Natural History of the Mammalia of India and Ceylon
Chapter 18
Campbell, in his notes to 'The Old Forest-Ranger,' gives the dimensions of a tiger of 9 ft. 5 in. of which the tail was only 2 ft. 10 in. From the other detailed measurements it must have been an enormous tiger. The number of caudal vertebrae in the tiger and lion should be twenty-six. I now regret that I did not carefully examine the osteology of all short-tailed tigers which I have come across, to see whether they had the full complement of vertebrae. The big tiger in the museum is short by the six terminal joints = three inches. This may have occurred during life, as in the case of the above-quoted panther; anyhow the tail should, I think, be thrown out of the calculation. Now as to the measurement of the head and body, I quite acknowledge that there must be a different standard for the sportsman and for the scientific naturalist. For the latter the only reliable data are derived from the bones. Bones cannot err. Except in very few abnormal conditions the whole skeleton is in _accurate_ proportion, and it has lately struck me that from a certain measurement of the skull a true estimate might be formed of the length of the skeleton, and approximately the size of the animal over the muscles. I at first thought of taking the length of the skull by a craniometer, and seeing what portion of the total length to the posterior edge of the sacrum it would be, but I soon discarded the idea on account of the variation in the supra-occipital process.
I then took the palatal measurement, from the outer edge of the border in which the incisors are set to the anterior inside edge of the brain-hole, or foramen magnum, and I find that this standard is sufficiently accurate, and is 5.50 of the length taken from the tip of the premaxillaries to the end of the sacrum. Therefore the length of this portion of any tiger's skull multiplied by 5.50 will give the measurement of the head and body of the skeleton.
For the purpose of working out these figures I applied to all my sporting friends for measurements of their largest skulls, with a view to settling the question about tigers exceeding eleven feet. The museum possesses the skeleton of a tiger which was considered one of the largest known, the cranial measurement of whose skull is 14.50 inches, but the Maharajah of Cooch Behar showed me one of his skulls which exceeded it, being 15 inches. Amongst others I wrote to Mr. J. Shillingford of Purneah, and he most kindly not only drew up for me a tabular statement of the dimensions of the finest skulls out of his magnificent collection, but sent down two for my inspection. Now in the long-waged war of opinion regarding the size of tigers I have always kept a reserved attitude, for if I have never myself killed, or have seen killed by others, a tiger exceeding ten feet, I felt that to be no reason for doubting the existence of tigers of eleven feet in length vouched for by men of equal and in some cases greater experience, although at the same time I did not approve of a system of measurement which left so much to conjecture.
There is much to be said on both sides, and, as much yet remains to be investigated, it is to be hoped that the search after the truth will be carried on in a judicial spirit. I have hitherto been ranged on the side of the moderate party; still I was bound to respect the opinion of Sir Joseph Fayrer, who, as not only as a sportsman but as an anatomist, was entitled to attention; and from my long personal acquaintance I should implicitly accept any statement made by him. Dr. Jerdon, whom I knew intimately, was not, I may safely assert, a great tiger shikari, and he based his opinion on evidence and with great caution. Mr. J. Shillingford, from whom I have received the greatest assistance in my recent investigations, and who has furnished me with much valuable information, is on the other hand the strenuous assertor of the existence of the eleven-foot tiger, and with the magnificent skulls before me, which he has sent down from Purneah, I cannot any longer doubt the size of the Bengal tiger, and that the animals to which they belonged were eleven feet, _measured sportsman fashion_--that is round the curves. The larger of the two skulls measures 15.25 inches taken between two squares, placed one at each end; a tape taken from the edge of the premaxillaries over the curve of the head gives 17.37 inches; the width across the zygomatic arches, 10.50.[10] The palatal measurement, which is the test I proposed for ascertaining the length of the skeleton, is 12.25, which would give 5 feet 7.37 inches; about 3-3/4 inches larger than the big skeleton in the Museum. This may seem very small for the body of an animal which is supposed to measure eleven feet, but I must remind my readers that the bones of the biggest tiger look very small when denuded of the muscles; and the present difficulty I have to contend with is how to strike the average rate for the allowance to be added to skeleton for muscles, the chief stumbling block being the system which has hitherto included the tail in the measurement. It all tigers had been measured as most other animals (except felines) are--i.e. head and body together, and then the tail separately--I might have had some more reliable data to go upon; but I hope in time to get some from such sportsmen as are interested in the subject. I have shown that the tail is not trustworthy as a proportional part of the total length; but from such calculations as I have been able to make from the very meagre materials on which I have to base them, I should allow one 2.50th part of the total length of skeleton for curves and muscles.
