Chapter 17

Such being the general ferment in the minds of naturalists, it is no wonder that they mustered strong in the rooms of the Linnaean Society, on the 1st of July of the year 1858, to hear two papers by authors living on opposite sides of the globe, working out their results independently, and yet professing to have discovered one and the same solution of all the problems connected with species. The one of these authors was an able naturalist, Mr. Wallace, who had been employed for some years in studying the productions of the islands of the Indian Archipelago, and who had forwarded a memoir embodying his views to Mr. Darwin, for communication to the Linnaean Society. On perusing the essay, Mr. Darwin was not a little surprised to find that it embodied some of the leading ideas of a great work which he had been preparing for twenty years, and parts of which, containing a development of the very same views, had been perused by his private friends fifteen or sixteen years before. Perplexed in what manner to do full justice both to his friend and to himself, Mr. Darwin placed the matter in the hands of Dr. Hooker and Sir Charles Lyell, by whose advice he communicated a brief abstract of his own views to the Linnaean Society, at the same time that Mr. Wallace's paper was read. Of that abstract, the work on the 'Origin of Species' is an enlargement; but a complete statement of Mr. Darwin's doctrine is looked for in the large and well-illustrated work which he is said to be preparing for publication.

The Darwinian hypothesis has the merit of being eminently simple and comprehensible in principle, and its essential positions may be stated in a very few words: all species have been produced by the development of varieties from common stocks; by the conversion of these, first into permanent races and then into new species, by the process of NATURAL SELECTION, which process is essentially identical with that artificial selection by which man has originated the races of domestic animals--the STRUGGLE FOR EXISTENCE taking the place of man, and exerting, in the case of natural selection, that selective action which he performs in artificial selection.

The evidence brought forward by Mr. Darwin in support of his hypothesis is of three kinds. First, he endeavours to prove that species may be originated by selection; secondly, he attempts to show that natural causes are competent to exert selection; and thirdly, he tries to prove that the most remarkable and apparently anomalous phenomena exhibited by the distribution, development, and mutual relations of species, can be shown to be deducible from the general doctrine of their origin, which he propounds, combined with the known facts of geological change; and that, even if all these phenomena are not at present explicable by it, none are necessarily inconsistent with it.

There cannot be a doubt that the method of inquiry which Mr. Darwin has adopted is not only rigorously in accordance with the canons of scientific logic, but that it is the only adequate method. Critics exclusively trained in classics or in mathematics, who have never determined a scientific fact in their lives by induction from experiment or observation, prate learnedly about Mr. Darwin's method, which is not inductive enough, not Baconian enough, forsooth, for them. But even if practical acquaintance with the process of scientific investigation is denied them, they may learn, by the perusal of Mr. Mill's admirable chapter "On the Deductive Method," that there are multitudes of scientific inquiries in which the method of pure induction helps the investigator but a very little way.

"The mode of investigation," says Mr. Mill, "which, from the proved inapplicability of direct methods of observation and experiment, remains to us as the main source of the knowledge we possess, or can acquire, respecting the conditions and laws of recurrence of the more complex phenomena, is called, in its most general expression, the deductive method, and consists of three operations: the first, one of direct induction; the second, of ratiocination; and the third, of verification."

Now, the conditions which have determined the existence of species are not only exceedingly complex, but, so far as the great majority of them are concerned, are necessarily beyond our cognizance. But what Mr. Darwin has attempted to do is in exact accordance with the rule laid down by Mr. Mill; he has endeavoured to determine certain great facts inductively, by observation and experiment; he has then reasoned from the data thus furnished; and lastly, he has tested the validity of his ratiocination by comparing his deductions with the observed facts of Nature. Inductively, Mr. Darwin endeavours to prove that species arise in a given way. Deductively, he desires to show that, if they arise in that way, the facts of distribution, development, classification, etc., may be accounted for, i.e. may be deduced from their mode of origin, combined with admitted changes in physical geography and climate, during an indefinite period. And this explanation, or coincidence of observed with deduced facts, is, so far as it extends, a verification of the Darwinian view.

There is no fault to be found with Mr. Darwin's method, then; but it is another question whether he has fulfilled all the conditions imposed by that method. Is it satisfactorily proved, in fact, that species may be originated by selection? that there is such a thing as natural selection? that none of the phenomena exhibited by species are inconsistent with the origin of species in this way? If these questions can be answered in the affirmative, Mr. Darwin's view steps out of the rank of hypotheses into those of proved theories; but, so long as the evidence at present adduced falls short of enforcing that affirmation, so long, to our minds, must the new doctrine be content to remain among the former--an extremely valuable, and in the highest degree probable, doctrine, indeed the only extant hypothesis which is worth anything in a scientific point of view; but still a hypothesis, and not yet the theory of species.

