Manual of the Trees of North America (Exclusive of Mexico) 2nd ed.
Part 96
Leaves thin, ovate, abruptly pointed at apex, obliquely truncate or unsymmetrically cordate at base, and coarsely serrate with long slender straight or slightly curved conspicuously glandular teeth, as they unfold, dark red and sparingly pubescent on the midrib and veins, glabrous at the end of a few days, without or rarely with small axillary tufts, dark green on the upper surface, pale yellow-green or glaucous (var. _glaucescens_ Sarg.) on the lower surface, 4′—4½′ long and 2½′—3½′ wide; petioles slender, glabrous, 2′—2½′ in length. Flowers opening early in June, about ⅓′ long, on hoary-tomentose pedicels, in broad usually 10 or 12, sometimes 30 or 40-flowered long-branched glabrous cymes; peduncle glabrous, the free portion ⅘′—1¼′ in length, its bract oblong, often slightly falcate, cuneate or rounded at base, rounded at apex, glabrous, 3′—4′ long, ½′—1¼′ wide, decurrent nearly to the base of the peduncle; sepals acute, rusty-tomentose on the outer surface, glabrous on the inner surface; petals oblong-ovate, narrowed at the rounded apex; staminodia oblong-obovate rounded at the broad apex; style glabrous. Fruit ripening in September, subglobose to depressed-globose, covered with rusty tomentum, ¼′—⅓′ in diameter.
Usually a small tree with pale furrowed or sometimes checkered bark, small spreading branches forming a narrow round-topped head, and slender glabrous orange or red-brown branchlets. Winter-buds ovoid, obtusely pointed, dull red, glabrous, ⅙′—⅕′ long.
Distribution. Central and southwestern Mississippi (Hinds and Adams Counties); Dallas County, Alabama; West Feliciana and Calcasieu Parishes, Louisiana, to the valley of the Brazos River, eastern Texas, and to Hempstead County (Fulton and McNab), southern Arkansas; the var. _glaucescens_ with the type, and near Page, Le Flore County, Oklahoma; in wet woods subject to overflow at San Augustine, San Augustine County, Texas, a variety (var. _brevipedunculata_ Sarg.), differs from the type in the less coarsely serrate smaller leaves glaucescent below, in the shorter free portion of the peduncle of the inflorescence and its broader bract. A tree 25°—30° high, with slender glabrous dark red-brown branchlets.
3. Tilia venulosa Sarg.
Leaves broad-ovate, abruptly acuminate at apex, cordate or unsymmetrically cordate or obliquely truncate or cordate at base, coarsely serrate with gland-tipped teeth pointing forward, covered when they unfold with pale tomentum, soon becoming pubescent, and glabrous before the flowers open, dark yellow-green on the upper surface, paler on the lower surface, 4′—4¾′ long and broad, with a prominent pale yellow midrib slightly villose on the upper side near the base, and 9 or 10 pairs of remote primary veins without axillary tufts and connected by conspicuous cross veinlets; petioles stout, glabrous, 1¾′—2′ in length. Flowers opening early in July, ⅓′ long, on slightly pubescent pedicels, in broad slender-branched nearly glabrous cymes; peduncle stout, glabrous, red, the free portion 1′—1½′ in length, its bract oblong to slightly obovate, gradually narrowed and rounded at base, rounded at apex, glabrous on the upper surface, pubescent below on the midrib and veins, 3½′—6′ long and 1¼′—1½′ wide, longer than the peduncle and decurrent nearly to its base or to within 1′—1½′ of its base; sepals ovate, acute, pale pubescent on the outer surface, villose and furnished at base on the inner surface with a tuft of long white hairs, a third shorter than the lanceolate acuminate petals; staminodia oblong-obovate, rounded at apex, about as long as the sepals; stigma slightly villose at base. Fruit ripening the end of September, subglobose, ¼′—⅓′ in diameter, covered with loose light brown pubescence.
