Physiological Researches on Life and Death
CHAPTER IX.
OF THE ORIGIN AND DEVELOPMENT OF THE ORGANIC LIFE.
We have just now seen that the animal life, which is inactive in the fœtus, is developed after birth: we have also followed up the particular laws of its development. On the contrary, the organic life comes into action almost as soon as the fœtus is conceived; for as soon as the least organization is apparent, the little heart will be seen protruding its blood on all sides. The heart is the first formed part, the first in action: now, as all the organic phenomena depend upon it, we may readily conceive in what way the functions of the inward life are thrown into exercise.
I. _Of the mode of the organic life in the fœtus._
Nevertheless, the organic life of the fœtus, is not the same as that which the adult is destined to enjoy. Let us enquire into the reason of this difference.
We have said that the organic life is the result of two great orders of functions, of those namely of assimilation and decomposition, so as to form an habitual circle of creation and destruction. Now in the fœtus this circle is singularly contracted.
For in the first place, the functions of assimilation are much fewer in number; the molecules before they arrive within the organs which they are destined to create, are not submitted to so many actions; they penetrate the fœtus already elaborated by the digestion, circulation, and respiration of the mother. Instead of traversing the apparatus of the digestive organs, which at this age appear to be almost inactive, they enter at once into the system of the circulation; the road which they have travelled is less, it is not requisite that they should be presented to the influence of respiration; and accordingly the fœtus of the mammalia has in its preliminary organization a near analogy with that of the adult reptile, in which but a small part of the blood at its issuing from the heart, is sent into the vessels of the lungs.[48]
The molecules of nourishment in this way pass almost directly from the circulating torrent into the nutritive system. The general process of assimilation, then, is much less complicated than that of the following age.
On the other hand, those functions which habitually decompose the organs, which clear the system of substances already become injurious and foreign to its nature, are at this age but very inactive. Neither the pulmonary exhalation, nor sweating, nor transpiration have as yet commenced: the bile, urine, and saliva are but small in quantity, if compared with what they are destined at a future time to be, so that the portion of blood from which they are to be made in the adult, in the fœtus is almost entirely expended on the system of the nutritive organs.
The organic system of the fœtus, then, is remarkable--on the one hand, for the extreme promptitude of its assimilation, a promptitude depending on the very small number of the functions concurring to that end; and on the other, for the extreme inertia of its decomposition, an inertia depending on the little activity of the different functions, which are the agents of this great process.
It is easy from the foregoing considerations to account for the rapidity which characterizes the growth of the fœtus; a rapidity which is manifestly out of all proportion with that which takes place at any other age. Indeed, while every thing is in favour of the progression of the nutritive matter towards the parts where it is destined to be put down, every thing at the same time seems to oblige such matter to remain in the place where it has been deposited, the emunctories of the system being wanting.
To the great simplicity of assimilation in the fœtus, we may add the great activity of the organs which contribute to it an activity, which depends upon the more considerable sum of vital power which they then partake. All the powers of the economy, indeed, appear to be concentrated upon the system of the circulation and nutrition; the functions of digestion, respiration, secretion and exhalation, are exercised but obscurely.[49]
If we now observe that the organs of the animal life, which are condemned to a necessary inaction, are the seat at the same time of a very small portion only of vital power (the surplus of this being thrown upon the organic life) it will be easy to perceive, that almost the whole of the powers which are afterwards to be developed upon the two systems in general, will be then concentrated upon those which serve to nourish and compose the different parts of the fœtus, and that in consequence the functions which concur to the process of nutrition and growth, must at that age be the seat of an extreme energy.
II. _Development of the organic life after birth._
Immediately after birth, the organic life of the child has a great addition made to it; its extent is almost doubled, for not only are many of the functions which did not before exist at such time added, but those which existed previously are much enlarged. Now in this remarkable revolution of things, a law directly the contrary of that which presides over the animal life is observed; for the organs of this life, whether they be newly brought into exercise, or simply receive an increase of action, need no education; they suddenly attain to a perfection, which those of the animal life do not acquire, otherwise than by long habitude. A rapid glance upon the development of this life, will be sufficient to convince us of the truth of the above observation.
At the instant of birth, digestion and respiration, with a great part of the exhalations and absorptions commence. Now after the first inspirations and expirations; after the elaboration in the stomach of the first milk, which is taken in by the infant, as soon as the exhalants of the lungs and the skin have once rejected some small portions of their respective fluids, the respiratory, the digestive and exhalant organs, have as perfect an action as they ever will have.
At the same time all the glands, which slept as it were, which poured forth but a very small quantity of fluid, are awakened from their torpor by the stimuli of the various substances which are applied to the mouths of their excretory ducts. The passage of the milk at the extremities of the stenonian and wartonian ducts, of the chyme at the end of the choledochus and the pancreatic duct, the contact of air with the orifice of the urethra, awaken into action the salivary glands, the pancreas, the liver, and the kidneys. The air in like manner upon the inner surface of the trachea and the nostrils, and the aliments upon that of the digestive passages, are the excitants which rouse these parts into action.[50]
It is then also that begin the various excretions of the system: now if we examine well the different organs which concur to the above mentioned phenomena, we shall find that they require no sort of education.