After much consideration, and with assuredly no bias against Mr. Darwin's views, it is our clear conviction that, as the evidence stands, it is not absolutely proven that a group of animals, having all the characters exhibited by species in Nature, has ever been originate by selection, whether artificial or natural. Groups having the morphological character of species, distinct and permanent races in fact, have been so produced over and over again; but there is no positive evidence, at present, that any group of animals has, by variation and selective breeding, given rise to another group which was, even in the least degree, infertile with the first. Mr. Darwin is perfectly aware of this weak point, and brings forward a multitude of ingenious and important arguments to diminish the force of the objection. We admit the value of these arguments to their fullest extent; nay, we will go so far as to express our belief that experiments, conducted by a skilful physiologist, would very probably obtain the desired production of mutually more or less infertile breeds from a common stock, in a comparatively few years; but still, as the case stands at present, this "little rift within the lute" is not to be disguised nor overlooked.

In the remainder of Mr. Darwin's argument our own private ingenuity has not hitherto enabled us to pick holes of any great importance; and judging by what we hear and read, other adventurers in the same field do not seem to have been much more fortunate. It has been urged, for instance, that in his chapters on the struggle for existence and on natural selection, Mr. Darwin does not so much prove that natural selection does occur, as that it must occur; but, in fact, no other sort of demonstration is attainable. A race does not attract our attention in Nature until it has, in all probability, existed for a considerable time, and then it is too late to inquire into the conditions of its origin. Again, it is said that there is no real analogy between the selection which takes place under domestication, by human influence, and any operation which can be effected by Nature, for man interferes intelligently. Reduced to its elements, this argument implies that an effect produced with trouble by an intelligent agent must, a fortiori, be more troublesome, if not impossible, to an unintelligent agent. Even putting aside the question whether Nature, acting as she does according to definite and invariable laws, can be rightly called an unintelligent agent, such a position as this is wholly untenable. Mix salt and sand, and it shall puzzle the wisest of men, with his mere natural appliances, to separate all the grains of sand from all the grains of salt; but a shower of rain will effect the same object in ten minutes. And so, while man may find it tax all his intelligence to separate any variety which arises, and to breed selectively from it, the destructive agencies incessantly at work in Nature, if they find one variety to be more soluble in circumstances than the other, will inevitably, in the long run, eliminate it.

A frequent and a just objection to the Lamarckian hypothesis of the transmutation of species is based upon the absence of transitional forms between many species. But against the Darwinian hypothesis this argument has no force. Indeed, one of the most valuable and suggestive parts of Mr. Darwin's work is that in which he proves, that the frequent absence of transitions is a necessary consequence of his doctrine, and that the stock whence two or more species have sprung, need in no respect be intermediate between these species. If any two species have arisen from a common stock in the same way as the carrier and the pouter, say, have arisen from the rock-pigeon, then the common stock of these two species need be no more intermediate between the two than the rock-pigeon is between the carrier and pouter. Clearly appreciate the force of this analogy, and all the arguments against the origin of species by selection, based on the absence of transitional forms, fall to the ground. And Mr. Darwin's position might, we think, have been even stronger than it is if he had not embarrassed himself with the aphorism, "Natura non facit saltum," which turns up so often in his pages. We believe, as we have said above, that Nature does make jumps now and then, and a recognition of the fact is of no small importance in disposing of many minor objections to the doctrine of transmutation.

But we must pause. The discussion of Mr. Darwin's arguments in detail would lead us far beyond the limits within which we proposed, at starting, to confine this article. Our object has been attained if we have given an intelligible, however brief, account of the established facts connected with species, and of the relation of the explanation of those facts offered by Mr. Darwin to the theoretical views held by his predecessors and his contemporaries, and, above all, to the requirements of scientific logic. We have ventured to point out that it does not, as yet, satisfy all those requirements; but we do not hesitate to assert that it is as superior to any preceding or contemporary hypothesis, in the extent of observational and experimental basis on which it rests, in its rigorously scientific method, and in its power of explaining biological phenomena, as was the hypothesis of Copernicus to the speculations of Ptolemy. But the planetary orbits turned out to be not quite circular after all, and, grand as was the service Copernicus rendered to science, Kepler and Newton had to come after him. What if the orbit of Darwinism should be a little too circular? What if species should offer residual phenomena, here and there, not explicable by natural selection? Twenty years hence naturalists may be in a position to say whether this is, or is not, the case; but in either event they will owe the author of 'The Origin of Species' an immense debt of gratitude. We should leave a very wrong impression on the reader's mind if we permitted him to suppose that the value of that work depends wholly on the ultimate justification of the theoretical views which it contains. On the contrary, if they were disproved to-morrow, the book would still be the best of its kind--the most compendious statement of well-sifted facts bearing on the doctrine of species that has ever appeared. The chapters on Variation, on the Struggle for Existence, on Instinct, on Hybridism, on the Imperfection of the Geological Record, on Geographical Distribution, have not only no equals, but, so far as our knowledge goes, no competitors, within the range of biological literature. And viewed as a whole, we do not believe that, since the publication of Von Baer's Researches on Development, thirty years ago, any work has appeared calculated to exert so large an influence, not only on the future of Biology, but in extending the domination of Science over regions of thought into which she has, as yet, hardly penetrated.