[Footnote 10: At Mr. Shillingford's request, I made over this skull to the Calcutta Museum.]
In addition to a careful study of De Blainville's 'Osteographie,' where the bones are figured in large size to scale, I have made many careful measurements of skulls belonging to myself and friends, and also of the skulls and skeletons in the Calcutta Museum (for most willing and valuable assistance in which I am indebted to Mr. J. Cockburn, who, in order to test my calculations, went twice over the ground); and I have adopted the following formula as a tentative measure. I quite expect to be criticised, but if the crude idea can be improved on by others I shall be glad.
I now give a tabular statement of four out of many calculations made, but I must state that in fixing an arbitrary standard of 36 inches for tail, I have understated the mark, for the tails of most tigers exceed that by an inch or two, though, on the other hand, some are less.
_Formula_.--Measure from the tip of the premaxillaries or outer insertion of the front teeth (incisors) along the palate to the nearest inner edge of the foramen magnum. Multiply the result by 5.50. This will give the length of the skeleton, excluding the tail. Divide this result by 2.50, and add the quotient to the length for the proportionate amount of muscles and gain in curves. Add 36 inches for tail.
| Palatal | Add one | | | measure- | 2.50th | | | ment | part of | | | multi- | last for | | | plied | curves | | | by | and | | Total. | 5.50. | muscles. | Tail. | inches ft. in. -------------------+----------+----------+-------+-------------- Mr. Shillingford's | 67.37 | 26.94 | 36.00 | 130.31 10 10 tiger | | | | | | | | Big tiger in | 63.52 | 25.40 | 36.00 | 124.92 10 4-3/4 museum | | | | | | | | Maharajah of Cooch | 66.00 | 26.40 | 36.00 | 128.40 10 8.4 Behar's tiger | | | | | | | | A medium-sized one | 55.75 | 23.10 | 36.00 | 116.85 9 8-3/4 of my own | | | | ---------------------------------------------------------------- Remarks: Mr. Shillingford's tiger's tail was over 3 ft. 2 in., which would make it 11 ft. The Maharajah writes to me that his measured on the ground 9 ft. 11 in. See further on. ----------------------------------------------------------------
It will be seen that my calculation is considerably out in the Cooch Behar tiger, so I asked the Maharajah to tell me, from the appearance of the skull, whether the animal was young or old. He sent it over to me, and I have no hesitation in saying that it was that of a young tiger, who, in another year, might have put on the extra nine inches; the parietal sutures, which in the old tiger (as in Mr. Shillingford's specimens) are completely obliterated, are in this one almost open. It must be remembered that the bones of the skull do not grow in the same ratio to the others, and that they attain their full size before those of the rest of the body. Therefore it is only in the case of the adult that accurate results can be calculated upon. Probably I have not done wisely in selecting a portion of the skull as a standard--a bone of the body, such as a femur or humerus might be more reliable--but I was driven to it by circumstances. Sportsmen, as a rule, do not keep anything but the skull, and for general purposes it would have been of no use my giving as a test what no one could get hold of except in a museum.
