Part 23

Distribution. Banks of the Potomac River, District of Columbia, and Alleghany County, Maryland to Natural, Rockbridge, Fairfax and Elizabeth Counties, Virginia; northern Kentucky; northern Tennessee; northeastern Mississippi (near Iuka, Tishamingo County); St. Clair and Madison Counties, Illinois; more abundant in Missouri from Pike County southward to southwestern Kansas, western Arkansas and eastern Oklahoma.

8. Salix lasiandra Benth. Yellow Willow.

Leaves lanceolate to ovate-lanceolate, acuminate and long-pointed at apex, cuneate or rounded at base, often slightly falcate, finely serrate, glabrous, dark green and lustrous above, pale or glaucous below, 1½′—3′ long, about ½′ wide, on vigorous summer shoots often 6′ or 7′ long and 1½′ wide; petioles slender, glabrous, glandular at apex, ¼′ in length, or on summer shoots stout and 1′—1½′ long; stipules reniform, caducous. Flowers: aments terminal on leafy puberulous branchlets, narrow-cylindric, 2½′—3′ in length; scales pale pubescent, those of the staminate ament lanceolate-acuminate to obovate and rounded at apex and entire, those of the pistillate ament obovate and usually dentate near the apex; stamens 5—9; filaments hairy below the middle; ovary rather abruptly narrowed above the middle and acuminate, long-stalked; style short with slightly emarginate lobes. Fruit light red-brown, ¼′ long; pedicels about 1/16′ in length.

Distribution. Valley of the Yukon River near Dawson, Yukon, Vancouver Island, and southward near the coast of Washington and Oregon, and on the western slope of the Sierra Nevada and on the coast ranges to southern California, ranging from the sea-level to altitudes of 8500° on the southern Sierra Nevada; in New Mexico (Glenwood, Soccoro County, and Santa Fé, Santa Fé County); in Colorado (Buena Vista, Chaffee County, _Alice Eastwood_). Passing into var. _caudata_ Sudw., distinguished by its caudate-acuminate leaves green on both surfaces, and by its bright yellow or orange-yellow branchlets, and ranging from northeastern Oregon and eastern Washington through Idaho, and from northern Wyoming to southern Colorado, Utah and Nevada.

A variety (var. _lancifolia_ Bebb), differing from the typical _S. lasiandra_ in the gray or rusty villose pubescence covering the branchlets during their first and sometimes their second season and the lower surface of the young leaves, is distributed from Dawson in the valley of the Yukon River southward to the valley of the upper Nesqually River, Washington, to the valley of the Willamette River (Salem, Oregon), to Santa Cruz, Santa Cruz County, and to the San Bernardino Mountains, California.

9. Salix lucida Muehl. Shining Willow.

Leaves ovate-lanceolate, or narrow lanceolate (f. _angustifolia_ Anders.), acuminate and long-pointed at apex, cuneate or rounded at base, finely serrate, 3′—5′ long, 1′—1½′ wide, covered when they unfold with scattered pale caducous hairs, at maturity coriaceous, smooth and lustrous, dark green above, paler below, with a broad yellow midrib, and slender primary veins arcuate and united near the margins; petioles stout, yellow, puberulous, glandular at the apex, with several dark or yellow conspicuous glands, ¼′—½′ long; stipules nearly semicircular, glandular-serrate, membranaceous, ⅛′—¼′ wide, often persistent during the summer. Flowers: aments erect, tomentose, on stout puberulous peduncles terminal on short leafy branchlets, the staminate oblong-cylindric, densely flowered, about 1½′ in length, the pistillate slender, elongated, 1½′—2′ long, often persistent until late in the season; scales oblong or obovate, rounded, entire, erose or dentate at apex, light yellow, nearly glabrous or coated on the outer surface with pale hairs, often ciliate on the margins; stamens usually 5, with elongated free filaments slightly hairy at base; ovary narrowly cylindric, long-stalked, elongated, glabrous, with nearly sessile emarginate stigmas. Fruit: cylindric, lustrous, about ⅓′ long.

A tree, occasionally 25° high, with a short trunk 6′—8′ in diameter, erect branches forming a broad round-topped symmetrical head, and stout glabrous branchlets dark orange color and lustrous in their first season, becoming darker and more or less tinged with red the following year; usually smaller and shrubby in habit. Winter-buds narrowly ovoid, acute, light orange-brown, lustrous, about ¼′ long. Bark thin, smooth, dark brown slightly tinged with red.

