The Book of the Fly A nature study of the house-fly and its kin, the fly plague and a cure

CHAPTER XIII

Chapter 138,731 wordsPublic domain

A CAMPAIGN OF EFFECTIVE WARFARE

Several authors of recent books, and lately also able lecturers, have done much to awaken people to a realisation of the dangers of our ever recurrent summer plague of flies. The advent of the petrol motor-car and other automobile vehicles has at the most but very slightly improved the state of affairs within town areas, where mews were formerly much more numerous. The public press has followed suit, but something more in the way of a sustained effort for hygienic reform is desirable. The terrible European war should not preclude consideration of the subject, for the scourges of fly-borne contagion have ever followed armies and rivalled the casualties of the very battlefield. Bands of enthusiasts everywhere should keep going a veritable anti-fly campaign as one of the most urgent needs of practical sanitation. Otherwise active support of the cause will soon languish and be obliterated amongst the multitudinous ever-changing questions of the day, political and other, which, as newspaper editors are persuaded, have the attention of the public for the time being. In spite of the incontestible prospects of universal benefit it may not be easy to engage a large body of public support without something like an organised propagandist movement.

If any readers of this booklet are disposed to join and form a central body with a view of ultimately founding an association for promoting the work of fly extermination, the writer will be glad to find or meet with an honorary secretary and helpers who will work in the cause and economise in the necessary expenditure of all contributions received. After the preliminary efforts of starting such an association, its work will be not only to urge the local sanitary authorities everywhere to adopt the best possible course of action, but also to incessantly move public opinion to compel Parliament to pass laws, capable of administration, for the public welfare in this matter.

The present booklet had its origin very many years ago in the author's idea of writing an account of the house-fly and its kindred, which would be interesting and more truthful than much then to be found in current literature. Such off-hand inconsiderate writing, as appears in the "Elements of Entomology," by W. S. Dallas, F.L.S., requires to be controverted; therein it is stated that the house-fly, which is "troublesome, does very little actual damage, for our only real grounds of complaint are to be summed up in the tickling sensation which its feet cause," &c. "In its larvæ state, however, it lives inoffensively enough in dung." It has now seemed timely to publish my long-delayed work, re-written with the object of more urgently interesting the general public in the cause of the anti-fly campaign. Still, the author trusts that both the deeper and the less entomologically inclined nature students will find therein not only useful, but also some novel information, given with not too much entomological technicality.

There is no English work sufficiently modern and comprehensive for a study of our native flies. In 1776, Moses Harris, who originated or elaborated the study of wing patterns, published his "Exposition of English Insects," in which more than 300 flies are figured and described; they have the old Linnæan classification and nomenclature, of course, and the work is scarce. All later attempts by English authors in the way of a more comprehensive student's guide book have been left incomplete. Another excellent, but expensive work, Curtis's "Genera of British Insects," contains about 250 illustrations and descriptions of flies; but most of these are rather rarities, and the amateur in search of a facile guide to the commoner objects of the country-side will be apt to be disappointed. For the sake of readers possibly eager of advancing further in the study, and in the absence of any commendable guide book, a short appendix has been added to the present work, for help in identifying more numerous species and those of many families and genera not mentioned in the foregoing pages. With the leave of the Northumberland, Durham, and Newcastle-upon-Tyne Natural History Society some valuable plates of illustrations are herewith reprinted, and explanatory notes are added, mainly from the volume of the Society's transactions for 1906, a most valuable work and compilation by the late Rev. W. J. Wingate, of Bishop Auckland. This learned entomologist has succeeded in giving a marvellously comprehensive amount of clear condensed guidance. It is a great privilege that the present booklet has been allowed to borrow from such a source of knowledge, valuable far beyond the locality of its authorship.

Other illustrations which have been borrowed appear with the leave of His Majesty's Office of Works, out of Reports to the Local Government on Public Health and Medical Subjects.

APPENDIX

INDEX TO TERMS AND SYMBOLS OF THE WINGATE FLY CHART, _PLATE_ I.

This Index, together with the following "Table of Wing-cells and Veins," the "Glossary," and _plates_, II, III, IV, V, VI and VII, will explain the theoretical Diagram entitled The Fly Chart, illustrating characteristic features and exterior parts, by the study of which the reader may learn to differentiate all the sixty families which contain species native to Great Britain.

A = the Head, comprising nine regions and parts (A, I to A, IX).

A, I, the Vertex, which contains (1) the Ocellar Triangle; (2) the Ocelli; (3) the Vertical Triangle, a coloured patch surrounding the ocellar triangle; (7) the Vertical Orbit, the sides of the vertex close to the compound eyes. The Bristles hereabouts are (4) the Vertical (inner and outer); (5) the Post-vertical; and (6) the Ocellar.

A, II, the Compound Eyes, large and often hairy.

A, III, the Frons is the forehead between the eyes; it contains (1) the Frontal Stripe; (2) the Frontal Orbits at the sides of the frontal stripe; (3) the Frontal Lunule (_plate_ VII, 5) a crescent immediately above the antennæ. The Bristles hereabouts are (4) one pair of Frontal; a row, or rows of (5) Fronto-orbital; and (6) Lower Fronto-orbital. The (7) Eye Margin is a narrow, often glistening, white line close to the eye.

A, IV, Antennæ, jointed processes springing from between the eyes just below the frons; those with numerous joints are illustrated in _plate_ II, others in _plate_ III. The Frontal Suture in the middle of the face is a little pit in which short antennæ often lie close together. The (1) Basal Joints are the two next the head; (2) the Flagellum, all the joints, when numerous, taken together; (3) the First Joint is next the head; (4) the Second Joint; (5) the third Joint (_plate_ III, 1-9) is sometimes ringed; (6) the Arista, or Seta, is a plume, spine, or bristle springing from the third joint at some point of its upper surface between the base and the tip.

A, V, the Face is the region above the so-called mouth and below the antennæ; the ridges bounding the face on each side are (1) the Facialia; here are (2) the Facial Bristles; (3) the Mystax (_pl._ VI, _fig._ 28) is peculiar.

A, VI, the Cheeks, spaces at the sides between A, II and A, V (I).

A, VII, the Jowls, parts below A, VI and the so-called mouth.

A, VIII, the Mouth, more or less of a cavity into which (3) the Proboscis (the jointed sucking trunk) can be sometimes partly withdrawn; the proboscis has a basal joint called the Rostrum (a snout); the other joint is called the Haustellum (a pump) and has the tip variously adapted as for mere surface suction, or for deep insertion, or for piercing. (1) the Peristome is the region surrounding the mouth; (2) the Vibrissæ are strong bristles close to the sides of the mouth, or A, V (I); (4) the Palpi, two feelers (or perhaps smelling organs) attached to the proboscis.

A, IX, the Back of the Head; here (I) Post-orbital Bristles are sometimes found, that is to say, behind the eyes, A, II.

B = the Thorax, the middle portion of the fly bearing the organs of locomotion.

B, I, the Dorsum. The diagram on _plate_ VI, _fig._ 40 illustrates the upper surface of this region; this part is bounded in front by (_a_) the Neck; behind by (_b_) the Scutellar Suture (15) or Seam; at the sides by (_c_) the Dorso-pleural Sutures (9-9) running from the front corners to the front of the Wing Bases; by (_d_) the Wing Bases (10-10); and by (_e_) the Scutellar Bridges, which are ridges (11-11) running with right and left inclines to the root of (6) the Scutellum. The Dorsum is crossed by (8) the Dorsal Suture, behind which are the Alar Regions (5-5); here on each side is a ridge (12-12) the Alar Frenum; rows of minute bristles on the centre are termed Acrostichal.

B, II, the Sides, or Pleuræ, specified separately in connection with—

B, III, the Breast, or Sternum; (1) the Prothorax is below the shoulder and above the fore-leg; (2) the Meso-pleura is contiguous to the prothorax and the shoulder; here in the upper front corner is the "Stigma," a prominent respiratory orifice, below which the large Stigmatical Bristle may be found; the (3) Ptero-pleura is below the wings and behind the meso-pleura; (4) the Meta-pleura is "a more or less tubercular piece below the region between the root of the wing and the haltere"; (5) the Sterno-pleura is above the middle leg and below the Sterno-pleural Suture, where it bounds the meso-pleura; (6) the Hypo-pleura is behind the sterno-pleura and above the hind leg.

B, IV, the Meta-Notum is the part of the thorax behind and beneath B, I (6), the scutellum.

