBUCK of Western Abyssinia and the Nile Valley, south to Uganda and British and German East Africa. In habits all species of waterbuck are very similar. They live generally, though not invariably, in herds of from ten to twenty individuals, and in such small herds there is seldom more than one full-grown male present. In the interior of South Africa the waterbuck is often met with amongst steep stony hills and at a distance of more than a mile from the nearest river. Speaking generally, however, this antelope may be said to frequent the near neighbourhood of water, but to prefer dry to swampy ground. When chased by dogs it always makes for water, and will plunge fearlessly into broad, deep rivers, regardless of crocodiles, to which ravenous reptiles it sometimes falls a victim. In South Africa waterbuck vary much in colour even in the same district, some being reddish brown, whilst others are of a very dark grey. The flesh of the waterbuck is coarse, and sometimes rather strongly tasted, and when in good condition the fat is very hard.

The REEDBUCKS are similar in essential characters to the waterbucks, but are of smaller size, and have more bushy tails, and naked spots on the sides of the head beneath the ears.

Of this group the COMMON REEDBUCK of South Africa is the best known. This animal stands 3 feet at the withers, and is of a soft greyish fawn-colour, with a large fluffy tail, which is always thrown up when the animal runs, exposing the white under-surface. The males alone carry horns, which curve backwards and then forwards, and attain a length of from 12 to 16 inches. Reedbucks are met with singly or in twos and threes, and never congregate in herds, though I have seen us many as eight, belonging probably to three or four families, feeding in close proximity to one another on young green grass.

Another member of the reedbuck group is the ROOI RHEBUCK of South Africa. This latter species, though a much smaller animal, is very similar to the common reedbuck in colour, shape, and general appearance; it is quite distinct in its habits and mode of life, as it lives in small herds of from four or five to fifteen head, amongst rugged stony hills, often far from water.

THE BLACKBUCK OF INDIA.

This handsome species is found throughout India wherever there are open cultivated plains. The male stands about 32 inches at the shoulder, and when full grown is of a glossy black colour, with the exception of a chestnut-coloured patch at the back of the neck, and some markings of the same colour about the face. The belly and insides of the limbs are pure white, the line between the black and white being very clearly defined. The whole body and frame are very compact, strong, and beautifully proportioned, and the head is carried high. The males alone carry horns, which are spiral in shape, annulated almost to the tips, and vary in length from 18 to 28 inches. Young bucks and does are fawn-coloured instead of black. These antelopes are usually met with in considerable herds on open plains in which cultivated tracts alternate with waste land, and they often do much damage to the natives' crops. When alarmed, they first execute a series of prodigious bounds into the air before finally settling down to a steady run. They are surprisingly fleet, and can seldom be overtaken by the fastest greyhounds, although they can be caught and pulled down without difficulty by trained cheetas, or, as they are often called, hunting-leopards.

The PALLA, which is found in Southern and Eastern Africa from Bechuanaland to Kordofan, is one of the most graceful of animals. It is a forest-loving species, and is never found far from water. Both sexes are of a general bright reddish brown, with white bellies. The males alone carry horns, which are very graceful in shape, and vary from 14 to upwards of 20 inches in length. The finest specimens of the palla are met with in the extreme southerly and most northerly portions of its range, the animals inhabiting the intermediate districts being smaller and carrying shorter horns. Pallas are gregarious, living in herds of from twenty to over one hundred. When alarmed, they bound over bushes or any other obstacles with the utmost ease and grace, and appear to get over the ground at a high rate of speed. They are, however, very commonly run down and torn to pieces by wild dogs, which hunt in packs, and are very destructive to African game.

Of far less graceful appearance than the two preceding species is the SAIGA, which, though structurally closely allied to the gazelles, has been placed by naturalists in a genus by itself.

This curious-looking animal, which is chiefly remarkable for its large swollen-looking nose and light-coloured horns, is an inhabitant of the steppes of South-eastern Europe and Western Asia. In height it stands about 30 inches at the withers, and is of a dull yellowish colour in summer, turning to nearly white in winter. The males alone carry horns, which are sometimes 13 or 14 inches long, and of a peculiar colour which has been likened to pale amber.

