mi. W Mariano Lake, 3 (MVZ); Gallup, 6 (USBS); Wingate, 6 (USBS);

Thoreau, 2 (USBS); Zuni River, 1 (USBS). _Sandoval County_: Juan Tafoya, 1 (USBS). _Valencia County_: 8 mi. SE Grants, 7 (UM); 9 mi. SSE Grants, 5 (UM); 2-1/2 mi. E El Morro P. O., Zuni Mts., 1 (LACM); Laguna, 1 (USBS); Laguna Indian Reservation, 7 (LACM). _Catron County_: Cougar Ranch, NE Adams Diggings, 1 (CAS); 10 mi. SW Quemado, 1 (USBS). _Socorro County_: 1 mi. S Bernardo, 1 (MVZ); Riley, 1 (USBS).

=Dipodomys ordii pallidus= Durrant and Setzer

_Dipodomys ordii pallidus_ Durrant and Setzer, Bull. Univ. Utah, 35 (no. 26):24, June 30, 1945.

_Type._--Male, adult, no. 3526, Museum of Zoology, University of Utah; Old Lincoln Highway, 18 mi. SW Orr's Ranch in Skull Valley, 4,400 ft., Tooele County, Utah; obtained on June 6, 1940, by S. D. Durrant, original no. 1905.

_Range._--Low valleys of west-central Utah in Tooele, Juab and Millard counties; marginal occurrences are: all in Utah, 18 mi. SW Orr's Ranch, 7 mi. S Fish Springs, Hinckley and Lynndyl.

_Diagnosis._--Size medium (see measurements). Color light, entire dorsal surface Light Pinkish Cinnamon, purest on sides and flanks, with but slight suffusion of black on upper parts; cheeks white; arietiform markings, pinnae of ears, plantar surfaces of hind feet, dorsal and ventral stripes of tail, brownish. Skull large; auditory bullae long, wide and well inflated; external auditory meatus elongate with a notch on the dorsal border; nasals long and flaring distally.

_Comparisons._--From _Dipodomys ordii celeripes_, _D. o. pallidus_ differs in: Size larger; color generally darker, although some specimens are lighter; skull larger; nasals longer and more flared distally; interorbital width greater; auditory bullae larger; external auditory meatus larger.

From _Dipodomys ordii fetosus_, _D. o. pallidus_ differs in: Size larger; color lighter; ventral stripe of tail indistinct as opposed to pronounced; skull larger; nasals longer and more flared distally; auditory bullae larger; external auditory meatus larger; palate shorter and broader.

For comparisons with _Dipodomys ordii marshalli_ and _Dipodomys ordii utahensis_ see accounts of those subspecies.

_Remarks._--Among named subspecies of _Dipodomys ordii_, _D. o. pallidus_ most closely resembles, morphologically, _D. a. marshalli_, its nearest geographic neighbor to the north and east. Intergradation with _Dipodomys ordii utahensis_ is noted in color and intermediacy of body size of specimens from Clover Creek. The majority of cranial characters, however, show these animals to be referable to _D. o. utahensis_ rather than to _D. o. pallidus_. Specimens taken at Lynndyl and Hinckley show intergradation in size of body, length and configuration of the nasals and the degree of inflation of the auditory bullae between _Dipodomys ordii celeripes_ and _D. o. pallidus_. The majority of characters studied show these latter animals to be referable to _D. o. pallidus_. Specimens from 35 and 60 miles west of Delta, in size of body and shape and inflation of the auditory bullae, are intermediate between _D. o. pallidus_ and _D. o. celeripes_. These specimens are here referred to _D. o. celeripes_.

_Specimens examined._--Total, 34, all from Utah, distributed as follows: _Tooele County_: Old Lincoln Highway, 18 mi. SW Orr's Ranch in Skull Valley, 4400 ft., 9 (UU). _Juab County_: Fish Springs, 4400 ft., 4 (UU); 7 mi. S Fish Springs, 4400 ft., 4 (UU). _Millard County_: 1 mi. N Lynndyl, 4768 ft., 5 (UU); Lynndyl, 4768 ft., 1 (UU); Hinckley, 4600 ft., 11 (UU).

=Dipodomys ordii nexilis= Goldman

_Dipodomys ordii nexilis_ Goldman, Journ. Washington, Acad. Sci., 23:470, October 5, 1933.

_Perodipus longipes_, Warren, Mammals of Colorado, p. 77, 1910 (part--the part from Coventry, Colorado).

_Dipodomys ordii longipes_, Moore, Journ. Mamm., 10:260, August, 1929 (part--the part from Monticello, Utah).

_Type._--Male, adult, no. 149938, U. S. Nat. Mus. Biol. Surv. Coll.; 5 mi. W Naturita, Montrose County, Colorado; obtained on July 20, 1907, by Merritt Cary, original no. 1068.

_Range._--West-central Colorado, southwest into San Juan County, Utah, north of the San Juan River; northwest into Grand County, Utah, to the Colorado River; and westward probably as far as the Colorado River; marginal occurrences are: in Utah, Cisco, 18 mi. NE Moab, and Blanding; in Colorado, 5 mi. W Naturita, and Coventry.