C = Appendages of the thorax.

C, I, Fore-legs, Mid-legs, and Hind-legs; every leg has nine joints [particularised illustrations on _plate_ VII]; (1) Coxæ are the joints attached to the body; (2) Trochanters very small and inconspicuous; (3) Femora or Thighs; (4) Tibiæ or Shins; (5) First Tarsal Joint or Metatarsus; (6) Second Tarsal Joint; (7) Third; (8) Fourth; (9) Fifth Tarsal Joint; (10) the Knees are where the Thighs and Shins join; (11) Claws, Foot-pads, and Empodium are appendages of the fifth tarsal joint.

C, II, Wings [_see_ Table of wing-cells and veins]; the quarters of the wing are the Fore and the Hind Basal Regions and the Fore and the Hind Apical Regions. The Stigma is a thickened portion of the wing at the middle of the fore margin.

C, III, the Halteres.

C, IV, Squamæ are scale-like appendages beneath the wings and above the halteres; they seem to have some attachment to the roots of the wings; a small inner lobe is called the antisquama, but it is a feature not of much use in classifying and differentiating species.

D = the Abdomen has numerous ringed sections; in the Sub-order, Cyclorrhapha in eight segments, but commonly divided into four or five conspicuous segments and three or four more smaller, less visible, and much modified in the sexual terminal segments.

D, I, the first abdominal segment is the Basal Segment.

D, II, the last of those more visible is the Anal Segment or Anus.

D, III, the abdominal termination, _see plate_ VI, _fig._ 17.

D, IV, the chief segments consist each of an Upper Plate joined to an Under Plate by Membranous Sides, often concealed.

D, V, Abdominal Bristles, _see plate_ VI, _fig._ 41.

E = Bristles, Hairs, etc.; fine short furry hairs are called Pubescence; "Hairs" are longer and stronger; Bristles are still stronger; but a very thick bristle is called a Spine, if on the costa, and a Spur, if on the leg; the larger bristles on B or on D are called Macrochætæ.

F = Stripes and Bands; a long marking, when parallel (or nearly so) to the major axis of the body, leg, or wing, is called a Stripe; when lying at right angles (or nearly so) it is called a Band. For "frontal stripe" _see_ A, III (1).

O = Cells of the wing [_see_ the following Table]; the rib-like nervures are termed Veins; these divide the area of the wing into more or less twelve compartments called Cells or Areolets.

V = Longitudinal Veins [_see_ the following Table]; two groupings within the margin can be recognised; the principal veins of the Fore Region are numbered, 1, 2, and 3, whilst those of the Hind Region are numbered, 4, 5, 6 and 7 respectively; some have branches, _a_, _b_, _c_, etc., when sections (separately referred to) may have index numerals attached.

X = Cross-Veins [_see_ the following Table]; one very significant X-vein termed X, 4 (or the Discal X) in the middle of the wing connects the longitudinal V, 4 to the V, 3, which are thus to be identified even in very abnormal patterns.

The cross-veins and cells are symbolised by having attached to the letters X and O respectively the indicative and qualifying figures and letters belonging to the V upon which they rest, that is to say, the V (or part of V) which is the lower boundary.

TABLE OF WING CELLS AND VEINS

O, 1.—Costal Cell (undivided), or Costal Areolet.

O, 1_a_^1.—The Humeral Cell }

O, 1_a_^2.—Second Costal Cell } Costal Cell, when divided.

O, 1_b_.—Subcostal Cell }

O, 2.—Marginal Cell (resting on V, 2).

O, 3.—Submarginal Cell (resting on V, 3).

O, 4^1.—Cell resting on 4^1 (the 1st part of V, 4), a "basal" cell.

O, 4^2.—Cell resting on 4^2 (the 2nd part of V, 4).

O, 4_b_.—Cell resting on 4_b_ (a lower branch of V, 4).

O, 5^1.—Cell resting on 5^1 (the 1st part of V, 5), a "basal" cell.

O, 5^2.—Cell resting on 5^2 (the 2nd part of V, 5), a "discal" cell.

O, 5^3.—Cell resting on 5^3 (the 3rd part of V, 5), a "marginal" cell.

O, 6.—The Anal Cell.

O, 7.—The Axillary Cell.

O, 8.—The Subaxillary Cell.

O, 9.—The Alula, or Axillary Lobe.

V, _c_ (or C).—The Costal Vein, or Costa, bounding the front Margin.

V, _c_1_a_.—Part of the Costa (often ciliated and spined) reaching to the end of V, 1_a_.

V, 1_a_.—The upper branch of the V, 1.

V, 1_b_.—The lower branch of the V, 1.

V, 2.—Second (longitudinal) Vein.

V, 2_b_.—Lower branch of V, 2.

V, 3.—Third or Cubital Vein.

V, _s_.—The "Vena Spuria," a thickening of the wing (characteristic of Syrphidæ), an imperfect V between V, 3 and V, 4 crossing X, 4.

V, _h_.—A "Hang" V (or Appendix) a more or less irregular incomplete V.

V, 4.—Fourth Vein, often much branching.

V, 5.—Fifth Vein.

V, 6.—The Anal Vein.

V, 7.—The Axillary Vein.

X, 1_a_.—The Humeral Cross-vein, connecting V, 1_a_ to V, _c_.

X, 1_b_.—The Subcostal X-vein, connecting V, 1_b_ to V, 1_a_.

X, 2.—Cross-vein connecting V, 2 to V, 1_b_.

X, 4.—Discal, Middle, or Central X-vein.

X, 5^1.—Lower Cross-vein, connecting V, 5^1 to V, 4.

X, 5^2.—Lower Marginal Cross-vein, connecting V, 5^2 to V, 4 (or to some branch of V, 4).

X, 6.—Anal Cross-vein.

GLOSSARY OF TERMS WITH INDEX REFERENCES

Abdomen D

Acrostichal B, I

Alar frena B, I (12-12)

Alula, or Axillary lobe O, 9

Anal Vein V, 6

Antennæ, or Horns A, IV

Anus D, II

Apical Region C, II

Arista, or Seta [_plate_ III] A, IV (6)

Axillary Vein V, 7

Basal Region C, II

Breast, or Sternum B, III

Bristles, Spines, Hair, etc. E

Calcar, Spur, or Thorn E

Cells of Wings, or Areolets O

Cheeks or Genæ A, VI

Costa V, _c_

Coxæ, or body-joints C, I (1)

Cubital Vein V, 3

Dorsum [_plate_ VI] B, I

Empodium, claws and pads C, I (11)

Eye Margin A, III (7)

Face and Facialia A, V

Femora, or Thighs C, I (3)

Forceps [_plate_ VI, _fig._ 17]

Frons A, III

Frontal Lunule [_plate_ VII, 5] A, III (3)

Frontal Stripe A, III (1)

Frontal Suture A, IV

Halteres C, III

Hang Vein, or Appendix V, _h_

Hypo-pleura B, III (6)

Jowls A, VII

Knees C, I (10)

Meso-pleura B, III (2)

Meta Notum B, IV

Meta-pleura B, III (4)

Meta-tarsus C, I (5)

Mouth A, VIII

Mystax A, V (3)

O = Cell of wing C, II

Ocelli A, I (2)

Ocellar Triangle A, I (1)

Orbits A, I (7) and A, III (2)

Palpi A, VIII (4)

Peristome A, VIII (1)

Pleuræ, or Side regions B, II

Proboscis A, VIII (3)

Prothorax B, III (1)

Ptero-pleura B, III (3)

Scutellum B, I (6)

Scutellar Bridges B, I (11-11)

Shoulders or Humeri [_plate_ VI, 40] B, I (1-1)

Squamæ, or Calyptræ C, IV

Sterno-pleura B, III (5)

Stigma of Pleura B, III (2)

Stigma of Wing C, II

Stripe or Band F

Sutures A, IV, and B, I

Tarsi, or Tarsal (foot) Joints C, I (5-9)

Tibiæ, or Shins C, I (4)

Thorax B

Trochanter C, I (2)

V = longitudinal Veins C, II

Vena Spuria V, _s_

Vertex, or Crown A, I

Vibrissæ, or Mouth Bristles A, VIII (2)

X = Cross-Veins of Wings C, II

ALPHABETIC LIST OF FAMILIES

Name Family Analytic No. Ref.