Distribution. Banks of streams and swamps; Newfoundland to the shores of Hudson’s Bay and northwestward to the valley of the Mackenzie River and the eastern base of the Rocky Mountains, southward to southern Pennsylvania, northeastern Iowa, the Turtle Mountains, North Dakota, and eastern Nebraska; very abundant at the north, rare southward; a variety from extreme northeastern New England and adjacent New Brunswick and Quebec (var. _intonsa_ Fernald) is distinguished by its often linear leaves rufous pubescent during the season on the under side of the veins and by its pubescent branchlets; a shrub or tree up to 25°.

10. Salix taxifolia H. B. K.

Leaves linear-lanceolate, narrowed at the ends, acute, slightly falcate, mucronate at the apex, entire or rarely obscurely dentate above the middle, coated as they unfold with long soft white hairs, at maturity pale gray-green, slightly puberulous, ⅓′—1⅓′ long, 1/12′—⅛′ wide, with a slender midrib, thin arcuate veins, and thickened slightly revolute margins; petioles stout, puberulous, rarely 1/12′ long; stipules ovate, acute, scarious, minute, caducous. Flowers: aments densely flowered, oblong-cylindric or subglobose, ¼′—½′ long, terminal, or terminal and axillary on the staminate plant, on short leafy branchlets; scales oblong or obovate, rounded or acute and sometimes apiculate at apex, coated on the outer surface with hoary tomentum and pubescent or glabrous on the inner; stamens 2, with free filaments hairy below the middle; ovary ovoid-conic, short-stalked or subsessile, villose, with nearly sessile deeply emarginate stigmas. Fruit cylindric, long-pointed, bright red-brown, more or less villose, short-stalked, about ¼′ long.

A tree, often 40°—50° high, with a trunk 18′ in diameter, erect and drooping branches forming a broad open head, and slender branchlets covered during their first season with hoary tomentum, becoming light reddish or purplish brown and much roughened by the elevated persistent leaf-scars. Winter-buds ovoid, acute, dark chestnut-brown, puberulous, about 1/16′ long and nearly as broad as long. Bark of the trunk ¾′—1′ thick, light gray-brown, and divided by deep fissures into broad flat ridges covered by minute closely appressed scales.

Distribution. Near El Paso, Texas; southwestern New Mexico, and along mountain streams in southern Arizona; southward through Mexico to Guatemala, and on the Sierra de la Victoria, Lower California.

11. Salix sessilifolia Nutt.

Leaves linear-lanceolate to elliptic-lanceolate, acute or acuminate at apex, cuneate at base, entire or furnished above the middle with a few remote apiculate glandular teeth, bluish green and thickly covered with silky white hairs most abundant on the lower side of the midrib, 1′—2′ long, ½′—¾′ wide, or on vigorous summer shoots often 4′ long and 1¼′ wide; petioles densely villose-pubescent, 1/16′—⅛′ in length; stipules ovate to lanceolate, acute, entire or denticulate. Flowers: aments appearing after the leaves, terminal on leafy branchlets, densely hoary-tomentose, 1½′—2½′ long; scales broadly elliptic, acute or rounded at apex, cuneate at base, densely villose-tomentose; stamens 2; filaments villose below the middle; ovary sessile, villose, the stigmas sessile, deeply 2-lobed. Fruit ovoid-acuminate, densely villose, pubescent.

A shrub or small tree occasionally 20° high, with short hairy tomentose branchlets.

Distribution. River banks, southwestern British Columbia; Whitcomb County, Washington, and on the Umpqua and Willamette Rivers, western Oregon. Southward passing into

Var. _Hindsiana_ Anders., a large shrub with numerous stems often 20° high, differing in its more linear or narrow lanceolate usually entire leaves on longer petioles, smaller aments and pubescent, not tomentose, branchlets; and distributed from the valleys of central California to southwestern Oregon. A shrubby form of _S. sessilifolia_ (var. _leucodendroides_ Schn.) with longer and broader leaves is common on the banks of streams in southern California.

12. Salix exigua Nutt.

Leaves lanceolate to oblanceolate, acuminate at the ends, often slightly falcate, minutely glandular-serrate above the middle, bluish green and glabrous above, covered below with appressed silky white hairs, 1½′—3′ long, ⅛′—¼′ wide, or on summer shoots sometimes 4½′ long and 1½′ wide; petioles glabrous, 1/16′ long or less; stipules minute or wanting. Flowers: aments terminal and solitary or terminal and axillary, on leafy glabrous branchlets, 1′—2′ in length; scales hoary pubescent, lanceolate and acute on staminate aments, often wider, obovate and rounded at the apex on pistillate aments; stamens, 2, filaments hairy below the middle; ovary sessile, villose, the stigmatic lobes sessile. Fruit ovoid, acuminate, glabrous.