I have always understood that the tiger of the plains grew to a greater size, that is in length, than the tiger of hilly country. I have never shot a tiger in Lower Bengal, therefore I cannot judge of the form of the beast, whether he be more lanky or not. If an eleven-foot Bengal tiger be anything like as robust in proportion as our Central Indian ones, I should say he was an enormous creature, but I believe the Central and Southern tiger to be the heavier one, and this is borne out by an illustration given by Mr. Shillingford in one of his able letters, which have called forth so much hostile criticism. He compares one of his largest with the measurement of a Southern India tiger:--
Locality of Tiger. | Purneah | Southern India Length. | 11 ft. 0 in. | 10 ft. 2 in. Girth of Chest. | 4 ft. 6 in. | 6 ft. 1 in. Girth of Head. | 2 ft. 10 in. | 3 ft. 5 in. Tail. | 3 ft. 4 in. | 3 ft. 1 in. Round Fore-arm. | 2 ft. 2 in. | 2 ft. 10 in. Height. | 3 ft. 7 in. | 3 ft. 9 in. Total of ft. and in. | 27 ft. 5 in. | 29 ft. 4 in. --------------------------------------------------------
The shorter tiger has an advantage of nearly two feet in all-round measurement.
Sir Joseph Fayrer has also been called in question for his belief in twelve feet tigers, but what he says is reasonable enough. "The tiger should be measured from the nose along the spine to the tip of the tail, as he lies dead on the spot where he fell, before the skin is removed. _One that is ten feet by this measurement is large, and the full-grown male does not often exceed this_, though no doubt larger individuals (males) are occasionally seen, and I have been informed by Indian sportsmen of reliability that they have seen and killed tigers over twelve feet in length." ('Royal Tiger of Bengal,' p. 29).
Sir Joseph Fayrer in a letter to _Nature_, June 27, 1878, brings forward the following evidence of large tigers shot by sportsmen whose names are well known in India.
Lieutenant-Colonel Boileau killed a tiger at Muteara in Oude, in 1861, over 12 feet; the skin when removed measured 13 feet 5 inches.
Sir George Yule has heard once of a 12-foot tiger fairly measured, but 11 feet odd inches is the largest he has killed, _and that twice or thrice_.
Colonel Ramsay (Commissioner) killed in Kumaon a tiger measuring 12 feet.
Sir Joseph Fayrer has seen and killed tigers over 10 feet, and one in Purneah 10 feet 8 inches, in 1869.
Colonel J. Sleeman does not remember having killed a tiger _over_ 10 feet 6 inches in the skin.
Colonel J. MacDonald has killed one 10 feet 4 inches.
The Honourable R. Drummond, C.S., killed a tiger 11 feet 9 inches, measured before being skinned.
Colonel Shakespeare killed one 11 feet 8 inches.
However, conceding that all this proves that tigers do reach occasionally to eleven and even twelve feet, it does not take away from the fact that the average length is between nine and ten feet, and anything up to eleven feet is rare, and up to twelve feet still more so.[11]
[Footnote 11: Since writing the above I have to thank "Meade Shell" for the measurements of the skull of a tiger 11 ft. 6 in. The palatal measurement is 12 inches, which, according to my formula, would give only 10 ft. 8 in.; but it must be remembered that I have allowed only 3 ft. for the tail, whereas such a tiger would probably have been from 3-1/2 to 4 ft., which would quite bring it up to the length vouched for. The tail of a skeleton of a much smaller tiger in the museum measures 3 ft. 3-1/2 in., which with skin and hair would certainly have been 3-1/2 ft. Until sportsmen begin to measure bodies and tails separately it will, I fear, be a difficult matter to fix on any correct formula.--R. A. S. See Appendix _C_.]
VARIETIES OF THE TIGER.--It is universally acknowledged that there is but one species of tiger. There are, however, several marked varieties. The distinction between the Central Asian and the Indian tiger is unmistakable. The coat of the Indian animal is of smooth, short hair; that of the Northern one of a deep furry pelage, of a much richer appearance.