Agromyzidæ LIII 127, 138 Anthomyidæ XXXIII 74 Asilidæ XVIII 40 Astiadæ LV 126 Bibionidæ IV 11 Bombylidæ XIX 42 Borboridæ LVI 106 Braulidæ LIX 143 Cecidomyidæ II 3 Chironomidæ VI 14 Chloropidæ LI 111 Conopidæ XXIX 54, 60 Cordyluridæ XXXIV 82 Culicidæ IX 18 Cyrtidæ XXII 38 Dixidæ X 19_a_ Dolicopodidæ XXIV 50, 65 Drosophilidæ L 114, 131 Empidæ XXIII 45, 49, 66_a_ Ephydridæ XLIX 113 Geomyzidæ XLVIII 132 Helomyzidæ XXXVI 80 Heteroneuridæ XXXVII 78 Hippoboscidæ LVIII 144 Leptidæ XVII 31, 37 Limnobidæ XII 23 Lonchæidæ XLIII 92 Lonchopteridæ XXV 51 Micropezidæ XL 103, 109 Milichidæ LII 129 Muscidæ XXXII 73 Mycetophilidæ III 10 Nycteribidæ LX 145 Œstridæ XXX 62 Opomyzidæ XLV 139 Orphnephilidae VII 19_b_ Ortalidæ XLI 87, 104 Phoridæ LVII 140 Phycodromidæ XXXV 89, 116 Phytomyzidæ LIV 68, 125 Piophilidæ XLVII 120 Pipunculidæ XXVII 58 Platypezidæ XXVI 53 Psilidæ XXXIX 101, 135 Psychodidæ VIII 16 Ptychopteridæ XI 21 Pulicidæ I 0 Rhyphidæ XIV 8 Sapromyzidæ XLIV 93 Scenopinidæ XXI 44 Sciomyzidæ XXXVIII 86, 95 Sepsidæ XLVI 83, 102, 121, 136 Simulidæ V 13 Stratiomyidæ XV 28 Syrphidæ XXVIII 59 Tabanidæ XVI 30 Tachinidæ XXXI 72 Therevidæ XX 43 Tipulidæ XIII 24 Trypetidæ XLII 97

PLATE II

_Family Type-Forms of Antennæ in the first section, or Flies with many-jointed Antennæ—NEMOCERA_

II.—CECIDOMYIDÆ

1. Epidosis longipes, ♂

2. Asphondylia sarothamni

III.—MYCETOPHILIDÆ

3. Sciara thomæ

4. Mycetophila cingulum

5. Mycetobia pallipes

6. Macrocera lutea

IV.—BIBIONIDÆ

7. Bibio marci, ♀

8. Scatopse notata

V.—SIMULIDÆ

9. Simulium reptans

VI.—CHIRONOMIDÆ

10. Chironomus plumosus

11. Ceratopogon femoratus, ⚲

12. Clunio marinus

IX.—CULICIDÆ

13. Mochlonyx velutinus

14. Corethra plumicornis, ♀

XII.—LIMNOBIDÆ

15. Limnophila dispar

16. Rhipidia maculata, ⚲

17. Erioptera flavescens

18. Trichocera hiemalis

XI.—PTYCHOPTERIDÆ

19. Ptychoptera contaminata

XIII.—TIPULIDÆ

20. Tipula gigantea

21. Ctenophora pectinicornis

XIV.—RHYPHIDÆ

22. Rhyphos cinctus

PLATE III

_Family Type-Forms of Antennæ in the second section, or Flies with three or few-jointed Antennæ_

XV.—STRATIOMYIDÆ

1. Pachygaster atra

2. Oxycera pulchella

3. Stratiomys chamæleon

4. Sargus cuprarius

5. Beris chalybeata

6. Nemotelus uliginosus

XVI.—TABANIDÆ

7. Chrysops cæcutiens

8. Hæmatopota pluvialis

9. Tabanus bovinus

XVII.—LEPTIDÆ

10. Chrysopilus auratus

11. Leptis scolopacea

12. Atherix ibis

13. Symphoromyia melaena

14. Spania nigra

XVIII.—ASILIDÆ

15. Leptogaster cylindrica

16. Dioctria œlandica

17. Isopogon brevirostris

18. Laphria marginata

19. Asilus crabroniformis

XIX.—BOMBYLIDÆ

20. Anthrax hottentota

21. Bombylius major

XX.—THEREVIDÆ

22. Thereva annulata

XXI.—SCENOPINIDÆ

23. Scenopinus fenestralis

XXIII.—EMPIDÆ

24a. Empis livida

24b. Empis borealis

25. Ardoptera irrorata

26. Ocydromia glabricula

27. Stilpon lunata

28a. Clinocera stagnalis

28b. Clinocera lota

29. Hybos grossipes

30. Tachista arrogans

31. Chersodromia hirta

32. Cyrtoma spuria

XXIV.—DOLICHOPODIDÆ

33. Psilopus wiedemanni

34a. Dolichopus longicornis

34b. Dolichopus discifer

35. Chrysotus gramineus

36. Rhaphium longicorne

37. Medeterus diadema

XXV.—LONCHOPTERIDÆ

38. Lonchoptera lutea

XXVI.—PLATYPEZIDÆ

39. Platycnema pulicaria

40. Platypeza picta

XXVII.—PIPUNCULIDÆ

41. Chalarus spurius

42. Pipunculus campestris

XXVIII.—SYRPHIDÆ

43. Volucella bombylans

44. Xylota sylvarum

45. Orthoneura elegans

46. Chrysotoxum arcuatum

XXIX.—CONOPIDÆ

47. Myopa buccata

48. Physocephala rufipes

XXXI.—TACHINIDÆ

49. Zophomyia temula

50. Sarcophaga carnaria

XXXII.—MUSCIDÆ

51. Calliphora vomitoria

XXXIV.—CORDYLURIDÆ

52. Coniosternum obscurum

53. Scatophaga stercoraria

XXXVIII.—SCIOMYZIDÆ

54. Tetanocera ferruginea

LVI.—BORBORIDÆ

55. Borborus nitidus

XXX.—ŒSTRIDÆ

56. Hypoderma bovis

LVII.—PHORIDÆ

57. Phora incrassata

PLATE IV

_Family Type-Forms of the Wings of NEMOCERA_

II.—CECIDOMYIDÆ

1. Cecidomyia rosaria

2. Lasioptera rubi

3. Catocha latipes

III.—MYCETOPHILIDÆ

4. Sciara thomæ

5. Mycetophila cingulum

6. Macrocera lutea

IV.—BIBIONIDÆ

7. Bibio pomonæ

V.—SIMULIDÆ

8. Simulium reptans

VI.—CHIRONOMIDÆ

9. Ceratopogon illustris

10. Chironomus plumosus

11. Tanypus monilis

VIII.—PSYCHODIDÆ

12. Psychoda sexpunctata

IX.—CULICIDÆ

13. Culex annulatus

X.—DIXIDÆ

14. Dixa maculata

XI.—PTYCHOPTERIDÆ

15. Ptychoptera contaminata

XIV.—RHYPHIDÆ

16. Rhyphus cinctus

XII.—LIMNOBIDÆ

17. Dicranomyia stigmatica

18. Rhipida maculata

19. Symplecta punctipennis

20. Erioptera fuscipennis

21. Limnophila nemoralis

22. Trichocera hiemalis

XIII.—TIPULIDÆ

23. Nephrotoma dorsalis

24. Dolichopeza sylvicola

25. Pachyrrhina crocata

26. Tipula gigantea

PLATE V

_Family Type-Forms of the Wings in the second section, or Flies with few-jointed Antennæ_

