Physiological Researches on Life and Death
CHAPTER II.
GENERAL DIFFERENCES OF THE TWO LIVES WITH REGARD TO THE OUTWARD FORM OF THEIR RESPECTIVE ORGANS.
The organs of the animal life are symmetrical, those of the organic life irregular in their conformation; in this circumstance consists the most essential of their differences. Such character, however, to some animals, and among the fish, to the sole and turbot especially, is not applicable; but in man it is exactly traced, as well as in all the genera which are nearest to him in perfection. In them alone am I about to examine it.
I. _Symmetry of the external forms of the animal life._[6]
Two globes in every respect the same, receive the impressions of light. Sounds and odours, have also their double analogous organ. A single membrane is affected to savours, but the median line is manifest upon it, and the two segments, which are indicated by it, are exactly similar. This line indeed is not every where to be seen in the skin, but it is every where implied. Nature, as it were, has forgotten to describe it, but from space to space she has laid down a number of points, which mark its passage. The cleft at the extremity of the nose, of the chin, and the middle of the lips, the umbilicus, the seam of the perineum, the projection of the spinous apophyses of the back, and the hollow at the posterior part of the neck are the principal points at which it is shewn.
The Nerves, which transmit the impressions received by the senses, are evidently assembled in symmetrical pairs.
The brain, the organ (on which the impressions of objects are received) is remarkable also for the regularity of its form. Its double parts are exactly alike, and even those which are single, are all of them symmetrically divided by the median line.
The Nerves again, which transmit to the agents of locomotion and of the voice, the volitions of the brain, the locomotive organs also, which are formed in a great degree of the muscular system, of the bony system, and its dependencies, these together with the larynx and its accessories, composing the double agents of volition, have all of them a regularity, a symmetry, which are invariable.
Such even is the truth of the character which I am now describing, that the muscles and the nerves immediately cease to be regular, as soon as they cease to appertain to the animal life. The heart, and the muscular fibres of the intestines are proofs of this assertion in the muscles; in the nerves, the great sympathetic, is an evidence of its truth.
We may conclude then from simple inspection, that Symmetry is the essential character of the organs of the animal life of man.
II. _Irregularity of the exterior forms of the organic life._
If at present we pass to the viscera of the organic life, we shall perceive a character directly the contrary of the former. The stomach, the intestines, the spleen, the liver, &c. are all of them irregularly disposed.
In the system of the circulation, the heart and the large vessels, such as the upper divisions of the aorta, the vena azygos, the vena portæ, and the arteria innominata have no one trace of symmetry. In the vessels of the extremities continual varieties are also observed, and when they occur, it is particularly remarkable that their existence on one side in no way affects the other side of the body.
The apparatus of respiration appears indeed at first to be exactly regular; nevertheless, the bronchi are dissimilar in length, diameter, and direction; three lobes compose one of the lungs, two the other: between these organs also, there is a manifest difference of volume; the two divisions of the pulmonary artery resemble each other neither in their course, nor in their diameter; and the mediastinum is sensibly directed to the left. We shall thus perceive that symmetry is here apparent only, and that the common law has no exception.
The organs of exhalation and absorption, the serous membranes, the thoracic duct, the great right lymphatic vessel, and the secondary absorbents of all the parts have a distribution universally unequal and irregular.
In the glandular system also we see the crypts, or mucous follicles disseminated in a disorderly manner in every part; the pancreas, the liver, the salivary glands themselves, though at first sight more symmetrical, are not exactly submitted to the median line; added to this, the kidneys differ from each other in their situation, in the length and size of their artery and vein, and in their frequent varieties more especially.[7]
From considerations so numerous we are led to a result exactly the reverse of the preceding one; namely, that the especial attribute of the organs of the interior life is irregularity of exterior form.
III. _Consequences resulting from the difference of exterior form in the organs of the two lives._
It follows from the preceding description, that the animal life is as it were double; that its phenomena performed as they are at the same time on the two sides of the body, compose a system in each of them independent of the opposite system; that there is a life to the right, a life to the left; that the one may exist, the other ceasing to do so, and that they are doubtless intended reciprocally to supply the place of each other.
The latter circumstance we may frequently observe in those morbid affections so common, where the animal sensibility and mobility are enfeebled, or annihilated on one side of the body, and capable of no affection whatever; where the man on one side is little more than the vegetable, while on the other he preserves his claim to the animal character. Undoubtedly those partial palsies, in which the median line, is the limit where the faculties of sensation and motion finish, and the origin from whence they begin can never be remarked so invariably in animals, which, like the oyster, have an irregular exterior.