End of The Origin of Species.

CRITICISMS ON "THE ORIGIN OF SPECIES".*

([Footnote] *'The Natural History Review', 1864.)

1. UEBER DIE DARWIN'SCHE SCHOPFUNGSTHEORIE; EIN VORTRAG, VON A. KOLLIKER. Leipzig, 1864.

2. EXAMINATION DU LIVRE DE M. DARWIN SUR L'ORIGINE DES ESPECES. PAR P. FLOURENS. Paris, 1864.

In the course of the present year several foreign commentaries upon Mr. Darwin's great work have made their appearance. Those who have perused that remarkable chapter of the 'Antiquity of Man,' in which Sir Charles Lyell draws a parallel between the development of species and that of languages, will be glad to hear that one of the most eminent philologers of Germany, Professor Schleicher, has, independently, published a most instructive and philosophical pamphlet (an excellent notice of which is to be found in the 'Reader', for February 27th of this year) supporting similar views with all the weight of his special knowledge and established authority as a linguist. Professor Haeckel, to whom Schleicher addresses himself, previously took occasion, in his splendid monograph on the 'Radiolaria',* to express his high appreciation of, and general concordance with, Mr. Darwin's views. ([Footnote] *'Die Radiolarien: eine Monographie', p. 231.)

But the most elaborate criticisms of the 'Origin of Species' which have appeared are two works of very widely different merit, the one by Professor Kolliker, the well-known anatomist and histologist of Wurzburg; the other by M. Flourens, Perpetual Secretary of the French Academy of Sciences.

Professor Kolliker's critical essay 'Upon the Darwinian Theory' is, like all that proceeds from the pen of that thoughtful and accomplished writer, worthy of the most careful consideration. It comprises a brief but clear sketch of Darwin's views, followed by an enumeration of the leading difficulties in the way of their acceptance; difficulties which would appear to be insurmountable to Professor Kolliker, inasmuch as he proposes to replace Mr. Darwin's Theory by one which he terms the 'Theory of Heterogeneous Generation.' We shall proceed to consider first the destructive, and secondly, the constructive portion of the essay.

We regret to find ourselves compelled to dissent very widely from many of Professor Kolliker's remarks; and from none more thoroughly than from those in which he seeks to define what we may term the philosophical position of Darwinism.

"Darwin," says Professor Kolliker, "is, in the fullest sense of the word, a Teleologist. He says quite distinctly (First Edition, pp. 199, 200) that every particular in the structure of an animal has been created for its benefit, and he regards the whole series of animal forms only from this point of view."

And again:

"7. The teleological general conception adopted by Darwin is a mistaken one.

"Varieties arise irrespectively of the notion of purpose, or of utility, according to general laws of Nature, and may be either useful, or hurtful, or indifferent.

"The assumption that an organism exists only on account of some definite end in view, and represents something more than the incorporation of a general idea, or law, implies a one-sided conception of the universe. Assuredly, every organ has, and every organism fulfils, its end, but its purpose is not the condition of its existence. Every organism is also sufficiently perfect for the purpose it serves, and in that, at least, it is useless to seek for a cause of its improvement."

It is singular how differently one and the same book will impress different minds. That which struck the present writer most forcibly on his first perusal of the 'Origin of Species' was the conviction that Teleology, as commonly understood, had received its deathblow at Mr. Darwin's hands. For the teleological argument runs thus: an organ or organism (A) is precisely fitted to perform a function or purpose (B); therefore it was specially constructed to perform that function. In Paley's famous illustration, the adaptation of all the parts of the watch to the function, or purpose, of showing the time, is held to be evidence that the watch was specially contrived to that end; on the ground, that the only cause we know of, competent to produce such an effect as a watch which shall keep time, is a contriving intelligence adapting the means directly to that end.