XV.—STRATIOMYIDÆ

1. Sargus cuprarius

XVI.—TABANIDÆ

2. Tabanus bovinus

XVII.—LEPTIDÆ

3. Leptis scolopacea

XVIII.—ASILIDÆ

4. Asilus crabroniformis

5. Isopogon brevirostris

XIX.—BOMBYLIDÆ

6. Bombylius major

7. Anthrax hottentota

XX.—THEREVIDÆ

8. Thereva nobilitata

XXI.—SCENOPINIDÆ

9. Scenopinus fenestralis

XXIII.—EMPIDÆ

10. Empis livida

11. Ocydromia glabricula

12. Stilpon lunata

XXII.—CYRTIDÆ

13. Acrocera globulus

XXIV.—DOLICHOPODIDÆ

14. Dolichopus plumipes

15. Medeterus diadema

XXV.—LONCHOPTERIDÆ

16. Lonchoptera lutea, male. The dotted line shows V, 6 in the female.

XXVI.—PLATYPEZIDÆ

17. Platypeza picta

XXVII.—PIPUNCULIDÆ

18. Verralia aucta

XXVIII.—SYRPHIDÆ

19. Syritta pipiens

19-1/2. Eumerus sabulonum

20. Eristalis tenax

21. Brachyopa bicolor

XXIX.—CONOPIDÆ

21-1/2. Physocephala rufipes

XXX.—ŒSTRIDÆ

22. Gastrophilus equi

23. Œstrus ovis

XXXI.—TACHINIDÆ

24. Gymnosoma rotundatum

25. Cercomyia thoracica

26. Dexiosoma caninum

27. Sarcophaga carnaria

XXXII.—MUSCIDÆ

28. Calliphora erythrocephala

29. Mesembrina meridiana

XXXIII.—ANTHOMYIDÆ

30. Anthomyia pluvialis

XXXIV.—CORDYLURIDÆ

31. Scatophaga stercoraria

XXXVIII.—SCIOMYZIDÆ

32. Tetanocera ferruginea

XXXIX.—PSILIDÆ

33. Psila fimentaria

XLII.—TRIPETIDÆ

34. Tephritis corniculata

LVII.—PHORIDÆ

35. Phora rufipes

XL.—MICROPEZIDÆ

36. Micropeza corrigiolata

LI.—CHLOROPIDÆ

37. Centor cereris

LIV.—PHYTOMYZIDÆ

38. Chromatomyia affinis

LVI.—BOBORIDÆ

39. Limosina fontinalis

LVIII.—HIPPOBOSCIDÆ

40. Hippobosca equina

PLATE VI

1. Head of Mycetophila to show the position of the two ocelli.

2. Fore-leg of Dilophus

3. Fore-leg of Bibio

4. Palpus of Limnobia

5. Palpus of Tipula

6. Palpus of Anopheles

7. Palpus of Culex

8. Foot of Dilophus

9. Foot of Leptis

10. Foot of Asilus

11. Foot of Thereva

12. Profile of Eumerus

13. Profile of Syritta

14. Profile of Helophilus

15. Profile of Volucella

16. Profile of Rhingia

17. End of abdomen of Dolichopus 4 (_a_) the forceps (_b_) the pennis (_c_) the lamellæ

18. Head of Psarus, viewed from above, showing frontal process

19. Shape of the proboscis in Empis

20. Shape of the proboscis in Hilara

21. Shape of the proboscis in Ocydromia

22. Shape of the proboscis in Euthyneura

23. Hind leg of Ramphomyia pennata

24. Hind leg of Sphœrocera subsultans

24-1/2. Palpus of Lispe tentaculata

25. Leg of Limnophila—tibia spurred

26. Leg of Erioptera—tibia not spurred

27. Head and double-jointed proboscis of Myopa

28. Head of Asilus—front view 4 (_a_) the hollowed vertex (_b_) the mystax

29. Head of Drymia

30. Head and proboscis of Stomoxys calcitrans

31. Dorsum of Culex—no cross-seam

32. Dorsum of Tipula—cross-seams (_a_—_a_), (disregarding characters other than cross-seams)

33. Dorsum of Muscid (disregarding characters other than cross-seams)

34. Extremity of abdomen of Pedicia rivosa

35. Head of Stenopteryx sunk in shoulders

36. Foot and claws of Hippobosca

37. Puparium of Calliphora

38. T-shaped emergence-orifice in the pupa of Dixa—dorsal aspect

39. A mummy-pupa (Tipula)

40. Diagram of the Dorsum 4 (1-1) Humeri or Shoulders

(2) Præ-sutural Central Region

(3) Post-sutural Central Region

(4-4) Post-humeral Region

(5-5) Alar Regions

(6) The Scutellum

(7) The Neck

(8) Dorsal Suture

(9-9) Dorso-pleural Sutures

(10-10) Wing bases

(11-11) Scutellar Bridges

(12-12) The Alar Frena

(13-13) Inner rows of Dorso-central Bristles

(14-14) Outer rows of Dorso-central Bristles

(15) The Scutellar Suture

41. Diagramatic Chætotaxy of an Abdominal Segment 4 (1-1) Side Fore-marginal bristles

(2-2) Side Hind-marginal bristles

(3) Central Fore-marginal bristles

(4) Central Hind-marginal bristles

(5) Discal bristles

PLATE VII

1. Profile of Dorycera graminum

2. Profile of Trigonometopus frontalis

3. Profile of Platystoma seminationis

4. Profile of Ceroxys crassipennis

5. Head of Chilosia maculata ♂

6. Fore-leg of Platychirus manicatus, ♂ (5 should be 6).

7. Fore-leg of Platychirus peltatus, ♂

8. Fore-leg of Platychirus scutatus, ♂

9. Fore-leg of Platychirus albimanus, ♂

10. Fore-leg of Platychirus clypeatus, ♂

11. Platychirus, ♂ and ♀, quadrate spotting

12. Melanostoma mellinum, ♀ triangular spotting

13. Larva of Microdon

14. Wing of Helomyza pallida

15 & 16. Diagram of a fly's leg. _Vide_ C, I, on page 91.

15. As viewed from behind

16. As viewed, horizontal section (through A-B) 4 (_b.f._) base of femur

(_t.f._) tip of femur

(_b.t._) base of tibia

(_t.t._) tip of tibia

(1) upper or outer side

(2) under or inner side

(3) hinder side

(4) fore side

(5) upper fore side

(6) upper hind side

(7) under hind side

(8) under fore side

17 & 18. Base of wing, illustrating the difference in O, 6^1 respectively in

17. Urophora

18. Trypeta

19. Profile of Drosophila

20. Profile of Ceratomyza denticornis

21. Profile of Tichomyza fusca

22. Antennæ of Notophila cinerea; note the "thorn" on second joint

23. Antennæ of Hydrellia griseola

24. Antennæ of Tichomyza fusca

25. Profile of Hydrellia griseola

26. Profile of Ochthera mantis

27. Wing of Borborus equinus

28. Wing of Limosina sylvatica

29. Section of a collector's net, showing 4

(_a_) the small end of a large net;

(_b_) a detachable muslin bag;

(_c_) elastic closing the bag;

(_d_) transparent celluloid end

30. A small bag, closed and detached

31. Box for carrying bags

NUMBERED LIST OF FAMILIES,

_Together with references to the Analytical Table and Notes._

The sequence of the numbering indicates some proximity of relationship, in most cases at least, in respect of some particulars.

Nos. Refs. FAMILIES—Notes

I —. PULICIDÆ: fleas regarded as flies with atrophied or undeveloped wings. About 30 native species.

II 3. CECIDOMYIDÆ: mostly gall-gnats, minute and midge-like, or very frail, slender and gnat-like. _Cecidomyia destructor_ is the notorious Hessian-fly, injurious to cereal crops.

III 10. MYCETOPHILIDÆ: fungus-gnats; 34 genera; many common species. The marvellous "army-worm," the larvæ of _Sciara militaris_, which (being not a feeder on fungi) may be classified either separately or as belonging to this family, has been observed travelling in Continental forests in millions, described as massed together with a viscous mucus in bands said to be as large as 5 or 6 inches wide, 1 inch deep, and 50 or more feet long!

IV 11. BIBIONIDÆ: 40 species of small and medium sizes; commonly the males are black, but the females of some species coloured; _Bibio marci_, of a largish size, pubescent, and black, sometimes appears in great numbers on fine spring days, as likewise do some smaller black species of the same genus; ocelli absent; semi-blind.

V 13. SIMULIDÆ: midge-like flies, mostly with wings, limbs, and bodies of short length; the largest, _Simulium reptans_, only 2-3 mm., is common; its larvæ are aquatic. Some foreign species are called sand-flies and are much dreaded poisonous blood-suckers.

VI 14. CHIRONOMIDÆ: midges and many small, slender and gnat-like flies; _Ceratopogon pulicaris_, _C. bipunctatus_, and _C. varius_ are our commoner native blood-sucking midges, of which the larvæ have been found under the damp bark of decadent trees; the larvæ of some others are aquatic, as also are all those of the long slender non-blood-sucking species, including those of the very common harlequin-fly, _Chironomus plumosus_, as also of several congerers, including _Ch. dorsalis_, of which the larvæ are known as "pond blood-worms."

VII 19_b_. ORPHNEPHILIDÆ (darkness-loving): all foreign flies, except one rare.