A tree, 60°—75° high, with stout red glabrous branchlets. Winter-buds ovoid, cylindric, obtusely pointed, dark red, ¼′—⅓′ in length.
Distribution. North Carolina, rocky “coves” in rich soil, Hickory Nut Gap, in the Blue Ridge, and near Saluda, Polk County, passing into var. _multinervis_ Sarg., differing from the type in its obliquely truncate, not cordate, leaves with 12 or 13 pairs of more crowded primary veins, ellipsoid fruit, slender branchlets, and smaller winter-buds; a single tree near Saluda, Polk County.
4. Tilia littoralis Sarg.
Leaves ovate, abruptly short-pointed and acute or acuminate at apex, unsymmetric and rounded on one side and cuneate on the other, or symmetric and cuneate or oblique and truncate at base, and finely serrate with straight or incurved glandular teeth, covered above when they unfold with scattered fascicled hairs and tomentose below, soon glabrous, and when the flowers open, thin, yellow-green, paler, rarely glaucous (var. _discolor_ Sarg.) on the lower than on the upper surface, 3′—4′ long and 1¾′—2′ wide, with a slender midrib and primary veins and small conspicuous tufts of rusty brown axillary hairs; petioles slender, glabrous, 1′—1½′ in length; leaves on young vigorous shoots broad-ovate, truncate or slightly cordate at base, more coarsely serrate, pubescent with fascicled hairs especially on the midrib and veins, 4′—5′ long and 3′—4′ wide; petioles densely pubescent. Flowers opening the middle of June, ⅓′ long, on pale tomentose pedicels, in small, compact, mostly 9—15-flowered, pubescent cymes; peduncle covered with scattered fascicled hairs, the free portion ⅗′—1′ long, its bract gradually narrowed and cuneate at base, rounded at apex, ciliate on the margins, pubescent on the midrib, otherwise glabrous, 2′—7′ long, ¼′—⅖′ wide, longer or shorter than and decurrent to the base or nearly to the base of the peduncle; sepals acuminate, pale pubescent on the outer surface, villose on the inner surface along the margins and at the base with long white hairs; petals acuminate; staminodia oblong-obovate, rounded at apex. Fruit ellipsoid to depressed-globose, apiculate, covered with pale brown tomentum, ¼′—⅓′ in diameter.
A tree with slender glabrous branchlets densely coated when they first appear with pale pubescence, soon glabrous, light reddish brown during their first summer, often bright red during their first winter, becoming purple the following year and ultimately light gray-brown. Winter-buds ovoid, glabrous or puberulous, bright red, about ⅕′ long and 1/12′—⅛′ in diameter.
Distribution. Georgia, shore of Colonel’s Island near the mouths of the North Newport and Medway Rivers, near Durham, Liberty County; the var. _discolor_ with the type
5. Tilia crenoserrata Sarg.
_Tilia floridana_ Sarg., not Small.
Leaves ovate, abruptly narrowed and acuminate at apex, usually oblique and unsymmetrically cordate or truncate or occasionally symmetrical and cordate at base, crenately serrate, the teeth tipped with minute glands, covered when they unfold with pale caducous tomentum, and at maturity dark green and lustrous above, glaucescent below, glabrous with the exception of minute axillary tufts of rusty hairs, mostly 3½′—5½′ long and 2¾′—3′ wide; petioles slender, glabrous, about 1¼′ in length. Flowers opening the middle of June, ⅓′ long, on hoary-tomentose pedicels, in compact mostly 10—18-flowered tomentose cymes; peduncle glabrous, the free portion 1′—1½′ in length, its bract oblong-obovate, cuneate at base, rounded at apex, glabrous, 3′—5′ long, usually about ⅘′ wide, decurrent nearly to the base of the peduncle; sepals acute, hoary-tomentose on the outer surface, coated with pale tomentum mixed with long white hairs on the inner surface; petals narrow-acuminate; staminodia oblong-obovate, notched at apex. Fruit ripening from the middle to the end of August, ellipsoid, conspicuously apiculate at apex, rusty-tomentose, ⅓′—⅖′ long and ¼′—⅓′ in diameter.