Suppose, however, that any one had been able to show that the watch had not been made directly by any person, but that it was the result of the modification of another watch which kept time but poorly; and that this again had proceeded from a structure which could hardly be called a watch at all--seeing that it had no figures on the dial and the hands were rudimentary; and that going back and back in time we came at last to a revolving barrel as the earliest traceable rudiment of the whole fabric. And imagine that it had been possible to show that all these changes had resulted, first, from a tendency of the structure to vary indefinitely; and secondly, from something in the surrounding world which helped all variations in the direction of an accurate time-keeper, and checked all those in other directions; then it is obvious that the force of Paley's argument would be gone. For it would be demonstrated that an apparatus thoroughly well adapted to a particular purpose might be the result of a method of trial and error worked by unintelligent agents, as well as of the direct application of the means appropriate to that end, by an intelligent agent.

Now it appears to us that what we have here, for illustration's sake, supposed to be done with the watch, is exactly what the establishment of Darwin's Theory will do for the organic world. For the notion that every organism has been created as it is and launched straight at a purpose, Mr. Darwin substitutes the conception of something which may fairly be termed a method of trial and error. Organisms vary incessantly; of these variations the few meet with surrounding conditions which suit them and thrive; the many are unsuited and become extinguished.

According to Teleology, each organism is like a rifle bullet fired straight at a mark; according to Darwin, organisms are like grapeshot of which one hits something and the rest fall wide.

For the teleologist an organism exists because it was made for the conditions in which it is found; for the Darwinian an organism exists because, out of many of its kind, it is the only one which has been able to persist in the conditions in which it is found.

Teleology implies that the organs of every organism are perfect and cannot be improved; the Darwinian theory simply affirms that they work well enough to enable the organism to hold its own against such competitors as it has met with, but admits the possibility of indefinite improvement. But an example may bring into clearer light the profound opposition between the ordinary teleological, and the Darwinian, conception.

Cats catch mice, small birds and the like, very well. Teleology tells us that they do so because they were expressly constructed for so doing--that they are perfect mousing apparatuses, so perfect and so delicately adjusted that no one of their organs could be altered, without the change involving the alteration of all the rest. Darwinism affirms on the contrary, that there was no express construction concerned in the matter; but that among the multitudinous variations of the Feline stock, many of which died out from want of power to resist opposing influences, some, the cats, were better fitted to catch mice than others, whence they throve and persisted, in proportion to the advantage over their fellows thus offered to them.

Far from imagining that cats exist IN ORDER to catch mice well, Darwinism supposes that cats exist BECAUSE they catch mice well--mousing being not the end, but the condition, of their existence. And if the cat type has long persisted as we know it, the interpretation of the fact upon Darwinian principles would be, not that the cats have remained invariable, but that such varieties as have incessantly occurred have been, on the whole, less fitted to get on in the world than the existing stock.

If we apprehend the spirit of the 'Origin of Species' rightly, then, nothing can be more entirely and absolutely opposed to Teleology, as it is commonly understood, than the Darwinian Theory. So far from being a "Teleologist in the fullest sense of the word," we would deny that he is a Teleologist in the ordinary sense at all; and we should say that, apart from his merits as a naturalist, he has rendered a most remarkable service to philosophical thought by enabling the student of Nature to recognise, to their fullest extent, those adaptations to purpose which are so striking in the organic world, and which Teleology has done good service in keeping before our minds, without being false to the fundamental principles of a scientific conception of the universe. The apparently diverging teachings of the Teleologist and of the Morphologist are reconciled by the Darwinian hypothesis.

But leaving our own impressions of the 'Origin of Species,' and turning to those passages especially cited by Professor Kolliker, we cannot admit that they bear the interpretation he puts upon them. Darwin, if we read him rightly, does 'not' affirm that every detail in the structure of an animal has been created for its benefit. His words are (p. 199):--

"The foregoing remarks lead me to say a few words on the protest lately made by some naturalists against the utilitarian doctrine that every detail of structure has been produced for the good of its possessor. They believe that very many structures have been created for beauty in the eyes of man, or for mere variety. This doctrine, if true, would be absolutely fatal to my theory--yet I fully admit that many structures are of no direct use to their possessor."

And after sundry illustrations and qualifications, he concludes (p. 200):--

"Hence every detail of structure in every living creature (making some little allowance for the direct action of physical conditions) may be viewed either as having been of special use to some ancestral form, or as being now of special use to the descendants of this form--either directly, or indirectly, through the complex laws of growth."