VIII 16. PSYCHODIDÆ: minute moth-like flies; they have been named owl-midges from a peculiar manner of holding their wings in repose.

IX 18. CULICIDÆ: the true gnats or mosquitoes; 5 genera, 18 native species; three common, including _Culex pipiens_; larvæ abundant about marshy land and everywhere in water from which _fish are absent_, even in brackish water, but particularly in the stagnant water of small pools and vessels.

X 19_a_. DIXIDÆ: a few small and gnat-like flies; larvæ aquatic.

XI 21. PTYCHOPTERIDÆ: larvæ in shallow muddy pools.

XII 23. LIMNOBIDÆ: 32 genera and more than 100 species; generally marsh and fen slender flies, large and small, rather resembling "crane-flies"; the larvæ live some in decaying vegetable matter, some in fungi, some are aquatic, and some are unknown; two species called "winter-gnats," genus _Trichocera_, are very common.

XIII 24. TIPULIDÆ: true crane-flies or daddy-long-legs; about 60 species; larvæ (leather-jackets) underground in turf, or in decadent matter.

XIV 8. RHYPHIDÆ: three native species; slender and of medium size; larvæ in rotting vegetable matter.

XV 28. STRATIOMYIDÆ: 12 genera, 40 or more species; proboscis imperfect; wings rather small in comparison with the body, which is free from hairiness; a few are those large and conspicuously bright-coloured flies, which are called "soldier-flies"; the larvæ of the genus _Stratiomys_ are known as "star-tailed" maggots.

XVI 30. TABANIDÆ: breeze-flies; ocelli absent; the short well-developed proboscis of the blood-sucking female pierces the skin of mammalia; the abdomen is somewhat pubescent, never hairy; semi-blind; larvæ in damp earth, predaceous.

XVII 31, 37. LEPTIDÆ: a family of few species resembling some Empidæ rather than the flies of any preceding family. _Leptis scolopacea_ is a large fly, common in meadows, yellowish body with black spots.

XVIII 40. ASILIDÆ: the "robber-flies," which are more boldly voracious than any other raptorial flies, preying on winged insects, large and small; terrestrial predaceous larvæ.

XIX 42. BOMBYLIDÆ: furry, hovering, bee-like flies, mostly very long-tongued; bodies very rotund but with legs singularly thin; larvæ, some parasitic on the larvæ of ground bees, some unknown.

XX 43. THEREVIDÆ: non-raptorial flies; smaller, shorter, and more feeble than Asilidæ; though differing much in wing-pattern superficially like some Leptidæ or some Empidæ.

XXI 44. SCENOPINIDÆ: a very limited family of smallish flies. _Scenopinus fenestralis_, the "window-fly" was formerly thought to breed amidst old carpets and musty neglected clothing, but its larvæ have been found to be therein predaceous devourers of the larvæ of clothes-moths and fleas; larvæ of other species feed on fungi.

XXII 38. CYRTIDÆ: quite unlike any of the preceeding or succeeding families; flies with diminutive heads and large rotund abdomens; proboscis very short or obsolete; the larvæ of some are said to be parasitic on spiders.

XXIII 45, 49, 66_a_. EMPIDÆ: an extensive family; about 200 species, mostly of small sizes; of slender habit, and of dull colours; there are 30 genera and the characteristics of some are peculiar; the life-history and habits of most are unrecorded; though with bodies of feeble appearance, some have the proboscis well developed and are predaceous on small insects. Amongst the most curious are some species of the genus _Hilara_, of which the males have the first tarsal joint of the fore-leg thickened and flattened; some species of this genus, not very uncommon, fly and float about carrying "veils" or small cob-web-like attachments.

XXIV 50, 65. DOLICHOPODIDÆ: long-legged; a large family of 42 genera; some species of medium size but mostly small, amongst which latter bright metallic colours, often golden green, are common. The life-history of only a few is known.

XXV 51. LONCHOPTERIDÆ: a few small slender flies with long pointed wings.

XXVI 53. PLATYPEZIDÆ: small flies; the peculiar broad flat larvæ of some of the genus _Platypeza_ have been found in fungi.

XXVII 58. PIPUNCULIDÆ: small and uncommon flies with abnormally large eyes.

XXVIII 59. SYRPHIDÆ: a most interesting group of 51 genera; "hover-flies," and other various, conspicuous, large and medium sized flies (only a few small), very distinctly characterised and differentiated; the wings show the "vena spuria," and the face is without a "frontal suture." The larvæ are very diversified; some are terrestrial, some aquatic, some insectivorous, some parasitic or commensal in the nests of _Hymenoptera_.

XXIX 54, 60. CONOPIDÆ: the wings show no "vena spuria"; the known larvæ are parasitic on _Hymenoptera_ and _Orthoptera_.

XXX 62. ŒSTRIDÆ: a few very distinct species; proboscis and mouth parts atrophied; very hairy except one very rare; parasites of mammalia.

XXXI 72. TACHINIDÆ: 100 genera, more than 250 native species; with great difficulty classed in divisions, which may rank as separate families; stoutness of body and abundance of strong hairs, or rather _bristles_, are so generally apparent that many species may be guessed to belong to this family rather than to the closely related Muscidæ. Many of the larvæ are parasitic on various insects.

XXXII 73. MUSCIDÆ: 15 genera, comprising 36 native species, mostly very common, and many superabundant throughout summer by reason of rapid breeding.

XXXIII 74. ANTHOMYIDÆ: 4 sub-families (MYDÆINÆ, ANTHOMYINÆ, HOMALOMYINÆ, CŒNOSINÆ), 36 genera, and nearly 300 native species; difficult to characterise, but many rather resemble the lesser house-fly in size, and more or less in appearance, habit, and life-history, but some seem attracted rather more to flowers and others drawn only towards dung.

XXXIV 82. CORDYLURIDÆ: 29 genera; absence of squamæ apparent; otherwise generic features and general sizes and bodily shapes of species vary considerably. The yellow cow-dung fly, _Scatophaga stercoraria_, is the commonest species of this large family.

XXXV 89, 116. PHYCODROMIDÆ: sea-shore flies.

XXXVI 80. HELOMYZIDÆ: 8 genera; the costa of wings very "pectinate"; wings large and abdomen small.

XXXVII 78. HETERONEURIDÆ: 3 native species; smallish elongated wings; the larvæ, which live in rotten wood, can jump, somewhat like the cheese-hopper maggots.

XXXVIII 86, 95. SCIOMYZIDÆ: 11 genera; slender flies with tinted brownish wings; larvæ aquatic.

XXXIX 101, 135. PSILIDÆ: 4 genera.

XL 103, 109. MICROPEZIDÆ: 6 native species.

XLI 87, 104. ORTALIDÆ: about 20 species; smallish flies; some common in pastures; legs short and stout.

XLII 97. TRIPETIDÆ: numerous small species; larvæ in stems of plants, or galls thereon, some leaf-miners.

XLIII 92. LONCHÆIDÆ: some are of a plump figure and a dark metallic blue or green, others more slender and yellow, grey, or black.

XLIV 93. SAPROMYZIDÆ: 21 species.

XLV 139. OPOMYZIDÆ: 5 species; life-history unknown.

XLVI 83, 102, 121, 136. SEPSIDÆ: several are common small dung-flies, with black bodies, somewhat ant-like.

XLVII 120. PIOPHILIDÆ: the "cheese-hopper" maggots are larvæ of _Piophila casei_; seven other species.

XLVIII 132. GEOMYZIDÆ: 12 species.

XLIX 113. EPHYDRIDÆ: nearly 100 species; life-history of few known.

L 114, 131. DROSOPHILIDÆ: larvæ leaf-miners. The pale yellow "fruit-fly," _Drosophila fenestrarum_, loves the cider-press and fermenting fruit.

LI 111. CHLOROPIDÆ: numerous family; in the genus _Chlorops_ are many beautiful little flies.

LII 129. MICHILIDÆ: 2 native species.

LIII 127, 138. AGROMYZIDÆ: some are leaf-miners.

LIV 68, 125. PHYTOMYZIDÆ: leaf-miners.

LV 126. ASTIADÆ: 2 species; life-history unknown.

LVI 106. BORBORIDÆ: first tarsal joint (C, 1, 5) of hind leg conspicuously thick and short; small species of the genera _Borborus_ and _Spherocera_ often abound in the dung-pits of mews. The larvæ of some have been found on rotting potatoes and on confervæ.