At the present day the saiga is only found in Europe on the plains between the Don and the Volga, but to the east of the Ural River its range extends over the Kirghiz Steppes and the high plains of all Western Siberia. Living in open country, and having the senses of hearing, sight, and scent all highly developed, the saiga is a difficult animal to approach, and can only be successfully stalked by an expert hunter. In summer it is usually met with in small, scattered bands, which, when driven southwards by snow and cold, are collected into considerable herds in the more southerly portions of its range. In very severe winters whole herds have been known to perish in snow-drifts, and in such inclement seasons large numbers are also killed by the natives. The flesh of the saiga is said to resemble mutton, and is held in much esteem.

THE GAZELLES.

We now come to the Gazelles, among which are comprised many of the best known and most beautiful of the small or medium-sized antelopes. In the true gazelles both sexes generally carry horns. Indeed, this rule is universal in those of Africa and Arabia; and there are only four species known--all Asiatic--in which the females are hornless: _viz._ the TIBETAN GAZELLE, PREJEVALSKI'S GAZELLE, the MONGOLIAN GAZELLE, and the PERSIAN GAZELLE.

The range of the various species belonging to this large group is very extensive, comprising the whole of Northern and Eastern Africa, Arabia, and Western and Central Asia, as well as Mongolia and India. The gazelles are inhabitants of the open plains and arid desert regions of the Old World, and, although sometimes met with in tracts of country where there is a certain amount of scattered bush or open stunted forest, are never found in any kind of jungle or thick cover.

On the sandy plains of North-western Africa are found the RED-FRONTED GAZELLE of Senegal and Gambia; the little-known MHORR GAZELLE of South-western Morocco; and the DAMA GAZELLE, a species which has been known to naturalists ever since the time of Buffon. A near ally of the last-named animal is the RED-NECKED GAZELLE of Dongola and Senaar. In North-eastern Africa are found the large and handsome SOEMMERRING'S GAZELLE; the ISABELLA GAZELLE, of the coastlands of the Red Sea; HEUGLIN'S GAZELLE; PELZELN'S GAZELLE, of the maritime plains of Northern Somaliland; and SPEKE'S GAZELLE, of the interior of the same country; whilst, farther south the group is represented by the large and beautiful GRANT'S GAZELLE, with its allies PETERS'S GAZELLE and THOMSON'S GAZELLE. The well-known DORCAS GAZELLE is an inhabitant of Morocco and Algeria, ranging through Egypt into Palestine and Syria; the MARICA GAZELLE, the MUSCAT GAZELLE, and the ARABIAN GAZELLE inhabit the deserts of Arabia; the EDMI GAZELLE is found in the mountain-ranges of Morocco, Algeria, and Tunis; while LODER'S GAZELLE inhabits the sandy tracts of the interior of Algeria and Tunis. In Asia, besides the four species of gazelle already enumerated in which the females are hornless, one other member of the group is met with. This is the INDIAN GAZELLE, a species very closely allied to the Arabian form.

Of the whole genus GRANT'S GAZELLE is the most beautiful. This handsome animal, which was first discovered by the explorers Speke and Grant in 1860, is an inhabitant of Eastern Africa, from the neighbourhood of Lake Rudolph southwards to Ugogo. In size the average height at the shoulder of males of this species is about 34 inches. The coat is close and short and of a general fawn colour, the rump and belly pure white, and the face marked with a rufous band from the horns to the nose and with streaks of white on each side. The upper surface of the tail is white, with a black and tufted tip. The horns, which are very elegant in shape, being first curved slightly forwards and then backwards, are much longer and more powerful than in any other gazelle, and attain a length of 30 inches in the males and 17 inches in the females.

Grant's gazelles, though they undoubtedly find their most congenial home in open country, have also been met with by recent travellers in bush-sprinkled wastes and stony, rugged hills. They are, however, never found in dense jungles or high mountains. They live in herds of from half a dozen to twenty or thirty individuals, though in certain localities as many as 200 have been seen together. They are fond of consorting with other game, such as Burchell's and Grevy's zebras, Coke's hartebeest, and the beisa oryx, and are often met with at long distances from the nearest water. They are keen-sighted and wary, and from the open character of the country in which they are usually encountered are often difficult to stalk. When in good condition, the meat of this gazelle is said to be excellent.