A shrub with stems 10° or 12° tall, or rarely a tree 25° high, with a trunk 5′ or 6′ in diameter, thin spreading branches forming a round-topped head, and slender glabrous red-brown branchlets. Bark of the trunk thin, longitudinally fissured, grayish brown.

Distribution. Southern Alberta and valley of the Fraser River (Clinton), British Columbia, southward through western Washington and Oregon to San Diego County, California, and southeastern Nevada, and eastward to southern Idaho, central Nevada and western Wyoming (Yellowstone National Park).

Apparently only truly a tree on the banks of the Palouse and other streams of eastern Washington.

Several shrubby forms of _S. exigua_ found in Nevada, Arizona, Colorado, western Nebraska and in Lower California are distinguished.

13. Salix longifolia Muehl. Sand Bar Willow.

_Salix fluviatalis_ Sarg., not Nutt.

Leaves linear-lanceolate, often somewhat falcate, gradually narrowed at the ends, long-pointed, dentate with small remote spreading callous glandular teeth, 2′—6′ long, ⅛′—⅓′ wide, when they unfold coated below with soft lustrous silky hairs, at maturity thin, glabrous, light yellow-green, darker on the upper than on the lower surface, with a yellow midrib, slender arcuate primary veins, and slender reticulate veinlets; petioles grooved, ⅛′—¼′ long; stipules ovate-lanceolate, foliaceous, about ¼′ long, deciduous. Flowers: aments cylindric on leafy branchlets, pubescent, the staminate about 1′ long, ⅓′ broad, terminal and axillary, the pistillate elongated, 2′ or 3′ long, about ¼′ broad; scales obovate-oblong, entire, erose or dentate above the middle, light yellow-green, densely villose on the outer surface, slightly hairy on the inner; stamens 2, with free filaments slightly hairy at the base; ovary oblong-cylindric, acute, short-stalked, glabrous or pubescent, with large sessile deeply lobed stigmas. Fruit light brown, glabrous or villose, about ¼′ long.

A tree, usually about 20° high, with a trunk only a few inches in diameter, spreading by stoloniferous roots into broad thickets, short slender erect branches, and slender glabrous light or dark orange-colored or purplish red branchlets, growing darker after their first season; occasionally 60°—70° high, with a trunk 2° in diameter; often a shrub not more than 5°—6° tall. Winter-buds narrowly ovoid, acute, chestnut-brown, about ⅛′ long. Bark ⅛′—¼′ thick, smooth, dark brown slightly tinged with red and covered with small closely appressed irregularly shaped scales. Wood light, soft, light brown tinged with red, with thin light brown sapwood.

Distribution. River banks, sand bars and alluvial flats; shores of Lake St. John, Quebec to Manitoba, and southward through western New England to northeastern Virginia, southern Ohio, Indiana and Illinois, western Kentucky, south Tennessee, to the mouth of the Mississippi River, and westward to southwestern South Dakota, southwestern Wyoming, northeastern Colorado, western Kansas and Oklahoma, and northern Texas.

From central and northwestern Texas to northeastern Mexico and southern New Mexico represented by var. _angustissima_ Anders., differing in the absence of a dorsal gland in the male flowers and in the silky pubescence of the young ovary.

In the northern Rocky Mountains region replaced by var. _pedunculata_ Anders., differing from the type in its narrower linear leaves, glabrous ovaries and longer pedicels of the fruit, and ranging from western South Dakota and northwestern Wyoming, through eastern Montana, Saskatchewan, and Alberta, to the valley of the Yukon River in the neighborhood of Dawson.

A shrubby form with leaves densely covered with silky pubescence (var. _Wheeleri_ Schn.) is distributed from New Brunswick to North Dakota, Nebraska and Beckham County, Oklahoma.