LVII 140. PHORIDÆ: a numerous family of small flies; 6 genera.

LVIII 144. HIPPOBOSCIDÆ: the females of some shed their wings, and all live a lice-like life attached to animals.

LIX 143. BRAULIDÆ: eyeless and with strangely atrophied body appendages; found in bee-hives.

LX 145. NYCTERIBIDÆ: wingless parasites on bats; with small heads held in a curious posture.

ANALYTICAL TABLE OF FAMILIES

ORDER—DIPTERA

1 (141). Insects with the head distinct from the thorax, with one pair of wings, one pair of halteres, 5-jointed tarsi, proboscis not spirally coiled; with a well defined pupal stage intermediate between the larval stage and the perfect insect.

1-1/2 (52-1/2). Frontal lunule absent.

SUB-ORDER I. ORTHORRHAPHA (comprising 25 families); flies with "mummy-pupæ," _see plate_ VI, figs. 38 and 39.

2 (25). Antennæ many-jointed (_plate_ II).

NEMATOCERA (thread-horns)—

3 (4). Vein endings round the margin (_plate_ IV, figs. 1, 2, 3) not more than 6.

CECIDOMYIDÆ, II

4 (3). Vein endings more than 6.

5 (20). Thorax (_plate_ VI, 31) without B, I, 8 conspicuous.

6 (15). Costa ending at apex of wing (_plate_ IV, 4-11).

7 (12). Ocelli as in _plate_ I, or as in _plates_ VI, I and II, 4.

8 (9). Discal cell (O, 4_b_ or O, 5^2) present (_plate_ IV, 16).

RHYPHIDÆ, XIV

9 (8). Discal cell absent (_plate_ IV, 4-7).

10 (11). Antennæ long (_plate_ II, 3-6); or, if short, coxæ elongated.

MYCETOPHILIDÆ, III

11 (10). Antennæ as in _plate_ II, 7 and 8; coxæ not elongated.

BIBIONIDÆ, IV

12 (7). Ocelli absent.

13 (14). Antennæ (_plate_ II, 9) shorter than B; wings (_plate_ IV, 9) broad.

SIMULIDÆ, V

14 (13). Antennæ (_plate_ II, 10, 11) longer; wings narrow (_plate_ IV 9-11).

CHIRONOMIDÆ, VI

15 (6). Costa (_plate_ IV, 12-14) around hind margin.

16 (17). Wings held roof-like, hairy; X only at base (_plate_ IV, 12).

PSYCHODIDÆ, VIII

17 (16). Wings flat over D; X on middle of wing.

18 (19). Veins hairy (_plate_ IV, 13).

CULICIDÆ, IX

19 (18). Veins bare.

19_a_ (19_b_). Antennæ long; V, 4 and V, 2 forked; X, 4 (_plate_ IV, 14) ending in V, 2.

DIXIDÆ, X

19_b_ (19_a_). Antennæ very short but 11-jointed; no veins forked.

ORPHNEPHILIDÆ, VII

20 (5). Thorax (_plate_ VI, 32) with a strong cross seam.

21 (22). V, 6 absent (_plate_ IV, 15).

PTYCHOPTERIDÆ, XI

22 (21). V, 6 present.

23 (24). V, 1_a_ ending in costa and (_plate_ IV, 17-22) united to V, 1_b_ by X, 1_b_; last joint of palpi (_plate_ VI, 4) never whip-like.

LIMNOBIDÆ, XII

24 (23). V, 1_a_ ending in V, 1_b_; and, except for X, 1_a_ at its base, not united by X (_plate_ IV, 23-26); palpi (_plate_ VI, 5).

TIPULIDÆ, XIII

25 (2). Antennæ short, 3-jointed (_plate_ III); 3rd joint sometimes ringed.

BRACHYCERA (short-horns)—

26 (140). Wing veining normal; X-veins present.

27 (32). Third antennal joint ringed; A, IV, 6, if present (_plate_ III, 1-9) apical.

28 (29). Costa ending at apex of wing (_plate_ V, 1).

STRATIOMYIDÆ, XV

29 (28). Costa around the hind margin (_plate_ V, 2).

30 (31). Squamæ very large.

TABANIDÆ, XVI

31 (30). Squamæ very small.

_Part of_ LEPTIDÆ, XVII (genus Xylophagus)

32 (27). Third joint of antennæ (_plate_ III, 10-55) not ringed.

33 (55). Arista terminal or wanting (_plate_ III, 10-26, 28-32, 35-40, 48)

34 (46). V, 3 forked (_plate_ V, 3-10, 13).

35 (45). O, 6^1 long, extending (or nearly) to hind margin, or entirely wanting (_plate_ V, 3-9, 13).

36 (39). C, I, (II) much enlarged, appearance of 3 foot-pads (_plate_ VI, 9)

37 (38). Squamæ small, not covering C, III; O, 6^1 long and distinct (_plate_ V, 3); long bodied flies, head of normal proportions.

_Part of_ LEPTIDÆ, XVII

38 (37). Squamæ very large, hiding the halteres; O, 6^1 absent (_plate_ V, 13); short round-bodied flies with very small heads.

CYRTIDÆ, XXII

39 (36). Empodium small, only 2 foot-pads apparent (_plate_ VI, 10, 11).

40 (41). Vertex hollowed out between the eyes (_plate_ VI, 28).

ASILIDÆ, XVIII

41 (40). Vertex flat or raised, eyes not protruding.

42 (43) (44). V, 4 ending in 3 branches (_plate_ V, 6, 7) between V, 3 and V, 5.

BOMBYLIDÆ, XIX

43 (42) (44). V, 4 ending in 4 branches (_plate_ V, 8) between V, 3 and V, 5.

THEREVIDÆ, XX

44 (42) (43). V, 4 ending in 2 branches (_plate_ V, 9); antennæ (_plate_ III, 23) without A, IV, 6.

SCENOPINIDÆ, XXI

45 (35). O, 6^1 short (_plate_ V, 10) not nearly reaching hind margin.

_Part of_ EMPIDÆ, XXIII

46 (34). V, 3 not forked (_plate_ V, 11-12, 14-17).

47 (52). Alulæ absent or rudimentary.

48 (51). Apex of the wing rounded.

49 (50). X, 4 placed generally beyond X, 5^1, at most only a little before it; V, 5 with 2 X-veins bounding a discal cell (_plate_ V, 11); or, if only one, then X, 5 is placed before, or, at most (_plate_ V, 12) only a little beyond X, 4.

_Part of_ EMPIDÆ, XXIII

50 (49). X, 4 placed very near the base of the wing; V, 5 with only one X-vein placed far beyond X, 4; no discal cell (_plate_ V, 14, 15).

_Part of_ DOLICHOPODIDÆ, XXIV

51 (48). Apex of the wing (_plate_ V, 16) pointed.

LONCHOPTERIDÆ, XXV

52 (47). Alulæ present.

52-1/2 (1-1/2). Frontal lunule (_plate_ VII, 5) present.

SUB-ORDER II. CYCLORRHAPHA (families XXVI—LX); flies which escape from a puparium, formed of the indurated larval skin, through a circular orifice made by pushing off its head end (_plate_ VI, 37).

PROBOSCIDEA—

53 (54). Antennæ (_plate_ III, 39, 40) with a long seta or bristle.

PLATYPEZIDÆ, XXVI

54 (53). Antennæ (_plate_ III, 48) with a short seta or spine.

_Part of_ CONOPIDÆ, XXIX

55 (33). Arista dorsal (_plate_ III, 27, 33, 34, 41-47, 49-55).

56 (61). O, 6^1 long (_plate_ V, 18-21-1/2) extending nearly to hind margin: D without strong bristles (macrochætæ).

57 (60). Proboscis of moderate length (_plate_ VI, 12-16) generally retracted.

58 (59). Vena spuria (_plate_ V, 18) absent; A, II very large; O, 4^2 open.

PIPUNCULIDÆ, XXVII

59 (58). Vena spuria present (_plate_ V, 19-21); O, 4^2. closed; A, II moderate.

SYRPHIDÆ, XXVIII

60 (57). Proboscis projecting far in front, very thin and long, often double-kneed (_plate_ VI, 27); no vena spuria (_plate_ V, 21-1/2).

_Part of_ CONOPIDÆ, XXIX

61 (56). O, 6^1 short, not nearly reaching margin, or absent (_plate_ V, 12, 22-39) abdomen often with strong bristles.