The nearest ally of the true gazelles is undoubtedly the SPRINGBUCK of South Africa. Owing to the protection which it has received of late years, this graceful antelope is now a common animal in many parts of South Africa, and in the north-western portions of the Cape Colony still sometimes collects into prodigious herds, which travel through the country in dense masses, destroying every vestige of grass on the line of their advance, and causing considerable anxiety to farmers, whose flocks of sheep and goats are sometimes swept away by the migrating springbucks. In former years the migration of these antelopes in countless thousands from the deserts of Namaqualand to the countries farther south was a common occurrence, an unerring instinct guiding the wandering herds to districts where rain had lately fallen and caused a new growth of green grass. The animals composing these migrating herds were called by the Dutch settlers of the Cape Colony "Trekbokken," or "travelling-bucks."

Two other antelopes, the DIBATAG and the GERENUK, are included in the present group; but both, whilst typically gazelline in certain respects, differ so much in other ways from all members of that group that each has been placed in a separate genus.

The DIBATAG is a very remarkable-looking antelope, only found in certain districts of Central Somaliland, where it was first discovered by Mr. T. W. H. Clarke in 1890. This species shows the face-markings of the gazelles, whilst the horns, which are only present in the males, much resemble in shape those of a reedbuck. They are rather short, attaining a length of only 11 or 12 inches, and their basal halves are strongly ringed in front. The neck of this antelope is singularly long and thin, and the tail, which is held curved forwards over the back when the animal is in motion, is also much elongated, and only tufted at the tip. The dibatag frequents sandy ground sparsely covered with low thorn-bushes, and lives in small families, being usually met with in twos or threes, whilst it is rare to find more than four or five consorting together.

The GERENUK, like the last-named animal, is an East African species, but has a more extended range, being found all over Somaliland, and thence southwards to the Tana Valley and the Kilimanjaro district of British East Africa. The most remarkable external characteristic of this species is the excessively long neck. The males alone carry horns, which attain an average length of 12 or 13 inches, and, though somewhat gazelle-like in shape, are more strongly crooked forwards at the points. The skull of this species is more dense and solid in structure than in the true gazelles, and the cheek-teeth are smaller in size.

Coming now to the Sable Antelope group, we find an assemblage of antelopes which are all of large size and handsome appearance, and in all of which both the males and females are horned. With the single exception of the BEATRIX ORYX, which inhabits Arabia, all these antelopes are denizens of Africa. One species of the group, the BLUEBUCK, which appears to have been entirely confined to the mountainous districts of the Cape Peninsula, became extinct during the first decade of the last century. Little is known as to the life history of this animal, but it was undoubtedly nearly allied to the larger and more handsomely marked ROAN ANTELOPE. This latter animal once had a more extensive range than any other antelope, as it was found in almost every part of Africa south of the Sahara, with the exception of the Congo forest region. It has now been exterminated in the more southerly portions of the country, but from the Limpopo to the Upper Nile, and thence to the Niger, it is still to be found wherever the surroundings are suitable to its requirements.

A large bull roan antelope will stand 4 feet 9 inches at the withers. The general colour of the body differs in individuals, even in the same district, varying from a very light shade of brown to dark grey or red-roan. The front and sides of the face are jet-black in the adult male, and dark reddish brown in the female, with two long white tufts of hair under the eyes. The muzzle and extremity of the lower jaw are white. The hair on the under side of the neck is long and coarse, and a stiff mane about 3 inches in length runs from behind the ears to the withers. The ears are very long, and in the females and young males tufted. The horns are curved backwards, and in the male are very stout and strong, attaining a length of from 26 to 34 inches. In the female the horns are shorter and slighter, and not so strongly ringed.

Roan antelope are usually met with in small herds of from six to a dozen members, and never congregate in large numbers. I do not think I have ever counted as many as thirty together. I have found them fairly common in certain districts, but nowhere very plentiful. They frequent open plains and thinly forested country, and are never found far away from water. Bucks often become savage when wounded, and will sometimes charge viciously if approached incautiously. They can use their horns with great dexterity, and play havoc with a pack of dogs.