A tree, 25°—30°, rarely 60° high, with a trunk 10′—12′ rarely 18′—20′ in diameter, and slender glabrous red-brown branchlets. Winter-buds ovoid, acute, dark dull red, glabrous, ⅙′—⅕′ long.
Distribution. Near Albany, Dougherty County, Georgia, to central Florida (Levy, Columbia, Alachua, Putnam, Seminole and Orange Counties).
6. Tilia floridana Ashe.
Leaves broad-ovate, acuminate or abruptly acuminate at apex, cordate or obliquely truncate at base and coarsely serrate with apiculate teeth, tinged with red and tomentose below when they unfold, fully grown and glabrous or nearly glabrous when the flowers open late in May or in early June, and at maturity thin, glabrous, dark yellow-green on the upper surface, pale or rarely covered below with a silvery white bloom (var. _hypoleuca_ Sarg.), 3½′—5′ long and 2½′—3½′ wide, with a slender midrib and primary veins; in the east usually without axillary tufts, often present and sometimes conspicuous westward; petioles slender, glabrous, ¾′—1′ in length. Flowers opening in early summer ⅕′—¼′ long, on hoary-tomentose rarely puberulous (var. _australis_ Sarg.) pedicels, in few-flowered rather compact pubescent corymbs; peduncle pubescent, the free portion 1½′—2½′ in length, its bract oblong-obovate to oblong, rounded at apex, often falcate, glabrous, 3′—6′ long, ½′—¾′ wide, decurrent nearly to the base of the peduncle; sepals narrow, ovate, acuminate, hoary-tomentose on the outer surface, sparingly villose on the inner surface, two-thirds as long as the lanceolate petals; staminodia oblong-obovate, acute, nearly as long as the petals; style glabrous. Fruit ripening in August and September, subglobose to ellipsoid, rusty-tomentose, ½′ in diameter.
A tree, 40°—50° high, with a trunk 12′—15′ in diameter, and slender glabrous red-brown or yellow branchlets. Winter-buds obtuse, dark red-brown, glabrous, about ⅙′ long.
Distribution. North Carolina (Polk County) to western Florida and westward through northern and central Alabama, central Mississippi, northern and western Louisiana, eastern and over the Edwards Plateau to Kerr, Bandera and Uvalde Counties, Texas, and through southern and western Arkansas to eastern Oklahoma, Missouri and eastern Kentucky; in northeastern Mexico; the var. australis in Blount County, Alabama. A variety (var. _oblongifolia_ Sarg.) with narrower more elongated leaves with more prominent tufts of axillary hairs occurs in Putnam, Leon and Gadsden Counties, Florida, on the bluffs of the Alabama River near Berlin, Dallas County, Alabama, in Hinds, Rankin and Adams Counties, Mississippi, in West Feliciana, Iberia (Avery Island) and Natchitoches Parishes, Louisiana, in Hempstead and Salina Counties, Arkansas, and in Harris, Anderson and Livingston Counties, Texas.