There is an idea which is also to be found stated as a fact in some works on natural history, that the Northern tiger is of a pale colour with few stripes, which arises from Swinhoe having so described some specimens from Northern China; but I have not found this to be confirmed in those skins from Central Asia which I have seen. Shortly before leaving London, in 1878, Mr. Charles Reuss, furrier, in Bond Street, showed me a beautiful skin with deep soft hair, abundantly striped on a rich burnt sienna ground, admirably relieved by the pure white of the lower parts. That light-coloured specimens are found is true, but I doubt whether they are more common than the others. Of the varieties in India it is more difficult to speak. Most sportsmen recognise two (some three)--the stout thick-set tiger of hilly country, and the long-bodied lankier one of the grass jungles in the plains. Such a division is in consonance with the ordinary laws of nature, which we also see carried out in the thick-set muscular forms of the human species in mountain tracts.
Some writers, however, go further, and attempt subdivisions more or less doubtful. I knew the late Captain J. Forsyth most intimately for years. We were in the same house for some time. I took an interest in his writings, and helped to illustrate his last work, and I can bear testimony to the general accuracy of his observations and the value of his book on the Highlands of Central India; but in some things he formed erroneous ideas, and his three divisions, based on the habits of the tiger, is, I think, open to objection, as tending to create an idea of at least two distinct varieties.
Native shikaris, he says, recognise two kinds--the _Lodhia Bagh_ and the _Oontia Bagh_ (which last I may remind my readers is one of the names of the lion). The former is the _game-killing_ tiger, retired in his habits, living chiefly among the hills, retreating readily from man. "He is a light-made beast, very active and enduring, and from this, as well as his shyness, generally difficult to bring to bag."
I grant his shyness and comparative harmlessness (I once met one almost face to face)--and the nature of the ground he inhabits increases the difficulty in securing him--but I do not think he physically differs from his brother in the cattle districts. Mr. Sanderson says one of the largest tigers he had killed was a pure game-killer.
"The cattle-lifter again," says Forsyth, "is usually an older and heavier animal (called _Oontia Bagh_, from his faintly striped coat, resembling the colour of a camel), very fleshy and indisposed to severe exertion."
His third division is the man-eater. However, this is merely a classification on the habits of the same animal. I think most Central India sportsmen will agree with me when I say that many a young tiger is a cattle-eater, with a rich coloured hide, although it often happens that an old tiger of the first division, when he finds his powers for game failing by reason of age or increased bulk, transfers himself from the borders of the forest to the vicinity of grazing lands and villages, and he ultimately may come into the third division by becoming a man-eater. So that the _Lodhia_ becomes the _Oontia_ (for very old tigers become lighter in colour), and may end by being an _Adam-khor_, or man-eater. Tigers roam a great deal at times, and if in their wanderings they come to a suitable locality with convenience of food and water, they abide there, provided there be no occupant with a prior claim and sufficient power to dispute the intrusion. We had ample proof of this at Seonee. Close to the station, that is, within a short ride, were several groups of hills which commanded the pasture lands of the town. Many a tiger has been killed there, the place of the slain one being occupied ere long by another. On the other hand, if a tiger be accommodated with lodgings to his liking, he will stay there for years, roaming a certain radius, but returning to his home; and it is the knowledge of this that so often enables the hunter to compass his destruction. As long therefore as there are human habitations, with their usual adjuncts of herds and flocks, within a dozen miles of the jungle tiger's haunts, so long there will always be the transition from the game-killer to the cattle-lifter and the man-eater. Colour and striping must also be thrown out of the question, for no two individuals of any variety agree, and the characteristics of shade and marking are common to all kinds. The only reliable data therefore are derived from measurements, and from these it may be proved that the grass-jungle tiger of Bengal, though the longer animal, is yet inferior in all round measurement and probably in weight to the tiger of hilly country--see Mr. Shillingford's comparison quoted by me above. Let also any one compare the following measurements of one given by Colonel Walter Campbell with a tiger of equal length shot in the grassy plains of Bengal:--
ft. in. Length from point of nose to end of tail . . . . . 9 5 Ditto of tail . . . . . . . . . . . . . . . . . . 2 10 Height from heel to shoulder . . . . . . . . . . . 3 2 Extreme length from shoulder to point of toe . . . 3 11 From elbow to point of toe . . . . . . . . . . . . 2 0 Girth of body just behind the shoulder . . . . . . 5 3 Ditto of forearm . . . . . . . . . . . . . . . . . 2 7 Ditto of neck . . . . . . . . . . . . . . . . . . 3 0 Circumference of head . . . . . . . . . . . . . . 3 3
This is a remarkably short-tailed tiger. If the concurrence of evidence establishes the difference beyond doubt, then we may say that there are two varieties in India--the hill tiger, _Felis tigris, var. montanus_; and the other, inhabiting the alluvial plains of great rivers, _Felis tigris, var. fluviatilis_. Dr. Anderson says he has examined skulls and skins of those inhabiting the hill ranges of Yunnan, and can detect no difference from the ordinary Indian species.