_Diagnosis._--Size medium (see measurements). Color dark, entire dorsal surface between (_a_) Cinnamon-Buff and Clay Color, purest on sides and flanks, upper parts strongly suffused with black; arietiform markings, pinnae of ears, plantar surfaces of hind feet, dorsal and ventral tail-stripes blackish. Lateral white stripes of tail less than one fourth of diameter of tail. Skull large; rostrum long and narrow; nasals long; auditory bullae strongly inflated; braincase not inflated; pterygoid fossae ovoid.

_Comparisons._--From _Dipodomys ordii sanrafaeli_, _D. o. nexilis_ differs in: Color darker; skull smaller in all measurements taken; auditory bullae noticeably smaller; width across zygomatic processes of maxillae less.

From _Dipodomys ordii longipes_, _D. o. nexilis_ differs in: Size smaller; color darker in all respects; rostrum wider; breadth across auditory bullae less; interorbital width greater; breadth across zygomatic processes of maxillae less; cutting edge of upper incisors wider; zygomatic arch lighter and more nearly straight.

Comparison with _Dipodomys ordii evexus_ is made in account of that subspecies.

_Remarks._--This animal is apparently not abundant at any place in its range. Two different attempts, by me, to obtain topotypes were unsuccessful. A single specimen was obtained and that was only a skull saved from a mutilated animal that was taken away from a rattlesnake. The habitat at the type locality is such that a person would not expect it to be inhabited by kangaroo rats. The soil is a heavy clay with a generous admixture of stones but in isolated spots there are light sandy soils which should be suitable for kangaroo rats. Even so, 500 traps set in the area of the type locality over a period of two nights yielded no _Dipodomys_.

This subspecies of _Dipodomys ordii_, inhabiting west-central Colorado and southeastern Utah, is darker than any other subspecies with which its range comes in contact. The races to both the north and south are larger, with the exception of the hind foot which is longer in _D. o. nexilis_ than in either _D. o. sanrafaeli_ or _D. o. longipes_. The affinities of _D. o. nexilis_ are with _D. o. longipes_ rather than with any of the other known subspecies of _Dipodomys ordii_.

Intergradation between _D. o. nexilis_ and _D. o. longipes_ is shown by animals from Bluff, San Juan County, Utah, which, however, are referable to the latter. In animals from sixteen miles northwest of Moab, Grand County, Utah, there is intergradation in size of skull and in color between _D. o. nexilis_ and _D. o. sanrafaeli_. The specimens are referred to _D. o. sanrafaeli_.

_D. o. nexilis_ is apparently prevented from extending its range to the northward by the presence there of _D. o. sanrafaeli_. To the south _D. o. nexilis_ encounters _D. o. longipes_ and the San Juan River. To the east it is limited by the Rocky Mountains and to the west by the deep canyons of the Colorado River.

Warren (1942:183) did not apply the name _D. o. nexilis_ to Ord kangaroo rats in Colorado. He listed specimens from Montrose, Montezuma and Mesa counties as being referable to _D. o. longipes_. The specimens from Montezuma County, Colorado, probably are _D. o. longipes_ and those from Montrose and Mesa counties (for which Warren gives no precise localities) are _D. o. nexilis_.

_Specimens examined._--Total, 35, distributed as follows:

=Utah=: _Grand County_: Cisco, 4 (CM); 18 mi. NE Moab, 6000 ft., 1 (UU). _San Juan County_: 15 mi. N Monticello, 4 (MVZ); Blanding, 1 (UU).

=Colorado=: _Delta County_: Hotchkiss, 1 (USBS). _Montrose County_: Paradox, 1 (DRD); Bedrock, 4 (AMNH); 5 mi. W Naturita, 2 (USBS); Coventry, 3 (1 USBS; 2 AMNH).

=Dipodomys ordii cupidineus= Goldman

_Dipodomys ordii cupidineus_ Goldman, Journ. Washington Acad. Sci., 14:372, September 19, 1924.

_Type._--Male, no. 243093, U. S. Nat. Mus. Biol. Surv. Coll.; Kanab Wash, at southern boundary of Kaibab Indian Reservation, Arizona; obtained on October 12, 1922, by E. A. Goldman, original no. 23384.

_Range._--Northeastern Arizona and south-central Utah; marginal occurrences are: in Utah, Escalante, near Paria and S of Virgin; in Arizona, 6 mi. N Wolf Hole, 20 mi. S Wolf Hole, 5 mi. S Trumbull Spring and 10 mi. S Jacobs Pools in Houserock Valley.

_Diagnosis._--Size large (see measurements). Color relatively dark, entire dorsal surface between (_16''_) Pinkish Cinnamon and Cinnamon-Buff, purest on sides and flanks, upper parts washed with black; arietiform markings, pinnae of ears, dorsal and ventral stripes of tail, plantar surfaces of hind feet, blackish. Skull large; auditory bullae well inflated; interorbital width relatively narrow; rostrum long and narrow; nasals long and slightly flared distally; pterygoid fossae ovoid; external auditory meatus small and ovoid; jugal light and relatively straight.

_Comparisons._--From _Dipodomys ordii chapmani_, _D. o. cupidineus_ differs in: Size larger; color lighter in all pigmented areas; skull larger; rostrum shorter and wider; nasals shorter; auditory bullae more inflated, particularly median tympanic portion; least interorbital width less.

From _Dipodomys ordii nexilis_, _D. o. cupidineus_ differs as follows: Size smaller; color lighter in all pigmented areas; skull smaller; rostrum relatively longer and narrower; least interorbital width less; auditory bullae less inflated; breadth across maxillary arches less; foramen magnum circular as opposed to ovoid; cutting edge of upper incisors narrower; zygomatic arches lighter.