7. Tilia Cocksii Sarg.
Leaves ovate, abruptly acuminate at apex, very oblique at the truncate or rounded base, dentate with small remote glandular apiculate teeth, covered when they unfold with loose floccose pubescence, nearly glabrous when fully grown early in April, when the flowers open the middle of May dark green and lustrous on the upper surface, pale blue-green and lustrous below, and at mid-summer when the fruit ripens, subcoriaceous, dark green and lustrous on the upper surface, paler on the lower surface, with slender primary veins without or occasionally with minute axillary tufts, and connected by conspicuous straight or curved veinlets, 3½′—4′ long and 2½′—3′ wide; petioles slender, glabrous, ¾′—1′ in length; leaves on leading summer branchlets sometimes obliquely cordate, more coarsely serrate, covered on the upper surface with short fascicled hairs, and floccose-pubescent on the lower surface, 4′—5′ long and 4′—4¾′ wide, their petioles puberulous. Flowers opening the middle of May, ¼′ long, on tomentose pedicels, in compact pubescent many-flowered cymes; peduncle slender, glabrous, the free portion only ⅗′—⅘′ in length, its bract oblong, occasionally slightly obovate, rounded at the ends, hoary-tomentose on the under surface and pubescent on the upper surface when it first appears, and when the flowers open puberulous below and glabrous above, 3½′—6′ long, ½′—⅗′ wide and shorter than and decurrent to the base of the peduncle; sepals ovate, acuminate, pale pubescent on the outer surface, villose at the base on the inner surface, a third shorter than the lanceolate acuminate petals; staminodia oblong-obovate, rounded at apex, about half the length of the petals; style glabrous. Fruit ripening the middle of July, globose to depressed-globose, covered with loose brown tomentum, ¼′ in diameter.
A small tree with slender dull red glabrous branchlets, the leading branchlets in summer more or less pubescent. Winter-buds ovoid, acute, dull red, glabrous or pubescent on leading shoots, ⅕′—¼′ long.
Distribution. Louisiana, river banks and low woods, Lake Charles and West Lake Charles, Calcasieu Parish.
8. Tilia neglecta Spach.
_Tilia Michauxii_ Sarg., not Nutt.
Leaves thick and firm, acute or abruptly narrowed and long-pointed at apex, obliquely concave or unsymmetrically cordate at base, coarsely serrate with straight apiculate teeth pointing forward, dark green, smooth, glabrous and lustrous above, covered below except on the midrib and veins more or less thickly with short gray pubescence often slightly tinged with brown, and furnished with conspicuous tufts of axillary hairs, usually 4′—5½′ long and 2½′—4½′ wide; petioles stout, glabrous, 1¼′—2½′ in length. Flowers opening in June and July about ⅖′ long, on pubescent or nearly glabrous pedicels, in long-branched slender glabrous mostly 5—15-flowered cymes; peduncle slender, glabrous, the free portion 1¼′—1½′ in length, its bract gradually narrowed and cuneate or unsymmetrically cuneate or rounded at base, rounded at apex, glabrous, 2¾′—4½′ long, ⅖′—⅘′ wide and longer than and decurrent nearly to the base or to within ⅗′ of the base of the peduncle; sepals broad-ovate, acute, ciliate on the margins, glabrous on the outer surface, covered on the inner surface with long white hairs, about half as long as the lanceolate petals rounded and notched at apex and rather longer than the spatulate staminodia; stamens included; style villose toward the base. Fruit ripening in September, ellipsoid, ovoid, obovoid, or depressed-globose, rounded or acute or rarely gradually narrowed and acuminate at apex, rarely 5-angled, covered with rusty or pale pubescence, usually about ⅓′ in diameter.
A tree, 75°—90° high, with a trunk sometimes 3° in diameter, smooth often pendulous branches forming a broad round head, and slender glabrous branchlets. Winter-buds ovoid, rounded at the narrowed apex, about ⅕′ long, with glabrous red-brown or light brown scales. Bark of the trunk about 1′ thick, deeply furrowed, pale reddish brown and covered with small thin scales.
Distribution. Rich moist soil, Province of Quebec, near Montreal, to the coast of Massachusetts and New York, through the middle states to the valley of the Potomac River and along the Appalachian Mountains to those of North Carolina, and to Iuka, Tishomingo County, Mississippi, and from central and western New York to northern Missouri.