The SABLE ANTELOPE, though considerably smaller than the roan, is yet a handsomer animal. In colour the adult male, when in high condition, is jet-black all over with the exception of the white face-markings and the snow-white of the belly and insides of the thighs. The mane is longer and more bushy than in the roan antelope, and often hangs down on either side over the withers. The horns, too, are much finer, and, sweeping backwards in a bold curve, are commonly upwards of 42 inches long, and have been known to reach 50 inches. The striking colour, large size, and horns of this creature make it one of the most-prized trophies of the sportsman. The skin, when prepared and laid down as a rug in halls or dwelling-rooms, is far more handsome than that of any deer. The female of this species is usually of a rich red-brown in colour instead of black as in the male. South of the Zambesi, however, old cows become almost absolutely black. North of the Zambesi both male and female sable antelopes are dark red in colour rather than black. The horns in the female are slighter and less curved than in the male, and are also considerably shorter, as a rule not measuring over 30 inches in length.

The range of the sable antelope extends from the northern districts of the Transvaal to German East Africa. In the country between the Limpopo and the Central Zambesi it used to be a very common animal, especially in the northern districts of Mashonaland. It is partial to open forests intersected by grassy, well-watered glades, and is never found on open plains entirely devoid of bush. It is usually met with in herds of from twelve to twenty individuals, but I have often seen as many as fifty, and once counted between seventy and eighty together. However large a herd of sable antelopes may be, it is very exceptional to find with it more than one fully adult male, from which fact I should judge that these animals are of a very jealous and pugnacious disposition. When wounded and brought to bay by dogs, a sable antelope defends itself with the utmost fury, using its long scimitar-shaped horns with most wonderful quickness and dexterity. If badly wounded it will lie down, otherwise it fights standing. Keeping its face to some of its foes, with a sideways twist of its head it will transfix and throw into the air any dog which attempts to attack it from behind. I have seen a wounded sable antelope, when lying down, drive one of its horns clean through a large dog deep into its own haunch, and I have had four valuable hounds killed and four others grievously wounded by one of these animals in less than a minute. I once knew a native hunter who was stabbed through the kidneys and killed by a sable antelope cow.

The nearest allies of the sable and roan antelopes are the various species of the genus Oryx. In this group are included the WHITE ORYX, which inhabits the desert regions of the interior of Northern Africa from Dongola to Senegal; the BEATRIX ORYX of Southern Arabia; the GEMSBUCK of South-western Africa; the BEISA, which is found in North-east Africa from Suakim southwards to the river Tana; and the TUFTED BEISA, which is very nearly related to the last-named species, whose place it takes south of the Tana River in certain districts of British and German East Africa. In general appearance there is a strong family resemblance between the different species of oryx. In all of them both sexes carry horns, which are considerably longer, though somewhat slighter, in the females than in the males. In the white oryx the horns are curved backwards; but in the other four species they are straight, or nearly so. In all the faces are conspicuously banded with black and white, and the tails long, with large dark terminal brushes. The two most desert-loving species, the white and the Beatrix oryx, are paler in general body-colour than the other three, and the latter animal is considerably smaller than any other member of the group, standing not more than 35 inches at the withers. The gemsbuck is the largest and undoubtedly the handsomest of the group, standing 4 feet at the shoulders; the horns of the females are often upwards of 40 inches long, and have been known to attain a length of 48 inches.

In habits all species of oryx seem to be very similar. They are denizens of the arid sun-scorched plains of Africa, which are not necessarily devoid of all kind of vegetation, but are often covered with stunted bush, and carry a plentiful crop of coarse grass after rain. Oryx usually run in herds of from four or five to fifteen or twenty, though the beisa, the most abundant of the group, has been met with in troops numbering 400 or 500 head. All the oryx are shy and wary, and in the open country they usually frequent are difficult to approach on foot. If pursued on horseback, they run at a steady gallop, which they can maintain for long distances, swinging their bushy black tails from side to side, and holding their heads in such a way that their long straight horns are only sloped slightly backwards. Fleet and enduring, however, as oryx undoubtedly are, I am of opinion that in these respects the gemsbuck of South Africa, at any rate, is inferior to all other large antelopes living in the same country, with the single exception of the eland. I have often, when mounted on a fast horse, galloped right up to herds of gemsbuck, and on two occasions have run antelopes of this species to an absolute standstill. Oryx of all species should be approached with caution when badly wounded, as they are liable to make short rushes, and can use their horns with great effect.