On the contrary the organic life is a single system, in which every thing is connected and concatenated; where the functions on one side cannot be interrupted, and those on the other subsist. A diseased liver influences the state of the stomach; if the colon on one side cease to act, that upon the other side cannot continue in action: the same attack, which arrests the circulation in the right side of the heart, will annihilate it also in the left side of the heart. Hence it follows, the internal organs on one side being supposed to suspend their functions, that those on the other must remain inactive, and death ensue.
This assertion, however, is a general one; it is only applicable to the sum of the organic life, and not to its isolated phenomena. Some of them in fact are double, and their place may be supplied--the kidneys and lungs are of this description.
I shall not enquire into the cause of this remarkable difference, which in man, and those animals which approach him the nearest, distinguishes the organs of the two lives. I shall only observe, that it enters essentially into the nature of their phenomena, and that the perfection of the animal functions is so connected with the general symmetry observed in their respective organs, that every thing which troubles such symmetry, will more or less impair the functions.
It is from thence, no doubt, that proceeds this other difference of the two lives, namely, that nature very rarely varies the usual conformation of the organs of the animal life. Grimaud has made this observation, but has not shewn the principle on which it depends.
It is a fact, which cannot have escaped any one the least accustomed to dissection, that the spleen, the liver, the stomach, the kidneys, the salivary glands, and others of the internal life, are frequently various in form, size, position, and direction. Such in the vascular system are these varieties, that scarcely will any two subjects be found exactly alike under the scalpel of the anatomist: the organs of absorption, the lymphatic glands in particular, are rarely the same either in number or volume, neither do the mucous glands in any way affect a fixed and analogous situation.
And not only is each particular system subject to frequent aberrations, but the whole of the organs of the internal life are sometimes found in the inverse of the natural order. Of this I have lately seen an instance.
Let us now consider the organs of the animal life, the senses, the brain, the voluntary muscles, and the larynx: here every thing is exact, precise, and rigourously determined. In these there is scarcely ever seen a variety of conformation; if there do exist any, the functions are troubled, disturbed, or destroyed: they remain unaltered in the organic life, whatever may be the disposition of the parts.
The difference with respect to action, in the organs of the two lives, depends, undoubtedly, upon the symmetry of the one, whose functions the least change of conformation would have disturbed, and on the irregularity of the other, with which these different changes very well agree.
The functions of every organ of the animal life are immediately connected with the resemblance of the organ to its fellow on the opposite side if double, or if single to its similarity of conformation in its two halves: from hence the influence of organic changes upon the derangement of the functions may be well conceived.
But this assertion will become more sensible, when I shall have pointed out the relations which exist between the symmetry and the irregularity of the organs, and the harmony and the discordance of their functions.
FOOTNOTES:
[6] It is rather to the external forms that symmetry appears to have been primitively attached, and it is in some measure accidentally and because the nature of their functions requires in general that they should be placed on the exterior, that the organs of relation are found modified in virtue of this law. In the example cited, of fishes without a bladder, the eyes, to lose nothing of their utility, must be differently placed, and on the face, which alone is in relation with the light; yet even in this case, the symmetry of external forms has been displaced rather than destroyed, and at the first examination it seems complete. When the organs of relation are found placed on the interior, they frequently exhibit some irregularity, and to take an example of a known animal, the organ of voice, in the male duck, is a very remarkable one; in man even, the wind-pipe is not symmetrical, after it arrives at the first division of the bronchia. On the contrary, among the organs of the other life, those which are prominent on the exterior, constantly present the symmetrical character, as the thyroid gland; the mammary glands, &c.
[7] If we deny symmetry to the kidneys, because they are not uniformly composed of the same number of lobes in children, we must deny it also to the brain, the two lobes of which never exhibit the same arrangement in their circumvolutions; if we deny it to the salivary glands, because one is larger than the other, we must deny it to the extremities, because the right is usually more developed than the left. If these examples are not enough, a host of others might be cited; such as, the atrabiliary capsules, the bladder, the different organs of generation and lactation, and the very regular arrangement of the mucous follicles in certain parts situated upon the median line, &c. As to the anomalies that are observed in the distribution of the blood-vessels, they are also observed very frequently, though in a less evident manner, in the distribution of the nervous branches.