For comparisons with _Dipodomys ordii cinderensis_, _Dipodomys ordii panguitchensis_ and _Dipodomys ordii longipes_ see accounts of those subspecies.

_Remarks._-This subspecies is prevented from intergrading, to the east, with _D. o. longipes_ by the deep canyons of the Colorado River, with _D. o. cinderensis_ by the Virgin River and with _D. o. panguitchensis_ by a series of high mountain ranges. Intergradation does, however, occur with _D. o. sanrafaeli_ in animals from Escalante, Garfield County, Utah, but they are referable to _D. o. cupidineus_. The Colorado River, farther to the north, again serves as a barrier in preventing intergradation between _D. o. nexilis_ and _D. o. cupidineus_.

_Specimens examined._--Total, 122, distributed as follows:

=Utah=: _Garfield County_: Mouth of Calf Creek, Escalante River, 3 (BYU); Ten Mile Spring, 3 (BYU); Escalante, 3 (BYU). _Washington County_: near Short Creek road, south of town of Virgin, 18 (RH). _Kane County_: near Paria, 1 (UU); Willow Tank Springs, 7 (BYU); 8 mi. NW Kanab, 4800 ft., 1 (UU); near Sand Dunes, 7 (RH); Kanab, 15 (12 MVZ; 2 BYU; 1 UU); 1 mi. S Kanab, 4400 ft., 2 (UU).

=Arizona=: _Mohave County_: near S Boundary Kaibab Indian Reservation, 4 (MVZ); 6 mi. N Wolf Hole, 8 (MVZ); 4 mi. N Wolf Hole; 3 mi. NW Diamond Butte, 7 (USBS); 20 mi. S Wolf Hole, 1 (USBS); 1 mi. W Diamond Butte, 1 (USBS); 6 mi. N Mt. Trumbull, 1 (USBS); Nixon Springs, 6250 ft., W Base Mt. Trumbull, 6 (3 USBS; 3 MVZ); 5 mi. S Trumbull Spring, 1 (USBS). _Coconino County_: Fredonia, 3 (USBS); 2 mi. W Lees Ferry, 3 (MVZ); Jacobs Pools, 4000 ft., Houserock Valley, 15 (MVZ); 10 mi. S Jacobs Pools, Houserock Valley, 2 (MVZ); Houserock Valley, 10 (USBS).

=Dipodomys ordii palmeri= (Allen)

_Dipodops ordii palmeri_ Allen, Bull. Mus. Comp. Zool., 8:187, 1881.

_Cricetodipus ordii palmeri_, Trouessart, Catalogus Mammalium, 1:581, 1897.

_Perodipus ordii palmeri_, Goldman, Proc. Biol. Soc. Washington, 30:113, May 23, 1917.

_Dipodomys ordii palmeri_, Grinnell, Journ. Mamm., 2:96, May 2, 1921.

_Cotypes._--Two males, adults, nos. 5886 and 5887, Mus. Comp. Zool.; San Luis Potosí, Mexico; obtained on May 1, 1878, and September 1, 1878, respectively, by Dr. Edward Palmer. (Types not seen.)

_Range._--Eastern Zacatecas, Aguascaliente, northern Jalisco, San Luis Potosí, Hidalgo, and probably Querétaro; marginal occurrences are: Zacatecas, Canitas and Berriozobal; Aguascaliente, 1 mi. N Chicalote; Jalisco, 9 mi. N Encarnación; Guanajuato, Celaya; Hidalgo, Irolo.

_Diagnosis._--Size small (see measurements). Color dark, entire dorsal surface (_h_) between Cinnamon and Sayal Brown, purest on sides and flanks, upper parts strongly suffused with black; posterior surfaces and fold of pinnae of ears, white; arietiform markings, plantar surfaces of hind feet, inside of pinnae of ears, dorsal and ventral stripes of tail, blackish. Skull small; nasals long; rostrum long and narrow; interorbital region relatively wide; degree of inflation of auditory bullae about average for species; zygomatic arches light and bowed laterally; pterygoid fossae subcircular; braincase but slightly vaulted.

_Comparisons._--From _Dipodomys ordii ordii_, _D. o. palmeri_ differs as follows: Size somewhat larger; color darker; skull larger; nasals longer and flaring distally; interorbital width greater; narrower across auditory bullae; interparietal region narrower; breadth across maxillary arches greater; zygomatic arches bowed laterally as opposed to relatively straight; pterygoid fossae more nearly circular; rostrum longer and narrower.

From _Dipodomys ordii sennetti_, _D. o. palmeri_ differs in larger size, darker color, longer tail and longer skull.

For comparisons with _Dipodomys ordii obscurus_ and _Dipodomys ordii fuscus_ see accounts of those subspecies.

_Remarks._--Specimens from Berriozobal, Zacatecas, in the width of the rostrum and the configuration of the nasals, are intermediate between _Dipodomys ordii fuscus_ and _D. o. palmeri_. In color these animals are more nearly like typical representatives of _D. o. palmeri_. The animals from Jalisco and Guanajuato are not typical but the characters differentiating them from _D. o. palmeri_ are merely modifications of such slight degree that they all have been placed with that subspecies.