14. Salix lasiolepis Benth. Arroyo Willow.

Leaves oblanceolate to lanceolate-oblong, often inequilateral and occasionally falcate, acute or acuminate or rarely rounded at apex, gradually or abruptly cuneate or rounded at base, entire or remotely serrate, pilose above and coated below with thick hoary tomentum when they unfold, at maturity thick and subcoriaceous, conspicuously reticulate-venulose, dark green and glabrous above, pale or glaucous and pubescent or puberulous below, 3′—6′ long, ½′—1′ wide, with a broad yellow midrib and slender arcuate veins forked and united within the slightly thickened and revolute margins; petioles slender, ⅛′—½′ long; stipules ovate, acute, coated with hoary tomentum, minute and caducous, or sometimes foliaceous, semilunar, acute or acuminate, entire or denticulate, dark green above, pale below, persistent. Flowers: aments erect, cylindric, slightly flexuose, densely flowered, nearly sessile on short tomentose branchlets, 1½′ long, the staminate ½′ thick, and nearly twice as thick as the pistillate; scales oblong-obovate, rounded or acute at the apex, dark-colored, clothed with long crisp white hairs, persistent under the fruit; stamens 2, with elongated glabrous filaments more or less united below the middle; ovary narrow, cylindric acute and long-pointed, dark green, glabrous, with a short style and broad nearly sessile stigmas. Fruit oblong-cylindric, light reddish brown, about ¼′ long.

A tree, 20°—35° high, with a trunk 3′—7′ in diameter, slender erect branches forming a loose open head, and stout branchlets coated at first with hoary tomentum, bright yellow or dark reddish brown and puberulous or pubescent during their first year, becoming darker and glabrous in their second season; or often at the north and at high altitudes a low shrub. Winter-buds ovoid, acute, compressed, contracted laterally into thin wing-like margins, light brownish yellow, glabrous or puberulous. Bark on young stems and on the branches thin, smooth, light gray-brown, becoming on old trunks dark, about ⅓′ thick, roughened by small lenticels and broken into broad flat irregularly connected ridges. Wood light, soft, close-grained, light brown, with thick nearly white sapwood; in southern California often used as fuel.

Distribution. Banks of streams in low moist ground; valley of the Klamath River, California, southward along the foothills of the Sierra Nevada, the central valley, and on the Coast Ranges to southern California; on Santa Catalina Island and on the mountains of southern Arizona; on the Sierra de Laguna, Lower California; occasionally ascending to altitudes of 4000° above the sea.

15. Salix Mackenzieana Barr.

Leaves lanceolate to oblanceolate, or elliptic, long-pointed at apex, cuneate or rounded at base, finely crenately serrate, reddish and pilose with caducous pale hairs when they unfold, at maturity thin and firm in texture, light green above, pale below, 1½′—2′ long, about ½′—¾′ wide, on summer shoots, often 4′ long and 1½′ wide, with a slender yellow midrib, arcuate veins, and obscure reticulate veinlets; petioles thin, yellow, about ⅓′ long; stipules reniform, conspicuously veined, about 1/16′ broad. Flowers: aments densely flowered, glabrous, erect, often more or less curved, about 1½′ long, terminal on short leafy branchlets; scales oblanceolate, acute, dark-colored; stamens 2, with elongated free glabrous filaments; ovary cylindric, long-stalked, elongated, gradually narrowed into a short style, with spreading emarginate stigmas. Fruit ovoid, acuminate, light brown, about ⅛′ long; pedicels about ⅙′ in length.

A small tree, with a slender trunk, upright branches forming a narrow shapely head, and slender branchlets marked with scattered lenticels, glabrous or slightly puberulous and often tinged with red when they first appear, soon becoming yellow and lustrous, growing lighter colored in their second year. Winter-buds ovoid, rounded on the back, compressed and acute at the apex, bright orange color, about ⅛′ long.

Distribution. Borders of streams and swamps; shores of Great Slave Lake southward through the region at the eastern base of the Rocky Mountains to Saskatchewan, northern Idaho, and northwestern Wyoming, and to western Nevada (Lake County; _M. S. Bebb_), and on the high Sierra Nevada in Calaveras and Mariposa Counties, California (_W. L. Jepson_).

16. Salix missouriensis Bebb.

Leaves lanceolate or oblanceolate, acuminate and long-pointed at apex, gradually narrowed from above the middle to the cuneate or rounded base, finely glandular-serrate, coated with pale hairs on the lower surface and pilose on the upper surface when they unfold, soon becoming nearly glabrous, at maturity thin and firm, dark green above, pale and often silvery white below, 4′—6′ long, 1′—1½′ wide, with slender veins often united near the margins and connected by coarse reticulate veinlets; petioles stout, pubescent or tomentose, ½′—¾′ long; stipules foliaceous, semicordate, pointed or rarely reniform and obtuse, serrate with incurved teeth, dark green and glabrous on the upper side, coated on the lower with hoary tomentum, reticulate-venulose, often ½′ long, deciduous or persistent during the season. Flowers: aments oblong-cylindric, densely flowered, appearing early in February on short leafy branchlets, the staminate 1½′ long and nearly ½′ wide and rather longer than the more slender pistillate aments becoming at maturity lax and 3′—4′ long; scales oblong-obovate, light green, and covered on the outer surface with long straight white hairs; stamens 2, with elongated free glabrous filaments; ovary cylindric, short-stalked, beaked, glabrous, with a short style and spreading entire or slightly emarginate stigmas. Fruit narrow, long-pointed, light reddish brown, ⅓′ in length; pedicels about half the length of the scales.