62 (63). Proboscis and palpi atrophied.

ŒSTRIDÆ, XXX

63 (62). A, VIII, 3 and 4 distinctly present.

64 (67). X, 5^1 absent, X 5^2 present, _i.e._, only one X-vein on V, 5 and that beyond X, 4.

65 (66). X, 4 very near base of wing (_plate_ V, 14, 15); X, 5 far beyond X, 4.

_Part of_ DOLICHOPODIDÆ, XXIV

66 (65). X, 4 more towards middle of wing (_plate_ V, 12, 36, 37) nearer X, 5.

66_a_ (66_b_). Proboscis directed straight downwards.

_Part of_ EMPIDÆ, XXIII (genus Stilpon)

66_b_ (66_a_). A, VIII, 3 retracted [ordinary Muscid type] pass on to 108 of this table.

67 (64). X, 5^1 present, that is, a X-vein on V, 5 before, or under X, 4; or no X-veins on V, 5.

68 (69). No X-veins on V, 5.

_Part of_ PHYTOMYZIDÆ, LIV

69 (68). At least X, 5^1 present, generally also X, 5^2.

70 (75). Squamæ distinctly present though sometimes small.

MUSCIDÆ CALYPTERÆ—

71 (74). V, 4 angled or bent towards V, 3 (_plate_ V, 24-29).

72 (73). D with E (macrochætæ and hairs) on middle segments; otherwise A, IV, 6 is bare or bare on the tip-half (_plate_ III, 49-50).

TACHINIDÆ, XXXI

73 (72). D without macrochætæ though often with rather strong hairs; A, IV, 6 fully plumed or combed (_plate_ III, 51).

MUSCIDÆ, XXXII

74 (71). V, 4 going straight or nearly straight to margin (_plate_ V, 30).

ANTHOMYIDÆ, XXXIII

75 (70). Squamæ absent or rudimentary.

MUSCIDÆ ACALYPTERÆ—

76 (105). V, 1_a_ distinctly present, nearly as thick as V, 1_b_, which generally ends in the fore margin at or beyond the middle of the wing (_plate_ V, 31, 32, 34).

77 (84). Vibrissæ present (_plate_ VIII, 2) together with more or less hair.

78 (79). V, 4^4 four or five times as long as V, 4^3 (_i.e._, X, 5^2 nearer X^4 than to wing margin).

HETERONEURIDÆ, XXXVII

79 (78). V, 4^4 about equal to V, 4^3, or only a little longer.

80 (81). Costa with spines in addition to the ciliation (_plate_ VII, 14).

HELOMYZIDÆ, XXXVI

81 (80). Costa without a row of spines, only ciliated.

82 (83). A, III (5) present, or, if absent, then body never shining black.

CORDYLURIDÆ, XXXIV

83 (82). Fronto-orbital bristles absent; body always shining black.

_Part of_ SEPSIDÆ, XLVI

84 (77). Vibrissæ absent; mouth with short hairs or bare (_plate_ VII, 1, 3, 4); or, if the hairs are strong, all of same length and strength.

85 (88). A, IV very long and horizontal; A, IV (4) as long or longer than A, IV (5) of which the upper edge is concave (_plates_ III, 54; VII, 1).

86 (87). Profile not triangular; face perpendicular or slightly retreating.

_Part of_ SCIOMYZIDÆ, XXXVIII

87 (86). Profile triangular; A, V retreating (_plate_ VII, 1) almost horizontally.

_Part of_ ORTALIDÆ, XLI

88 (85). Antennæ not long, or, if long, drooping; A, IV (4) shorter than A, IV (5) of which the upper edge is straight or convex.

89 (90). Brown or black shore flies with flat horny thorax and with strong mouth hairs.

_Part of_ PHYCODROMIDÆ, XXXV

90 (89). Otherwise than 89.

91 (94). O, 5^1 and O, 6^1 very small and indistinct.

92 (93). Only one fronto-orbital bristle on each side; tibiæ all without the præ-apical bristle.

LONCHÆIDÆ, XLIII

93 (92). A, III (5) on each side; the præ-apical E on some of the C, I (4).

SAPROMYZIDÆ, XLIV

94 (91). O, 5^1 and O, 6^1 fairly large and quite distinct.

95 (96). Hind tibiæ with a præ-apical bristle.

_Part of_ SCIOMYZIDÆ, XXXVIII

96 (95). Hind tibiæ without the præ-apical bristle.

97 (98). A, III (6) present and close to A, II; V, 1_a_ sharply bent up at its end (_plate_ V, 34). Wings usually spotted; ovipositor protruding.

TRYPETIDÆ, XLII

98 (97). Lower fronto-orbital bristles absent; V, 1_a_ gently bent up.

99 (104). Legs long; abdomen narrow and long, contracted at base.

100 (103). V, 4 straight or nearly so. Shining black species.

101 (102). V, 6 reaching to the wing margin; abdomen narrow, more than twice as long as the thorax.

_Part of_ PSILIDÆ, XXXIX

102 (101). V, 6 not nearly reaching margin; D rather broad.

_Part of_ SEPSIDÆ, XLVI

103 (100). V, 4 much bent towards V, 3. Brown species.

_Part of_ MICROPEZIDÆ, XL (genus Calobata)

104 (99). Legs normal; D comparatively short and broad, never contracted at base.

_Part of_ ORTALIDÆ, XLI

105 (76). V, 1_a_ absent or rudimentary, much thinner than V, 1_b_, which latter ends at or before the middle of the fore margin.

106 (107). Hind C, I (5) thickened and shorter than the next joint.

BORBORIDÆ, LVI

107 (106). Hind metatarsus longer than and not thicker than C, I(6).

108 (115). X, 5^1 absent, X, 5^2 present (_i.e._ only one X-vein on V, 5 and that placed beyond X, 4 (_plate_ V, 36).

109 (110). Legs long and thin; abdomen very long.

_Part of_ MICROPEZIDÆ, XL (genus Micropeza)

110 (109). Legs short and strong; D oval or elliptical.

111 (112). A, III (6) absent; A, IV (5) round, or, if lengthened, then A in profile is triangular.

CHLOROPIDÆ, LI

112 (111). A, III (6) present; A, IV (5) generally long or oval; A in profile never triangular.

113 (114). Head square; face strongly convex (_plate_ VII, 21); O, 6^1 absent. Not orange coloured species.

EPHYDRIDÆ, XLIX

114 (113). Head round or semi-circular; face not arched (_plate_ VII, 19); O, 6^1 present, or, if absent, then the species orange coloured.

_Part of_ DROSOPHILIDÆ, L.

115 (108). X, 5^1 present, or, if absent, then X, 5^2 also absent (_i.e._, either two X-veins or no X-veins on V, 5) (_plate_ V, 33, 38).

116 (117). Scutellum of the male extraordinarily long and squared off behind; costa thickened from the end of V, 1 to the wing tip.

_Part of_ PHYCODROMIDÆ (genus Thyreophora)

117 (116). B, I (6) usually short, and somewhat triangular; costa normal.

118 (133). Mouth bristles present.

119 (122). Fronto-orbital bristles absent.

120 (121). Wings without a black spot near the tip.

PIOPHILIDÆ, XLVII

121 (120). Wings with a black spot near the tip.

_Part of_ SEPSIDÆ, XLVI

122 (119). Fronto-orbital bristles present.

123 (128). X, 5^2 before middle of wing (both X, 4 and X, 5^2 near the base and exceedingly near together) or no X-vein on V, 5.

124 (127). X, 5^2 immediately under X, 4 (or nearer the base), or X, 5^1 and X, 5^2 both absent (_plate_ V, 38).

125 (126). Arista bare or nearly so.

_Part of_ PHYTOMYZIDÆ, LIV

126 (125). Arista with long hairs on one side.

ASTIADÆ, LV

127 (124). X, 5^2 present and placed further from base than X, 4.

_Part of_ AGROMYZIDÆ, LIII

128 (123). X, 5^2 at or beyond the middle; X, 4 and X, 5^2 neither very near to the wing base nor to one another.

129 (130). Arista bare; body short and broad.

MILICHIDÆ, LII

130 (129). Arista plumose, combed, or pubescent, or, if bare, then the body always narrow and long.

131 (132). Arista thinly but long haired, plumose, or combed.

_Part of_ DROSOPHILIDÆ, L

132 (131). Arista thickly and short haired, pubescent, or bare.

GEOMYZIDÆ, XLVIII

133 (118). Mouth bristles absent.