Nearly related to the antelopes of the Oryx group in many essential characteristics, yet at once distinguishable by its spiral horns and broad reindeer-like feet, the desert-haunting ADDAX has been placed in a separate genus, of which it is the sole representative.

This remarkable animal stands about 38 inches in height at the withers, and varies in general colour at different seasons of the year, from brownish grey to a reddish hue. The forehead is covered with a thick growth of bushy black hair, beneath which there is a patch of white extending across the nose to under the eyes. The hindquarters, tail, and legs are white. The horns are spiral, and are present in both sexes. In the male they attain a length of about 28 inches in a straight line, and almost 36 inches following the spiral. In the female they are thinner and less spirally curved. The addax is confined to the desert regions of Northern Africa from Dongola to Senegal, and the broad, rounded hoofs, so unlike those of any other antelope, would seem to show that it inhabits countries where the soil is deep, soft sand.

Very little is known of the life history or habits of this antelope. It is said to associate in pairs or small herds, and to be entirely independent of water, though it travels great distances over the desert in the track of thunder-storms for the sake of the young herbage which grows so quickly wherever rain falls in those thirsty regions. It is killed in considerable numbers by the Arabs for the sake of its flesh and hide, and is either stalked or hunted on horseback, with the help of greyhounds, by Europeans.

The last of the sub-families into which modern naturalists have divided the antelopes of the world comprises some of the handsomest species of the whole group, and includes the largest of all antelopes, the Eland, as well as such small and beautifully marked creatures as the Harnessed Bushbucks.

With one exception--the Nilgai--all the members of this sub-family are denizens of the great African Continent.

The NILGAI, or BLUE BULL, is an inhabitant of India, and is found throughout the greater portion of the peninsula, from the base of the Himalaya to the south of Mysore. It is an animal of large size, standing about 4 feet 6 inches at the shoulder. In general colour the male is of a dark iron-grey, the female tawny fawn. White spots on the cheeks and just above the hoofs on the fore and hind feet are the outward signs of its affinity to the African harnessed antelopes. The male alone carries horns, which are nearly straight and very small for the size of the animal, rarely exceeding 9 inches in length.

Passing now to the Harnessed Antelopes of Africa, our attention is first claimed by the BUSHBUCKS. Excluding the Inyala and the Broad-horned Antelope, we find several forms of the smaller bushbucks recognised by naturalists: _viz._ the HARNESSED ANTELOPE of the forest regions of Western Africa; the CAPE BUSHBUCK of South Africa; CUMMING'S BUSHBUCK of Eastern Africa; and the DECULA BUSHBUCK of Abyssinia. The various forms of bushbuck vary in general colour from very dark brown to various shades of grey-brown, yellow-brown, and rich red. In all species the young are more or less striped and spotted; but whereas in some forms the adult animals lose their stripes and spots almost entirely, in others the adults are more richly marked than immature specimens. For my part, I am inclined to believe that, if large series of bushbuck-skins were collected from every district throughout Africa, it would be found that all the varieties of this animal at present accepted as distinct species would be found to grade into one another in such a way that only one true species could be recognised.

The bushbucks vary in height at the shoulder from 28 inches to 33 inches, and only the males carry horns, which are nearly straight, with a close spiral twist, and measure in adult animals from 10 inches to 18 inches in length.

Bushbucks are not found in open country, but live in forest or thick bush near the bank of a river, stream, or lake, and are never met with far from water. They are very partial to wooded ravines amongst broken, mountainous country, provided such districts are well watered; and are very solitary in their habits, both males and females being usually found alone, though the latter are often accompanied by a kid or half-grown animal. They are shy and retiring, and should be looked for between daylight and sunrise, or late in the evening, as they are very nocturnal in their habits, and lie concealed in long grass or thick bush during the heat of the day. Their call resembles the bark of a dog, and may often be heard at nights.