_Specimens examined._--Total, 56, distributed as follows:

=Zacatecas=: Canitas, 3 (USBS); Berriozobal, 10 (USBS).

=San Luis Potosí=: Potrero Santa Ana, 7.6 mi. S Matehuala, 2 (MVZ); Jesus Maria, 12 (USBS); 2 mi. NW San Luis Potosí, 8 (MVZ).

=Aguascalientes=: 1 mi. N Chicalote, 1900 m., 4 (MVZ).

=Jalisco=: 9 mi. N Encarnación, 1900 m., 1 (MVZ).

=Guanajuato=: Celaya, 8 (USBS).

=Hidalgo=: Ixmiquilpan, 1 (USBS); Irolo, 12 (USBS).

CONCLUSIONS

1. There are thirty-five recognizable subspecies of the species _Dipodomys ordii_ of which four are herein, for the first time, recognized by name. Three subspecies, _D. o. oklahomae_, _D. o. sennetti_ and _D. o. compactus_, previously were regarded by most authors as full species.

2. The species _Dipodomys ordii_ is divisible into six complexes, or groups, of subspecies on both geographic and morphological bases.

3. _Dipodomys ordii_ is the most generalized Recent species of the genus.

4. The extremes of geographic variation in _Dipodomys ordii_ are greater than in any other species of the genus.

5. Color, at least in _Dipodomys ordii_, does not seem to be correlated with amount of moisture but rather with color of soil.

6. Clinal variation, from north to south, is shown in the decreasing length of the nasals. This decrease in length of nasals and resultant decrease in size of the nasal chamber may be correlated with the decrease of humidity of the environment.

7. Subspeciation has been enhanced by the late Quaternary mountain building which was prevalent over the western United States.

8. In general, the most primitive kinds of _Dipodomys ordii_ occur at the periphery of the range of the species.

9. Natural selection plus geographical and ecological isolation has undoubtedly been operative in speciation and in subspeciation.

10. _Dipodomys deserti_ is found to be the most specialized species in the genus.

11. Six, rather than nine, groups of species are recognized, on the basis of morphology, as comprising the genus _Dipodomys_.

12. The center of dispersal for the genus _Dipodomys_ appears to have been in the southwestern United States and the adjoining part of Mexico. A secondary center of differentiation is apparent in the low, hot valleys of central California.

13. Parallel development of species is noted between _Dipodomys_ of the parental center and _Dipodomys_ of the isolated valleys of central California; _Dipodomys_ in the California center is the less specialized.

14. The trend of the dipodomyines, as indicated by the fossil record, has been toward a saltatorial specialization with consequent morphological changes.

15. The morphological change in the direction of saltatorial specialization is clearly evident in the compacting and aligning of the viscera as well as in the lengthening of the distal segments of the hind legs and the tail, tufting of the tail, enlargement of the auditory bullae, shortening of the neck and fusion of the cervical vertebrae for stability and other modifications of the skeleton.

TABLE 7

MEASUREMENTS (IN MILLIMETERS) OF DIPODOMYS ORDII

Key:

A Total length B Length of tail C Length of hind foot D Greatest length of skull E Greatest breadth across bullae F Breadth across maxillary arches G Width of rostrum H Length of nasals I Least interorbital width J Basilar length