I shall not inquire into the reason of this difference in the development of the two lives. I shall only observe that it is out of the power of any one of the inward organs, to acquire a marked degree of superiority over any other, for the same reason that they all of them attain, immediately upon entering into action, as great a perfection as at any time they are destined to possess.
Nevertheless there is nothing more common than the predominance of one system of the organic life over the other systems; this is sometimes the vascular, sometimes the pulmonary apparatus, at other times the organs of digestion, and the liver especially, have the greater degree of development, and decide on the particular temperament of the individual; but the cause of this sort of constitution depends on primitive organization, on the structure of the parts, on their conformation. Such superiority is by no means the effect of exercise or habit, for the fœtus and the child display the same phenomena, in as much reality though less apparently indeed, than adolescence, or manhood.
In the same way, the debility of any particular system of the internal functions, may depend either on original constitution, or on some accidental vice or disease, by which, while the others have remained untouched, its constitution may have been impaired.
Such then is the great difference of the two lives of the animal, with respect to inequality of perfection in the organs. In the animal life, the predominance or inferiority of one system, with relation to the others, depends almost entirely upon its activity or inertia, on its habitude of acting or not acting. In the organic life on the contrary, such states are immediately connected with the texture of the organs, and never with their education.
From hence also we have the reason why physical temperament, and moral character, are not susceptible of change from education, which so prodigiously modifies the actions of the animal life, for as we have seen, they both of them belong to the organic life.
The character is, if I may so express myself, the physiognomy of passions; temperament, that of the internal functions: now the one and the other being at all ages the same, having a direction which habitude and exercise can never alter, it is manifest that they must ever be withdrawn from the influence of education. The violence of the temperament may indeed be moderated, for the powers of the judgment, and reflection may be augmented, and the animal life strengthened in such way as to give it a capacity of resisting the impulses of the organic life; but to attempt an immediate alteration of the character, or of the passions, which are its habitual expressions, is an enterprise analogous to that of the physician, who should attempt to elevate or depress, (and that, for the entire life of the patient,) the ordinary contracting powers of the heart and arteries.
We should observe to such physician, that the circulation and respiration, are not under the dominion of the will; and that they cannot be modified excepting in passing into a state of disease. The same observation might be made to those, who imagine that the character, and consequently the passions may be modified.
FOOTNOTES:
[48] I am persuaded that the still very obscure theory of the fœtus might be elucidated by that of animals who have a similar organization. For example, in the frog, in whom but little blood goes through the lungs, the heart is a simple organ, with a single auricle and ventricle; there is a communication or rather continuity between the two systems, venous and arterial, whilst in the mammalia, the vessels in which the red blood circulates do not communicate with those which carry the black blood, except it be by the capillaries.
In the fœtus, the foramen ovale and the ductus arteriosus also render very evidently the arteries and veins continuous; in the fœtus the heart is likewise a simple organ, not forming, notwithstanding its partitions, but one cavity whilst it is double after birth. The two kinds of blood mix at this period, as in reptiles. Now, I shall prove hereafter, that in the child who has breathed, this mixture would soon be fatal; that the black blood, circulating in the arteries, would very quickly produce asphyxia in the animal. Whence arises then this difference? It cannot be studied in the fœtus; it is necessary perhaps to search for it in frogs, salamanders and other reptiles, which can, by their organization, be a long time deprived of air without dying, a phenomenon which approximates them to the mammalia while living in the womb of the mother. Till these very important researches are made, the history of respiration will be incomplete.
[49] This explanation is no doubt ingenious, but it is insufficient, since the causes which Bichat assigns for the rapidity of the growth of the fœtus cease entirely at the moment of birth, and yet the growth continues for a long time after to be as rapid.
[50] When two phenomena are seen to follow each other immediately, we are naturally led to consider one as the cause of the other. _Post hoc ergo propter hoc._ It is a form of reasoning which is very often abused. Food taken into the mouth touches the orifice of the salivary ducts, the fluid flows out, and it is then concluded that the salivary gland has been excited by the impression made on the extremity of its canal. At the moment of birth, the orifice of the urethra is exposed to the contact of the air, and soon the kidneys begin to secrete; then it is the impression of the air on the urethra that has produced their action. But is not this contact of the food in the one case, and of the air in the other an accidental and purely accessory circumstance? Do we believe, that if by any cause the opening of the prepuce was entirely obliterated, the secretion of urine would be prevented? Do we not know that if instead of taking into the mouth savoury food, it is brought near to it, the saliva flows not less, or in vulgar language the mouth waters? There is however no contact, there is not any mechanical or chemical impression in the orifice of the salivary ducts.