The BROAD-HORNED ANTELOPE is only found in the forests of the West African coast range, from Liberia to Gaboon. The male of this species is a very handsome animal, standing about 43 inches at the withers, and is a bright chestnut-red in general colour, with a white spinal stripe extending from the withers to the root of the tail, and fourteen or fifteen white stripes on the shoulders, flanks, and hindquarters. The ears are large and rounded, and the horns very massive, and about 30 inches in length, measured over the single spiral twist. There are two or three large white spots on the cheeks, and a broad white arrow-shaped mark across the nose below the eyes. The female is similar in coloration to the male, but smaller and hornless.

Little or nothing is known as to the habits of this very beautiful antelope. Du Chaillu, who met with it in the interior of Gaboon between 1856 and 1859, says that it is "very shy, swift of foot, and exceedingly graceful in its motions"; but he does not tell us whether it lives in pairs like the bushbucks, or in small herds like some of its other near allies.

The INYALA is another bush-loving antelope closely allied to the bushbucks. In this species the general colour of the adult male is a deep dark grey, that of the female and young male bright yellow-red, and both sexes are beautifully striped with narrow white bands on the body and haunches. In the male long dark hair hangs from the throat, chest, and each side of the belly, and fringes the front of the thigh almost to the hock, and the back of it up to the root of the tail. The ears are large and rounded; and the horns, which are only present in the male, attain a length of about 2 feet in a straight line, and 30 inches along their spiral curve. The standing height at the shoulder of males of this species is about 42 inches.

This most beautiful antelope has a very restricted range, being only found in a narrow belt of coastland extending from St. Lucia Bay to the Sabi River, in South-east Africa, and in a still smaller area in the neighbourhood of the Upper Shiri River, in British Central Africa.

Before the acquisition of firearms by the natives in South-east Africa, the inyala was very plentiful in Northern Zululand and Amatongaland, and was then to be met with in herds of from ten to twenty individuals; whilst the males, which at certain seasons of the year separated from the females, were in the habit of consorting together in bands of from five to eight. Constant persecution by the natives in Amatongaland and the countries farther north very much reduced the numbers of inyalas in those districts a long time ago; but in Zululand, where this animal has been strictly protected by the British authorities for the last twenty years, it was still plentiful up to 1896, when the rinderpest swept over the country, and committed such sad ravages amongst all the tragelaphine antelopes that it is to be feared the inyala can now no longer be found anywhere in any considerable numbers. Where I met with these antelopes some years ago, in the country to the south of Delagoa Bay, I found them living either alone or in pairs like bushbucks. They frequented dense thickets in the immediate neighbourhood of a river or lagoon, and I never saw one in anything like open country or far away from water. Their tracks showed me that at night they were accustomed to feed in open spaces in the bush, but they always retired to the jungle again at daylight, as they had become very wary and cunning through constant persecution at the hands of the natives.

Closely allied to the bush-antelopes of the present group are the swamp-haunting SITATUNGAS. Three species of these have been described,--one from East Africa, named after Captain Speke; another from tropical West Africa; and a third from Lake Ngami and the Chobi River, named after the present writer.

There is very little difference between the adult males of these three species, except that in the West African form the coat is of a darker colour than in the other two. The main difference consists in the fact that, whereas the female of Selous' sitatunga is light brown in colour like the male, and the newly born young are very dark blackish brown (the colour of a mole), beautifully striped and spotted with pale yellow, the female and young of the other two forms are red in ground-colour, with white spots and stripes. However, personally I am of opinion that there is only one true species of sitatunga in all Africa, and that the differences between the various forms are superficial, and would be found to grade one into the other, if a sufficiently large series of skins of all ages and both sexes could be gathered together from all parts of the continent. In the Barotse Valley, on the Upper Zambesi, my friend Major R. T. Coryndon informs me that both red and brown female sitatungas are met with. On the Lower Chobi and Lake Ngami region the females are never red, but always of the same brown colour as the males, whilst on the Congo all the females are red.

The male sitatunga stands about 3 feet 6 inches at the shoulder, and varies in general colour in different localities from light to dark brown. The adult females are either red with a few faint stripes and spots, or light brown, only retaining very faint traces of any stripes or spots. The young are, both in tropical West and Central East Africa, red, striped, and spotted with white; but in South-west Africa dark blackish brown, with spots and stripes of yellowish white. The hoofs are excessively long, and the skin which covers the back of the pastern is hairless, and of a very thick and horny consistency. The males alone carry horns, which are of the same character as in the inyala, but more spiral and longer, having been known to attain a length of 28 inches in a straight line and 35 inches over the curve.