=================+=====+=====+====+====+====+====+===+====+====+==== | A | B | C | D | E | F | G | H | I | J -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. richardsoni_ 1-1/2 mi. N Beaver, Oklahoma (KU) [M] Mean (4) |251.2|131.5|40.5|41.0|24.7|22.0|4.3|14.9|13.0|26.3 Maximum |256 |136 |41 |41.8|25.1|22.3|4.5|15.4|13.2|27.3 Minimum |243 |133 |40 |40.4|24.4|21.5|4.2|14.6|12.7|26.0 | | | | | | | | | | [F] KU 17962 |253 |135 |42 |40.6|25.4|22.6|4.6|15.1|14.2|25.7 KU 17963 |246 |133 |40 |39.4|24.0| ...|4.3|14.1|12.5|25.6 KU 17964 |255 |136 |41 |41.2|25.5|22.1|4.5|15.1|13.8|26.2 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. oklahomae_ 2-1/4 mi. S Norman, Oklahoma (USBS) [M] Mean (4) |239.7|130.0|39.5|39.0|23.9|21.5|4.3|13.7|13.3|24.9 Maximum |254 |140 |42 |40.3|24.6|22.4|4.4|14.3|13.6|25.8 Minimum |227 |125 |37 |38.0|23.0|20.9|4.2|13.4|12.8|23.9 | | | | | | | | | | [F] USBS 265456|245 |133 |40 |39.1|24.5|22.0|4.4|14.4|13.5|25.3 OU 20140 |234 |120 |39 |38.7|23.8|21.7|4.4|14.3|13.5|24.8 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. compactus_ 19 mi. S Port Aransas, Mustang Island, Texas (TCWC) [M] Mean (10) |234.6|119.5|39.3|37.0|21.8|19.4|4.1|13.8|11.9|24.2 Maximum |251 |134 |41 |37.9|22.5|20.8|4.2|14.6|12.3|25.3 Minimum |228 |112 |38 |35.6|21.3|18.2|3.9|12.9|11.6|22.8 | | | | | | | | | | [F] Mean (7) |231.0|116.0|39.0|37.1|22.4|19.7|4.0|13.9| ...|24.2 Maximum |235 |120 |41 |38.2|22.9|20.5|4.1|14.7| ...|24.8 Minimum |224 |110 |38 |36.0|21.7|19.1|3.8|13.2| ...|23.8 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. sennetti_ 2 mi. S Riviera, Texas (TCWC) [M] Mean (5) |217.6|112.2|35.8|37.2|23.4|20.1|4.0|13.6|13.1|24.2 Maximum |222 |115 |38 |38.2|24.1|20.7|4.3|14.4|13.2|24.6 Minimum |208 |104 |34 |36.3|23.0|19.4|3.8|13.0|12.6|23.8 | | | | | | | | | | [F] Mean (4) |218.3|112.0|36.0|37.2|23.3|20.1|4.0|13.9|13.1|24.1 Maximum |226 |115 |37 |37.7|23.9|20.8|4.1|14.4|13.4|24.4 Minimum |209 |108 |35 |36.7|22.8|19.0|3.9|13.3|12.6|23.5 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. evexus_ Salida, Colorado (AMNH) [M] AMNH 28802 |267.0|140.0|42.0| ...|23.0| ...|4.2| ...| ...|25.0 AMNH 28804 |271 |156 |43 |39.3|23.5|22.1|4.4|14.6|14.0|25.8 AMNH 28805 |252 |137 |39 | ...| ...|21.5|4.2| ...| ...|24.5 | | | | | | | | | | [F] Mean (4) |261.0|147.0|42.2|38.0|23.5|21.6|4.3|14.1|13.4|24.6 Maximum |264 |149 |43 |38.1|24.0|22.1|4.4|14.2|13.8|25.0 Minimum |258 |144 |42 |37.9|22.7|20.8|4.2|14.0|13.0|24.3 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. medius_ Santa Rosa, New Mexico (USBS) [M] Mean (6) |258.5|143.3|38.0|39.5|25.2|21.4|4.1|14.2|12.9|24.8 Maximum |266 |151 |40 |40.6|25.9|22.5|4.2|15.2|13.1|25.6 Minimum |251 |132 |37 |38.9|24.8|20.6|4.0|13.5|12.8|24.1 | | | | | | | | | | [F] USBS 118527|261.0|145.0|40.0|40.4|25.4|22.2|4.5|15.7|13.5|25.7 USBS 127310| ...| ...| ...|38.1|24.3|20.8|4.1|13.5|13.5| ... -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. obscurus_ Rio Sestin, Durango (AMNH) [M] Mean (6) | ...| ...| ...|36.3|22.9|19.7|3.7|13.1|12.6|22.8 Maximum | ...| ...| ...|38.0|23.4|21.0|3.9|13.5|13.0|23.9 Minimum | ...| ...| ...|35.2|22.6|19.1|3.6|11.9|12.4|21.7 | | | | | | | | | | [F] AMNH 20945 | ...| ...| ...|36.5|22.8|19.9|3.8|13.1|12.9|22.3 AMNH 20951 | ...| ...| ...|36.0|22.3|19.6|3.5|13.0| ...|22.5 AMNH 20958 | ...| ...| ...