9. Tilia caroliniana Mill.
Leaves ovate, oblique and truncate or cordate at base, abruptly long-pointed at apex, coarsely dentate with broad apiculate glandular teeth pointing forward, and coated below with a rusty or pale easily detached pubescence of fascicled hairs, coated when they unfold with hoary tomentum, soon glabrous on the upper surface, and at maturity dark yellow-green and lustrous above, 2¾′—4½′ long and 2½′—5′ wide; petioles stout, glabrous, 1′—1½′ in length. Flowers opening the middle of June, ¼′ long, on slender pubescent pedicels, in small stout-branched pubescent mostly 8—15-flowered cymes; peduncle slender, pubescent, the free portion ¾′—1¼′ long, its bract oblong-obovate, cuneate at base, rounded or acute at apex, nearly glabrous on the upper surface when it first appears, pubescent becoming glabrous or almost glabrous below, 4′—5′ long and ⅘′ wide, longer or shorter than and decurrent to the base or nearly to the base of the peduncle; sepals ovate, acuminate, ciliate on the margins, brown and covered with pale pubescence on the outer surface, coated on the inner surface with long white hairs; petals lanceolate, acuminate, a third longer than the sepals; staminodia oblong-obovate, rounded at apex, rather shorter than the petals; style tomentose at base or glabrous. Fruit subglobose, ellipsoid or obovoid, ⅛′ in diameter.
A large tree with slender red-brown glabrous or slightly pubescent branchlets. Winter-buds ovoid, acute, glabrous or rarely pubescent, about ¼′ long.
Distribution. Coast of North Carolina (Wrightsville Beach and the neighborhood of Wilmington, New Hanover County), southward in the immediate neighborhood of the coast to Liberty County, Georgia; western Louisiana to southern Arkansas (Hempstead and Clark Counties) common, and through eastern Texas to the Edwards Plateau (near Boerne, Kendall County); in Orizaba. Passing into
Tilia caroliniana var. rhoophila Sarg.
Differing from the type in its pubescent branchlets and winter-buds, its usually larger leaves, and in its tomentose corymbs of more numerous flowers. Leaves broad-ovate, abruptly short-pointed and acuminate at apex, oblique and truncate or cordate at base, coarsely serrate with broad apiculate teeth pointing forward, dark green and lustrous on the upper surface, pale and thickly covered on the lower surface with persistent white or brownish pubescence, 4′—5′ long and 2½′—5′ wide, with a slender midrib and primary veins pubescent on the lower side, and small conspicuous axillary tufts of pale hairs; petioles stout, thickly coated with pubescence, 1′—1¾′ in length; leaves on vigorous shoots often 6′ long, and 5½′ wide, and occasionally 10′ long and 9′ wide. Flowers ¼′ long, on short hoary-tomentose pedicels, in wide thin-branched pubescent many-flowered (sometimes 50) cymes; peduncle thickly covered with fascicled hairs, the free portion ¼′ long, its bract oblong, unequally rounded at base, rounded at apex, glabrous on the upper surface, pubescent on the lower surface, 4′—6′ long, 1′—2′ wide, usually shorter than and decurrent nearly to the base of the peduncle; sepals acuminate, coated on the outer surface with pale or slightly rusty pubescence, villose and furnished at base on the inner surface with tufts of long hairs; petals lanceolate, acuminate and ciliate at apex, about a third longer than the sepals; staminodia spatulate, acute, about half the length of the petals; style coated at base with long white hairs. Fruit subglobose, covered with rusty tomentum, about ⅓′ in diameter.
A tree with slender branchlets thickly coated during their first year with pale pubescence, dark red-brown or gray and puberulous during their second season. Winter-buds covered with pale pubescence.
Distribution. Western Louisiana (Calcasieu and Jefferson Davis Parishes) to Hempstead County, Arkansas, and through eastern Texas to the valley of the upper Guadalupe River, Kerr County.