The sitatunga is an inhabitant of the extensive swamps which exist in many parts of the interior of Africa. It may be said to live in the water, as it passes its life in flooded beds of reeds and papyrus, into the muddy bottoms of which its long hoofs, when splayed out, prevent it from sinking. When forced out into dry ground by heavy floods, the formation of its feet so hinders it in running that it can be overtaken and speared by a native on foot. I was informed by the natives on the Chobi River that, when the floods enabled them to paddle their canoes through the reed-beds, they often killed considerable numbers of the sitatungas. These animals, they said, when they saw a canoe approaching, would often not attempt to seek safety by flight, but would sink down in the water, submerging their whole bodies, and leaving only their nostrils above the surface, and in this position were easily speared.

The sitatunga is not gregarious, but is met with singly or in pairs. The hair is long, but soft and silky; and the skins are much sought after by the natives for blankets.

In addition to the bushbucks and sitatungas, two more very notable spiral-horned African antelopes remain to be mentioned--namely, the GREATER KUDU and the LESSER KUDU.

The GREATER KUDU is one of the most magnificent-looking of the whole family of antelopes, and is an animal of large size, an adult male standing 4 feet 9 inches and upwards at the withers. The general colour of this species is light brown to dark grey, the old males looking much darker than females or younger animals, because the scantiness of their coats shows the dark colour of the skin beneath. On each side of the body and hind-quarters there are several white stripes, which vary in number from four to eight or nine. As in all this group of antelopes, there are two or three cheek-spots, as well as an arrow-shaped white mark across the nose, below the eyes. In the male there is a slight mane on the back of the neck, and a fringe of long white and blackish-brown hair intermixed, extending from the throat to the chest. The ears are very large and rounded, and the male is adorned with magnificent spiral horns, which have been known to attain a length of 48 inches in a straight line from base to tip, and 64 inches over the curve.

The greater kudu once had a very wide range, which extended from the central portions of the Cape Colony to Angola on the west, and on the east throughout East Africa up to Abyssinia; but, with the single exception of the buffalo, no species of wild animal suffered more from the terrible scourge of rinderpest which recently swept over the continent than this lordly antelope, and it has almost ceased to exist in many districts of South and South Central Africa, where up to 1896 it was still very numerous.

The greater kudu is a bush-loving antelope, and very partial to wooded hills, though it is also plentiful in the neighbourhood of rivers which flow through level tracts of country covered with forest and bush. In my own experience it is never found at any great distance from water. It eats leaves and wild fruits as well as grass, and lives in small herds or families, never, I believe, congregating in large numbers. In Southern Africa, at any rate, it was always exceptional to see more than twenty greater kudus together, and I have never seen more than thirty. At certain seasons of the year the males leave the females, and live alone or several together. I once saw nine magnificently horned kudus standing on the bank of the Chobi, and I have often seen four or five males of this species consorting together. As a rule the greater kudu is met with in hilly country or in bush so dense that a horse cannot gallop through it at full speed; but if met with in open ground, a good horse can overtake an old male without much difficulty. The females are much lighter and faster, and cannot be overtaken in any kind of ground.

The greater kudu is one of the most timid and inoffensive of animals, and when attacked by dogs will not make the slightest attempt to defend itself either with its horns or by kicking.

The LESSER KUDU in general colour nearly resembles its larger relative, but is much smaller, the males only standing about 40 inches at the withers, and it lacks the long fringe of hair under the throat. The white stripes on the body and hindquarters are, however, more numerous--from eleven to fourteen; and the horns, which are only present in the males, are less divergent, and with the spiral curvature much closer than in the greater kudu.

The lesser kudu is an inhabitant of Somaliland and the maritime districts of British East, Africa. It frequents thick scrubby jungle, and is said to be exceedingly watchful and wary. It lives either in pairs or in small families, but never congregates in large herds. Like all the tragelaphine antelopes, this species is a leaf-eater, and feeds principally during the night, lying up in thick bush during the heat of the day.