A tree, 40°—50° high, with a tall straight trunk 10′—12′ or rarely 18′ in diameter, rather slender upright slightly spreading branches forming a narrow open symmetrical head, and slender branchlets marked by small scattered orange-colored lenticels, light green and coated during their first year with thick pale pubescence, becoming reddish brown and glabrous or puberulous in their second winter. Winter-buds ovoid, round, or flattened, acute at the apex, reddish brown, hoary-tomentose, nearly 1′ long. Bark thin, smooth, light gray, slightly tinged with red, and covered with minute closely appressed plate-like scales. Wood dark red-brown, with thin pale sapwood; durable, used for fence-posts.

Distribution. Deep sandy alluvial bottom-lands of the Missouri River in southwestern Nebraska to western Missouri; through northeastern Kansas and eastern Oklahoma to Cache Creek, Comanche County (_G. W. Stevens_); and from the neighborhood of St. Louis to southeastern and western Iowa.

17. Salix pyrifolia Anders.

_Salix balsamifera_ Barr.

Leaves ovate to oblong-lanceolate, acute at apex, broad and rounded and usually subcordate at base, finely glandular serrulate, balsamic particularly while young, when they unfold thin, pellucid, red and coated below with long slender caducous hairs, at maturity thin and firm, dark green above, pale and glaucous below, 2′—4′ long, 1′—1½′ wide, with a yellow midrib and conspicuous reticulate veinlets; petioles reddish or yellow, ⅓′—½′ long; stipules often wanting or on vigorous shoots foliaceous, broadly ovate and acute. Flowers: aments cylindric, 1′—1½′ long, on short leafy branchlets, the staminate 1′—1¼′ long and ¾′ in diameter and shorter and broader than the pistillate ament; scales obovate, rose-colored, coated with long white hairs; stamens 2, with free filaments and reddish ultimately yellow anthers; ovary narrow-ovoid, long-stalked, gradually contracted above the middle, with a short style and emarginate stigmas. Fruit ovoid-conic, ¼′ long, dark orange color; pedicels ⅙′ in length.

Usually a shrub, often making clumps of crowded slender erect stems generally destitute of branches except near the top, rarely arborescent, with a height of 25°, a trunk 12′—14′ in diameter, erect branches, and comparatively stout reddish brown branchlets becoming olive-green in their second year and marked with narrow slightly raised leaf-scars. Winter-buds acute, much-compressed, bright scarlet, very lustrous, about ¼′ long. Bark thin, smooth, dull gray.

Distribution. Cold wet bogs; Newfoundland and the coast of Labrador to the valley of the Saskatchewan and the Mackenzie, and British Columbia, and to northern Maine, New Hampshire, Vermont, New York, Michigan, and northeastern South Dakota; reported to become arborescent only near Fort Kent on the St. John River, Aroostook, Maine.

18. Salix amplifolia Cov.

Leaves oval to broadly obovate, rounded or broadly pointed at apex, gradually or abruptly narrowed at the cuneate base, dentate-serrulate or entire, densely villose when they unfold, with long matted white hairs, at maturity nearly glabrous, pale yellow-green above, slightly glaucous below, 2′—2½′ long, 1′—1½′ wide, with a midrib broad and hoary-tomentose toward the base of the leaf and thin and glabrous above the middle; petioles slender, tomentose. Flowers: aments appearing about the middle of June, stout, pedunculate, tomentose, on leafy branchlets, the staminate 1½′—2′ long and shorter than the pistillate; scales oblanceolate or lanceolate, dark brown or nearly black, covered with long pale hairs; stamens 2, with slender elongated glabrous filaments; ovary ovoid-lanceolate, short-stalked, glabrous or slightly pubescent, gradually narrowed into the elongated slender style crowned with a 2-lobed slender stigma. Fruit ovoid-lanceolate, glabrous, short-stalked, ¼′ long.

A tree, occasionally 25° high, with a trunk a foot in diameter, and stout branchlets conspicuously roughened by the large elevated U-shaped leaf-scars, and marked by occasional pale lenticels, coated at first with thick villose pubescence, becoming during their second and third years dark dull reddish purple.

Distribution. Sand dunes on the shores of Yakutat Bay and Disenchantment Bay, Alaska.

19. Salix alaxensis Cov. Feltleaf Willow.