The tigress goes with young for about fifteen weeks, and produces from two to five at a birth. I remember once seeing four perfectly formed cubs, which would have been born in a day or two, cut from a tigress shot by my brother-in-law Col. W. B. Thomson in the hills adjoining the station of Seonee. I had got off an elephant, and, running up the glen on hearing the shots, came unpleasantly close to her in her dying throes. When about to bring forth, the tigress avoids the male, and hides her young from him. The native shikaris say that the tiger kills the young ones if he finds them. The mother is a most affectionate parent as a rule, and sometimes exhibits strange fits of jealousy at interference with her young. I heard an instance of this some years ago from my brother, Mr. H. B. Sterndale, who, as one of the Municipal Commissioners of Delhi, took a great interest in the collection of animals in the Queen's Gardens there. Both tiger and leopard cubs had been born in the gardens, and the mother of the latter shewed no uneasiness at her offspring being handled by strangers as they crept through the bars and strayed about; but one day, a tiger cub having done the same, the tigress exhibited great restlessness, and, on the little one's return, in a sudden accession of jealous fury she dashed her paw on it and killed it. I am indebted to Mr. Shillingford for a long list of tigresses with cubs killed during the years 1866 to 1880. Out of 53 cubs (18 mothers) 29 were males and 22 females, the sex of two cubs not being given. This tends to prove that there are an equal number of each sex born--in fact here the advantage is on the side of the males. I have heard it asserted that tigresses are more common, and native shikaris account for it by saying that the male tiger kills the cubs of his own sex; but I have not seen anything to justify this assertion, or the fact of there being a preponderance of females. Mr. Sanderson, however, writes: "Male and female cubs appear to be in about equal proportions. How it is that amongst mature animals tigresses predominate so markedly I am unable to say."
Tigresses have young at all seasons of the year, and they breed apparently only once in three years, which is about the time the cubs remain with their mother.
For the following interesting memorandum I have to thank Mr. Shillingford:--
Feet. "Cubs one year old measure . . . . . . Males 4-1/2 to 5-1/2 Females 4 to 5 Ditto two years old . . . . . . . . . Males 5-1/2 to 7 Females 5 to 6-1/2 Ditto three years old . . . . . . . . Males 7 to 8-1/2 Females 6-1/2 to 7-1/2
"When they reach three years of age they lose their 'milk' canines, which are replaced by the permanent fangs, and at this period the mother leaves them to cater for themselves."
The cubs are interesting pets if taken from the mother very young. I have reared several, but only kept one for any length of time. I have given a full description of Zalim and his ways in 'Seonee.' He was found by my camp followers with another in a nullah, and brought to me. The other cub died, but Zalim lived to grow up into a very fine tiger, and was sent to England. I never allowed him to taste raw flesh. He had a little cooked meat every day, and as much milk as he liked to drink, and he throve well on this diet. When he was too large to be allowed to roam about unconfined I had a stout buffalo-leather collar made for his neck, and he was chained to a stump near the cook-room door. With grown-up people he was perfectly tame, but I noticed he got restless when children approached him, and so made up my mind to part with him before he did any mischief.