There remains to be mentioned but one other group of antelopes, the ELANDS, large, heavily built animals, which belong to the present group, but differ from all species of kudu, sitatunga, and bushbuck, inasmuch as both sexes are horned. There are two forms of the COMMON ELAND--namely, the grey variety of South-western Africa, and the striped animal, which is found in the countries farther north and east. The two forms grade one into the other, and are absolutely identical in their habits and mode of life, the differences between them being merely superficial. To the south of the twenty-third parallel of south latitude all elands are of a uniform fawn colour, except the old animals, which look dark grey, from the fact that the scantiness of their coats allows the dark colour of the skin to show through the hair. Old males, when standing in the shade of a tree, appear to be of a deep blue-grey in colour, and are known to the colonists of South Africa as "blue bulls." In Rhodesia, South-east Africa, and the countries to the north of the Zambesi, all the elands are bright chestnut-red when young, with a black line down the centre of the back from the withers to the tail, broad black patches on the backs of the fore legs above the knees, and eight or nine white stripes on each side. When they grow old, the ruddiness of the ground-colour gradually fades, the black markings on the fore legs die out, and the white stripes become indistinguishable at a short distance, the old bulls looking deep blue-grey in general colour. Every intermediate stage of colouring between the unstriped and the highly coloured forms of eland is to be found in the district lying between the central portions of the Kalahari Desert and the Zambesi River. Old male elands south of the Zambesi develop a growth of long, bristly black hair on the forehead, which often hangs over their eyes and extends half-way down their noses. North of the Zambesi this growth of hair is not nearly so luxuriant.

I have carefully measured the standing height at the withers of many old male elands in the interior of South Africa, and found that it varied from 5 feet 8 inches to 5 feet 10 inches. The horns of bulls in their prime measure from 26 inches to 33 inches in length, but old bulls wear their horns down very much. The cows carry longer, though thinner horns than the bulls.

The range of the eland once extended from Cape Agulhas to the White Nile, but it has become extinct in many districts of Southern Africa, and in almost every other portion of its range has, like all other tragelaphine antelopes, suffered so cruelly from the recent visitation of rinderpest that it has now become a scarce animal all over Africa.

During the rainy season elands are usually met with in small herds of from four or five to ten individuals; but towards the end of the dry season they collect into large herds, and at such times I have often seen from fifty to over two hundred of these animals in one troop.

In my experience elands live for two-thirds of the year in forest or bush-covered country, or amongst rugged hills; and in such localities they are difficult to overtake on horseback; but in the middle of the dry season, as soon as they smell the smoke of the grass fires lighted by the natives on the open plateaux, they leave their retreats, and, collecting in herds, wander out on to the treeless plains in search of young grass. They then fall an easy prey to a mounted hunter, especially the heavy old bulls, which can be run to a standstill with ease by a very moderate horse.

The flesh of the eland is excellent when the animal is in good condition, as at such a time these animals become very fat, especially the old bulls, whose hearts become encased in a mass of fat which will often weigh 20 lbs. It is a mistake, however, to think that eland-meat is always good; for towards the end of the dry season, when there is little grass to be got, they feed extensively on the leaves of certain bushes, and their meat at such times becomes very poor and tasteless.

Besides the common eland of Southern, Central, and Eastern Africa, another distinct species is met with in Senegal and the Gambia Colony. This is the DERBIAN ELAND, about which animal our knowledge is still very slight, as I believe that it has never yet been shot nor its habits studied by a European traveller. A good many skulls and horns and a few skins have been obtained from natives, from which it appears that in general colour this species is of a rich reddish-fawn colour, becoming nearly white below, the middle of the belly being black. The neck is covered with long hair of a dark brown or black colour, blacker towards the shoulder than in front. A broad black stripe extends all down the centre of the back from the neck to the root of the tail, and there are large black patches on the backs and inner sides of the fore legs above the knees. On each side of the body and haunches there are thirteen or fourteen narrow white stripes. The horns are larger and more massive and divergent than in the common eland.

The Derbian eland is said to be a forest-loving animal, never of its own accord coming out into the plains. It lives in small herds, is very shy and not at all abundant, and browses on the leaves and young shoots of various trees and bushes.

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