134 (137). O, 5^1 and O, 6^1 large and very prominent.

135 (136). Antennæ moderately long or very long, male genitalia not prominent.

_Part of_ PSILIDÆ, XXXIX

136 (135). Antennæ short; male genitalia prominent, and club-shaped.

_Part of_ SEPSIDÆ, XLVI (genus Tetanura)

137 (134). O, 5^1 and O, 6^1 small and not very prominent.

138 (139). Light grey or silvery species.

_Part of_ AGROMYZIDÆ, LIII

139 (138). Orange or brownish-red species.

_Part of_ OPOMYZIDÆ, XLV

140 (26). Wings with two short very thick veins (in fore-basal region) whence spring a few very fine unbranched veins. X-veins absent (_plate_ V, 35).

PHORIDÆ, LVII

141 (1) (52-3/4). Body of a horny or leathery nature; head sunk into a hollow in the thorax (_plate_ VI, 35). No proper proboscis or palpi; claws very large (_plate_ VI, 36). Perfect insect parasitic.

EPROBOSCIDEA—

142 (145). Head fairly large; wings sometimes shed or absent.

143 (144). Eyes, ocelli, scutellum, halteres, and wings all absent; last tarsal joint combed with bristles.

BRAULIDÆ, LIX

144 (143). Eyes and scutellum present; last tarsal joint with well developed claws.

HIPPOBOSCIDÆ, LVIII

145 (142). Head small, when at rest thrown back upon the dorsum; always wingless.

NYCTERIBIDÆ, LX

INDEX

Air-sacks, breathing by, 46

ANTHOMYIDA, 10, 17

_Asilus crabroniformis_, 55

Aphides, plant-lice, etc., 8

Beelzebub, Lord-of-flies, 2

BIBIONIDÆ, semi-blindness of, 108

Birds, food for, 80

Black death, 2

Blood-sucking flies, kinds of— genus, _chrysops_, 24 genus, _hæmatobia_, 16 genus, _hæmatopota_, 23 genus, _lyperosia_, 17 genus, _tabanus_, 23 genus, _therioplectes_, 23

Blood-sucking midges and gnats —common species— _ceratopogon pulicaris_, 108 _ceratopogon bipunctatus_, 108

_ceratopogon varius_, 108 (also others of the same genus and of the allied genus, _anopheles_) _culex pipiens_, 108

Blue-bottles, 12, 18, 33, 36, 51

_Borborus equinus_, 34

Bot-fly of the horse, 26

Breeze-flies, "blinden," 10, 23, 29, 69

Bridgewater Treatises, 1

Cabbage-fly, 17

_Calliphora erythrocephala_ } _see_ blue-bottles

_Calliphora vomitoria_ } _see_ blue-bottles

Celery-fly, 17

Chelifers, 55

_Chrysops cæcutiens_, 24 _Chrysops relicta_, 24

_Conops flavipes_, 22

Contagion, fly-borne, 84

CORDYLURIDA, 10

Cow-dung fly, the yellow, 19

Crane-flies, or Daddy-long-legs, 109

Cremation of refuse, 41, 61, 65, 70

_Cyrtoneura simplex_, 17

DIPTERÆ, nearly 3,000 species, 10

Dragon-flies, 55

Drone-flies, 20, 22

Dun-flies, _see_ Gad-flies

Dust-bins, preferably open, 65, 70

Dust destructor furnace, imperative, 65

Economic value of the fly, 41

Egg-traps, domestic refuse as, 65, 72

EMPIDÆ, 54

_Empusa muscæ_, 56

_Eristalis tenax_, 21

Exclusion from rooms, 76

_Fannia canicularis_, 7 _Fannia scalaris_, 14, 18

Filth-flies, 51

Fleas, 108

Flecked flesh, 29

Fly chart, the Wingate, 88

Fly-traps, 66

Forest (or spider) fly, 22

Fungus-flies, 51

Futurist's Credo, 39

Gad-flies, 23

Garden pests, vegetarian, 17

Gargantuan jokes, 32

_Gastrophilus_, _see Œstrus_

Gentles, apparatus for breeding, 80

Gentles, green, 82

_Glossina morsitans_, 62

Green-bottle flies, 20

Grey blow-flies, 19

_Hæmatobia stimulans_, 16

_Hæmatopota crassicornis_, 23 _Hæmatopota italica_, 23 _Hæmatopota pluvialis_, 23

Hibernation, 15, 80

HIPPOBOSCIDÆ, 22

Horse-flies, _see_ Forest-flies

House-fly fungus, 56

House-fly, larva of, 35

House-fly, wing pattern of, 12

Hover-flies, 20

Human intestinal myiasis, 25, 26

Humble Creatures, 1, 34 _hydrotæa dentipes_, 17 _hydrotæa irritans_, 17

Instar, pupa, or chrysalid, 40

Larvæ, _see_ Maggots

Lesser house-fly, larva of, 36 Lesser house-fly, wing pattern, 12

Louse-fly, tick, or ked, 22

_Lucilia Cæsar_, 20 _Lucilia sericata_, 20

_Lyperosia irritans_, 17

Maggots, feeding in company, 36 Maggots, insectivorous, 21 Maggots, jumping or hopping, 111, 112 Maggots, myiasic possibilities, 28, 37, 82 Maggots, parasitic on _lepidoptera_, etc., 111 Maggots, predaceous, 109 Maggots, "rat-tail," 21 Maggots, "star-tail," 109

Meal-worms, 80

_Merodon narcissi_, 21

Messengers of peril, 78

Metamorphosis, 37

MICRODON, 22

_Musca corvina_, 14, 17

_Muscina stabulans_, 18, 27

Nostril-fly of the sheep, 4, 26, 28

Ocelli, visual importance of, 45

Odour of humanity, attracted by, 33

Œstrid flies, egg-laying aggressions, 31, 69

_Œstrus hominis_, discredited, 28

_Œstrus (Gastrophilus) equi_, 27

_Œstrus (Gastrophilus) hæmorrhoidalis_, 26

_Œstrus nasalis_ 4

_Œstrus ovis_, 28 (_Œstrus ovis hypoderma_) _bovis_, _see_ worble-flies (_Œstrus ovis hypoderma_) _lineatum_, _see_ worble-flies

Onion-fly, 17

Owl-midges, moth-like, 109

Pestering flies, commoner kinds of— _cyrtoneura simplex_, 17 _fannia canicularis_, 7 _fannia scalaris_, 18 _musca domestica_, 7 _musca corvina_, 17 _muscina stabulans_, 18 and other ANTHOMYIDÆ

Poisons, effective insecticide, 73

_Polietes lardaria_, 18

Pond blood-worms, 108

Robber-flies, 54, 109

Root-fly, the, 17

_Sarcophaga carnaria_, 19

_Scatophaga stercoraria_, 19

Scavenging services, 78

_Scenopinus fenestralis_, 110

Sex differentiation, 45

Soldier-flies, 22, 109

Spiders, flies parasitic on, 110

Stable-fly, 12, 13, 16, 51

Stable manure, 67

_Stomoxys calcitrans_, 12, 13, 16, 51

STRATIOMYIDÆ, 22, 109

Swallow, attracted to breed, 53

Syrphidæ, 20

_Syrphus_, 21

TABANIDÆ, 10, 22, _see also_ blood-sucking flies

TACHINIDÆ, 10, 19

_Therioplectes_, _see_ blood-sucking flies

TIPULIDÆ, 109

Traps for flies, some ineffective, 72

Turnip-fly, a beetle, 17

Veins of wings, 11

Voltairean atheism, controversy of, 5

VOLUCELLA, 21

Warning signals, flies as, 78

Westminster Hospital and myiasis, 28

Window-fly, the, 110

Window screens, 77

Wingate's nomenclature symbols, 12

Wing patterns contrasted, 12

Worble-flies, 26, 28, 69

WOODS & SONS LTD., PRINTERS, LONDON, N.

Transcribers note.

This work is presented as is, with no changes to the spelling, except for confirmed typos. Printer's errors, where obvious, and original spelling have been retained.

Variations in hyphenation and compound words have been preserved.

Italicized words and phrases in the text version are presented by surrounding the text with underscores.

The Caret character (^) is used to denote superscripts.

Table of Contents, List of Illustrations, etc. Discrepancies: many of the items in the front matter sections do not match the titles in the book's content section.

End of Project Gutenberg's The Book of the Fly, by G. Hurlstone Hardy