|35.9|22.7|19.4|3.4|12.6|12.3|22.7 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. terrosus_ Jordan, Montana [M] AMNH 41442 |280.0|155.0|44.0|42.7|26.5|23.6|4.0|15.2|14.5|27.0 AMNH 41443 |267.0|155.0|40.0|40.5|24.8|21.1|4.0|14.5|13.1|25.1 AMNH 41444 |279.0|162.0|41.0|40.8|25.7|21.6|4.4|14.6|13.4|25.9 | | | | | | | | | | [F] AMNH 41441 |265.0|149.0|40.5|41.4|25.4|22.2|4.3|14.7|13.0|26.8 MVZ 25658 |273.0|154.0|41.0|41.3|25.1|22.4|4.0|14.9|13.8|26.0 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. fremonti_ Torrey, Utah [M] CM 15663 |250.0|141.0|40.0|37.0|23.3|19.3|3.6|13.5|12.0|23.2 CM 15670 |248.0|136.0|38.0|37.3|23.8|19.8|3.8|13.7|11.6|23.5 | | | | | | | | | | [F] CM 15666 |258.0|141.0|39.0|37.5|23.5|19.9|3.7|13.8|11.9|23.5 CM 15667 |252.0|140.0|39.0|36.8|23.5|20.5|3.7|13.5|11.5|23.2 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. uintensis_ 2 mi. N Fruitland, Utah [M] CM 11634 |253.0|140.0|40.0|37.9|23.3|20.3|3.9|13.5|12.5|23.9 CM 11640 |260.0|150.0|41.0|38.2|24.2|20.6|4.0|13.2|12.5|24.5 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. sanrafaeli_ 12 mi. SW Green River, Utah [M] CM 15649 |265.0|144.0|41.0| ...| ...| ...|...|14.1| ...| ... | | | | | | | | | | [F] CM 15647 |253.0|138.0|42.0| ...| ...| ...|...|13.5| ...| ... -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. panguitchensis_ 1 mi. S Panguitch, Utah [M] RH 2151 |257.0|145.0|41.0| ...| ...| ...|...|13.3|12.0| ... RH 2152 |252.0|135.0|40.0| ...| ...| ...|...|13.5|11.9| ... | | | | | | | | | | [F] RH 2153 |240.0|132.0|38.0| ...| ...| ...|...|12.7|11.3| ... -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. monoensis_ 5 mi. N Benton Station, California (MVZ) [M] MVZ 26993 |232.0|125.0|40.0|36.6|23.7|20.0|3.7|13.0| ...|23.4 MVZ 26995 |240.0|125.0|39.0|36.8|23.0|19.8|3.6|13.4|11.5|23.3 MVZ 26997 |230.0|122.0|39.0|37.2|23.2|20.1|3.6|13.4|12.0|23.9 | | | | | | | | | | [F] Mean (5) |228.0|125.4|38.4|36.6|23.1|19.7|3.5|13.1|11.8|23.4 Maximum |240.0|137.0|39.0|37.2|24.2|20.0|3.7|13.2|12.2|23.6 Minimum |220.0|120.0|38.0|36.0|22.6|19.3|3.4|13.0|11.5|23.0 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. ordii_ Near El Paso, Texas (USNM) [M] Mean (5) | ...| ...| ...|37.1|24.1|19.6|3.7|13.0|12.7|23.5 Maximum | ...| ...| ...|38.1|24.8|20.5|3.8|13.2|13.3|24.3 Minimum | ...| ...| ...|36.3|23.4|18.8|3.5|12.8|12.1|22.4 | | | | | | | | | | [F] Mean (6) |235.2|128.0|37.4|37.3|24.0|19.6|3.7|13.3|12.8|23.5 Maximum |245.0|136.0|39.0|38.0|24.1|20.0|3.8|13.5|13.4|24.4 Minimum |222.0|120.0|35.0|36.3|24.0|19.1|3.6|13.0|12.4|23.0 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. luteolus_ 1 mi. NE Casper, Wyoming (KU) [M] Mean (12) |265.6|152.2|42.2|38.9|24.1|20.8|4.3|13.9|13.0|24.6 Maximum |281.0|163.0|43.0|39.5|25.0|22.1|4.4|14.5|13.7|25.7 Minimum |254.0|145.0|42.0|37.5|23.8|19.9|4.2|13.0|12.5|24.0 | | | | | | | | | | [F] Mean (7) |260.7|148.0|41.0|38.6|24.2|20.9|4.3|13.9|12.9|24.7 Maximum |269.0|153.0|43.0|40.5|25.7|21.9|4.4|14.9|13.8|25.5 Minimum |250.0|139.0|40.0|37.6|23.0|20.0|4.2|13.3|12.5|24.0 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. extractus_ 1 mi. E Samalayuca, Chihuahua (MVZ) [M] Mean (11) |235.1|128.2|37.3|37.5|24.1|20.1|3.5|13.2|13.0|23.4 Maximum |251.0|142.0|39.0|38.6|24.6|20.8|3.8|13.8|13.5|24.4 Minimum |224.0|121.0|35.0|36.2|23.2|19.3|3.3|12.6|12.4|22.3 | | | | | | | | | | [F] MVZ 76568 |251.0|132.0|39.0|39.6|25.4|22.2|3.7|13.8|14.1|24.8 MVZ 76569 |244.0|140.0|37.0|37.3|23.6|20.0|3.6|13.5|12.8|24.0 MVZ 76570 |230.0|120.0|37.0|35.7|23.4|19.5|3.2|12.4|12.1|22.5 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. chapmani_ Camp Verde, Arizona (USBS) [M] Mean (6) |258.0|148.0|39.0|38.9|24.2|21.1|3.8|14.2|13.5|24.3 Maximum |268.0|157.0|41.0|40.1|24.7|22.1|3.9|14.8|13.9|24.7 Minimum |244.0|138.0|38.0|38.1|23.6|20.5|3.8|13.3|13.1|24.1 | | | | | | | | | | [F] Mean (4) |255.4|151.0|38.2|37.1|23.9|20.1|3.8|13.4|13.5|23.3 Maximum |262.0|152.0|39.0|37.7|24.3|20.1|3.9|13.7|14.9|23.5 Minimum |245.0|144.0|37.0|36.0|23.7|20.1|3.6|13.1|12.6|23.0 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. montanus_ Fort Garland, Colorado (USBS) [M] Mean (11) |252.0|140.5|41.0|37.7|24.2|20.1|4.1|13.6|12.8|23.8 Maximum |263.0|150.0|42.0|39.0|25.0|21.0|4.5|14.0|13.2|24.7 Minimum |233.0|131.0|39.0|36.6|23.6|19.0|3.9|12.7|12.4|23.0 | | | | | | | | | | [F] Mean (11) |256.0|141.0|40.8|37.2|23.9|19.6|4.1|13.4|12.5|23.7 Maximum |259.0|145.0|42.0|38.4|24.4|21.4|4.3|14.1|13.3|24.4 Minimum |237.0|132.0|40.0|36.0|23.0|18.2|3.9|12.3|11.9|22.6 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. cinderensis_ 11 mi. SE Lund, Utah [M] MVZ 102059 |244.0|135.0|41.0|35.7|23.2|19.2|3.6|12.2|11.2|22.7 MVZ 102056 |237.0|130.0|41.0|36.4|23.7|20.6|3.5|13.0|12.1|23.1 | | | | | | | | | | [F] MVZ 102057 |229.0|122.0|41.0|36.3|23.5|20.2|3.6|13.0|11.5|23.4 MVZ 102058 |241.0|127.0|40.0|36.8|24.0|20.0|3.6|13.2|11.7|23.2 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. fetosus_ 2 mi. N Panaca, Nevada (MVZ) [M] Mean (4) |242.5|133.7|42.3|37.4|23.7|20.0|3.7|13.3|11.7|23.7 Maximum |249.0|140.0|43.0|37.7|24.0|20.3|3.8|13.9|12.0|23.8 Minimum |233.0|126.0|42.0|36.8|23.2|19.5|3.7|12.7|11.5|23.6 | | | | | | | | | | [F] Mean (4) |229.0|125.5|40.6|35.8|23.3|19.5|3.6|12.8|11.3|22.5 Maximum |235.0|129.0|41.5|37.0|24.1|20.0|3.6|13.3|11.6|23.2 Minimum |224.0|122.0|40.0|34.3|23.0|19.0|3.6|11.9|11.0|21.9 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. utahensis_ Ogden, Utah [M] MVZ 44005 |243.0|133.0|40.5|35.9|23.1|19.2|3.5|12.8|12.4|22.4 | | | | | | | | | | [F] MVZ 44006 |238.0|133.0|38.0|36.1| ...|19.8|3.7|13.5|12.2|22.9 MVZ 44007 |240.0|137.0|39.0|35.6|22.9| ...|3.6|12.8|11.7|22.4 MVZ 44008 |241.0|134.0|40.0|36.4|23.5|19.8|3.6|13.3|12.3| ... -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. columbianus_ Umatilla, Oregon [M] MVZ 45317 |243.0|140.0|40.0|37.2|23.6|20.0|3.8|13.5|12.2|23.4 MVZ 45318 |236.0|137.0|40.0|36.8|23.5|20.1|3.6|13.2| ...|23.4 MVZ 45322 |240.0|132.0|40.0|36.4|23.4|19.9|3.7|13.0|12.1|23.3 | | | | | | | | | | [F] MVZ 45314 |239.0|130.0|39.0|35.8|22.7|19.8|3.7|13.2|11.9|22.7 MVZ 45315 |251.0|140.0|41.0|36.6|23.8|19.6|3.6|13.3|12.0|23.1 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. idoneus_ 12 mi. W Lerdo, Durango [M] UM 90027 | ...| ...|35.0|37.3|24.3|20.7|3.8|13.0|14.1|23.1 UM 90029 |230.0|129.0|35.0|37.2|23.5|20.0|3.6|13.5|13.4|23.3 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. priscus_ 33 mi. S Bitter Creek, Wyoming (KU) [M] Mean (7) |259.0|148.0|44.0|39.1|24.3|20.7|4.1|14.3|13.1|24.9 Maximum |265.0|152.0|45.0|40.4|25.1|21.2|4.3|15.2|13.6|25.5 Minimum |251.0|144.0|43.0|38.0|23.7|20.0|4.0|13.8|12.7|23.7 | | | | | | | | | | [F] Mean (4) |257.0|147.0|43.0|39.4|24.6|20.8|4.2|14.3|13.1|24.7 Maximum |264.0|152.0|45.0|40.4|25.2|21.9|4.3|14.9|13.3|25.2 Minimum |249.0|138.0|40.0|38.1|23.5|20.1|4.1|14.0|12.7|24.0 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. celeripes_ Trout Creek, Utah (UU) [M] Mean (4) |208.7|109.0|40.2|35.3|22.9|19.3|3.6|12.6|11.2| ... Maximum |220.0|115.0|41.0|36.6|23.6|20.2|3.8|13.5|11.5| ... Minimum |203.0|100.0|39.0|34.0|22.1|18.4|3.5|11.7|11.1| ... | | | | | | | | | | [F] UU 1957 |219.0|110.0|39.0|35.3|23.4|19.1|3.6|11.9|11.8|22.4 UU 1961 |223.0|120.0|40.0|37.0|23.6|20.4|3.6|13.1|12.1|23.9 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. cineraceus_ Dolphin Island, Great Salt Lake, Utah [F] USNM 263893|228.0|129.0|39.0|37.2|23.4|19.7|3.5|13.6|11.6|23.0 USNM 263894|230.0|132.0|38.0|37.1|23.7|20.1|3.6|13.4|11.9|23.6 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. marshalli_ Stansbury Iland, Great Salt Lake, Utah [M] UU 2968 |238.0|128.0|40.0|36.4|23.5| ...|3.5|13.0|12.0|23.2 UU 2969 |241.0|136.0|40.0|36.3|23.7|19.2|3.7|12.9| ...|23.0 | | | | | | | | | | [F] UU 2972 | ...| ...| ...