10. Tilia texana Sarg.
Leaves thin, oblong-ovate, abruptly contracted into a long slender acuminate point, cordate or obliquely cordate at base, finely dentate with broad apiculate teeth, early in the season pubescent above with scattered fascicled hairs and covered below with brownish slightly attached pubescence, and in the autumn light yellow-green, lustrous and nearly glabrous on the upper surface, slightly pubescent on the lower surface, 4′—5½′ long and 3¼′—5′ wide, with a slender midrib and primary veins sparingly villose on the upper side and nearly glabrous on the lower side, and small axillary tufts of brownish hairs; petioles slender, pubescent with fascicled hairs, 1′—1½′ in length; leaves on vigorous shoots often furnished with one or two large lateral acuminate serrate lobes, more coarsely dentate and more thickly covered on the lower surface with pubescence, often 5½′—6′ long and 3½′—6′ wide. Flowers opening the middle of June, ¼′ long, on slender tomentose pedicels, in small villose-pubescent mostly 7—10-flowered cymes; peduncle slender, slightly villose-pubescent, the free portion 1¼′—1½′ in length, its bract oblong-ovate to slightly obovate, unsymmetrically cuneate at base, rounded and occasionally lobed at apex, glabrous on the upper surface, densely pubescent early in the season, later becoming nearly glabrous on the lower surface, 3′—6′ long and ¾′—1¼′ wide, longer or shorter than the peduncle and decurrent to its base or to within 1½′ of its base; sepals ovate, acute, pale pubescent on the outer surface, covered on the inner surface with white hairs longer and more abundant near the base; petals lanceolate, acuminate, a third longer than the sepals; staminodia linear-lanceolate, acuminate; style hoary-tomentose at base. Fruit ellipsoid, covered with rusty brown tomentum, ⅓′ long and ¼′ broad.
A small tree with slender branchlets thickly covered during their first season with close pale pubescence, and pale and puberulous or glabrous in their second year; on vigorous terminal branchlets often with thicker, light rusty brown pubescence. Winter-buds ovoid, obtusely pointed, thickly covered with pale pubescence, ¼′ long.
Distribution. Texas, Brazos and Cherokee Counties, on Spring Creek near Boerne, Kendall County, and on the rocky banks of the Guadalupe River at Kerrville, Kerr County.
11. Tilia phanera Sarg.
Leaves semiorbicular to broad-ovate, deeply and usually symmetrically cordate at base, abruptly short-pointed at apex, finely dentate with straight or incurved apiculate teeth, glabrous above when they unfold with the exception of a few hairs on the midrib and veins, and thickly coated below with hoary tomentum, and at maturity thin, blue-green, smooth and lustrous on the upper surface, paler and often brownish and coated with a floccose easily detached pubescence of fascicled hairs or scabrate (var. _scabrida_ Sarg.) on the lower surface, 2′—4′ wide and usually rather broader than long, with a slender midrib and primary veins pubescent on the lower side, and small axillary clusters of rusty brown hairs; petioles slender, coated when they first appear with hoary tomentum, glabrous or slightly pubescent in the autumn, 1′—1½′ in length. Flowers opening the middle of June, ⅕′ long, on tomentose pedicels, in compact villose mostly 16—20-flowered cymes; peduncle villose, the free portion 1¼′ in length, its bract obovate, cuneate at base, broad and rounded at apex, floccose-pubescent on the lower surface, nearly glabrous on the upper surface, 3′—3½′ long and ½′—1′ wide, longer than the peduncle and decurrent to its base or to within ¼′ of its base; sepals acuminate, pale pubescent on the outer surface, villose on the margins and furnished at base on the inner surface with a tuft of long white hairs, broader and shorter than the lanceolate acuminate petals; staminodia oblong-obovate, rounded at apex, style glabrous except at the base. Fruit ripening the end of September, ellipsoid, covered with rusty tomentum, ⅓′—⅖′ long and ¼′ wide, on a stout, densely floccose-pubescent pedicel.
A tree with slender light gray-brown often zigzag branchlets covered when they first appear with fascicled hairs and deciduous during their first summer. Winter-buds ovoid, obtusely pointed, terete, reddish brown, glabrous, ⅙′—⅕′ long.
Distribution. Texas, banks of Spring Creek, near Boerne, Kendall County; the var. _scabrida_ on a low limestone bluff of the Blanco River, near Blanco, Blanco County, near College Station, Brazos County, and at Velasco, Brazoria County.
12. Tilia lasioclada Sarg.