|35.9|23.5|19.9|3.6|12.9|11.9|23.1 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. inaquosus_ 11 mi. E and 1 mi. N Jungo, Nevada (MVZ) [M] Mean (4) |243.0|137.0|40.0|36.6|23.5|19.8|3.7|13.5|11.8|23.1 Maximum |247.0|140.0|40.5|36.8|24.0|20.0|3.8|13.8|12.1|23.5 Minimum |238.0|131.0|39.0|36.4|23.3|19.5|3.6|13.1|11.5|23.0 | | | | | | | | | | [F] MVZ 73577 |242.0|139.0|40.5|36.4|23.9|19.2|3.7|13.2|11.4|22.9 MVZ 73579 |231.0|130.0|40.5|36.3|23.2|19.0|3.5|12.7|11.0|23.2 MVZ 73582 |254.0|138.0|41.0|38.2|23.9|20.0|3.6|13.9|12.2| ... -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. attenuatus_ Johnson's Ranch, Texas [M] TCWC 3633 |235.0|126.0|41.0|37.3|24.1|20.1|3.7|13.7| ...|23.4 TCWC 3634 |237.0|136.0|37.0|35.5|22.1|18.8|3.5|13.2|12.2|22.3 UM 79121 |245.0|143.0|37.3|35.7|22.3|19.1|3.6|12.4|12.9|23.3 | | | | | | | | | | [F] UM 79122 |238.0|140.0|38.0|34.6|22.5|18.5|3.5|11.9|13.0|21.9 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. fuscus_ Juamave, Tamaulipas (USBS) [M] Mean (4) |248.0|149.0|38.5|37.6|24.1|20.2|3.6|12.6|12.7|23.6 Maximum |254.0|152.0|40.0|38.3|24.7|20.4|3.7|13.1|12.9|23.7 Minimum |241.0|146.0|38.0|37.0|23.5|19.7|3.6|12.3|12.5|23.6 | | | | | | | | | | [F] Mean (4) |244.7|146.2|38.5|37.4|24.1|19.9|3.4|12.9|12.7|23.2 Maximum |254.0|155.0|40.0|37.7|24.5|20.5|3.6|13.0|12.9|23.4 Minimum |240.0|138.0|38.0|37.2|23.7|19.2|3.3|12.7|12.4|23.2 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. longipes_ Kayenta, Arizona [M] USBS 247915|255.0|145.0|41.0|39.3|25.5|20.7|3.9|13.8|13.0|24.3 USBS 247916|242.0|138.0|40.0|38.4|25.4|20.6|3.8|13.3|12.7|24.6 | | | | | | | | | | [F] USBS 247552|255.0|135.0|40.0|39.9|25.7|21.2|4.0|13.8| ...|25.4 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. pallidus_ 18 mi. SW Orr's Ranch, Utah (UU) [M] Mean (4) |236.6|131.3|42.3|37.6|24.2|20.4|3.8|13.4|11.8|23.6 Maximum |240.0|134.0|43.0|37.9|24.6|20.9|4.0|13.6|12.1|24.2 Minimum |230.0|128.0|41.0|37.0|23.7|19.8|3.7|12.9|11.5|23.4 | | | | | | | | | | [F] UU 3528 |229.0|132.0|40.0|36.6|23.5|20.0|3.7|12.5|12.0| ... -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. nexilis_ 5 mi. W Naturita, Colorado [M] USBS 149940|261.0|156.0|44.0|38.7|25.1|18.9|4.1|13.8|11.4|24.6 | | | | | | | | | | [F] USBS 149941|265.0|142.0|45.0|40.5|26.3|21.5|4.2|14.4|13.1|25.3 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. cupidineus_ Houserock Valley, Arizona (USBS) [M] Mean (6) |239.0|138 0|39.8|36.9|24.1|19.3|3.8|12.8|12.0|23.6 Maximum |244.0|146.0|41.0|37.5|24.9|20.1|4.0|13.0|12.6|24.3 Minimum |225.0|130.0|38.5|36.3|23.5|18.7|3.6|12.6|11.7|23.2 | | | | | | | | | | [F] Mean (4) |244.7|141.0|40.0|37.4|24.2|19.7|3.8|13.2|12.3|23.9 Maximum |247.0|147.0|40.5|37.7|24.4|20.0|3.9|13.5|12.6|24.6 Minimum |240.0|137.0|39.5|37.1|23.9|19.5|3.7|12.9|12.0|23.7 -----------------+-----+-----+----+----+----+----+---+----+----+---- _D. o. palmeri_ Jesus María, San Luis Potosí (USBS) [M] Mean (6) |242.0|141.6|37.6|36.7|23.6|20.0|3.3|12.9|12.9|22.6 Maximum |255.0|157.0|39.0|37.3|24.4|20.8|3.6|13.6|13.4|23.0 Minimum |229.0|129.0|35.0|35.9|23.2|19.4|3.0|12.5|12.6|22.0 | | | | | | | | | | [F] Mean (6) |239.5|139.6|37.1|36.4|23.8|19.9|3.4|12.8|13.1|22.7 Maximum |252.0|148.0|38.0|37.3|23.9|20.0|3.6|13.3|13.2|23.5 Minimum |225.0|128.0|36.0|35.9|23.7|19.5|3.3|12.2|13.0|22.0 -----------------+-----+-----+----+----+----+----+---+----+----+----

LITERATURE CITED

ANDERSON, R. M. 1947. Catalogue of Canadian Recent mammals. National Museum of Canada, Bull. 102, Biol. Ser., 31, pp. v + 238, January 24, 1947.

BAILEY, V. 1905. Biological survey of Texas. N. Amer. Fauna, 25:1-222, 24 figs., Government Printing Office, October 24, 1905.

BENSON, S. B. 1933. Concealing coloration among some desert rodents of the southwestern United States. Univ. California Publ. Zool., 40:1-70, 2 pls., 8 figs. in text, June 13, 1933.

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_Transmitted May 15, 1948._

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Transcriber's note:

Gesperrt text is indicated by ~swung dashes~.