Part 34

These clusters are larger or smaller, according to the species of the vorticellæ which form them, as well as owing to the concurrence of many other circumstances. To obtain a clear idea of the figure of these animals, it is best to observe the smaller clusters, as in the larger they are often rendered less distinct on account of the number.

The length of those which are represented at Fig. 30, is about the 240th of an inch; they are of a bell-shape. The anterior part a c generally appears open, the posterior part is fixed to a stem or pedicle, b e; it is by the extremity of this pedicle that the vorticella fastens itself to any substance. It appears in the microscope of a brownish colour, excepting at the smaller end b, where it is transparent, as well as the whole pedicle b e. When the anterior part a c is open, a very lively motion may be perceived about its edges; and when it presents itself in a particular manner, something very much resembling the little wheels of a mill, moving with great velocity, may be discovered on both sides of the edges of this anterior part.

These vorticellæ are able to contract themselves suddenly. They may be made to do this, either by touching them, or moving the substance to which they are fixed. When they contract, the edges of the anterior parts are drawn quite into the body; on resuming their former posture, the edges may be seen to come forth, and put themselves in motion as before. Minute substances that float in the water are often forced down into these openings, and sometimes are thrown out again.

They are capable of swimming about singly, but their form is in that case considerably different from that which they have when they are fixed. To see regularly in what manner the clusters are formed, and in what way these little creatures multiply, it is best to observe one that is fixed by itself.

The pedicle of a single vorticella is at first short, but it soon grows longer, and then begins to multiply, that is, to divide or split itself into two lengthways. To effect this, the lips are first drawn into the body, the anterior part closes and becomes round, and loses its bell shape, the motion about the lips ceases, though a small degree of motion may be perceived within the body. The anterior end flattens gradually, and spreads wider in proportion as it grows smaller. It then gradually splits down the middle, that is, from the middle of the head to the pedicle, so that in a little time two separate round bodies appear to be joined to the end of the pedicle that before supported but one.

The mouth or anterior part of each of these bodies now opens by degrees; and in proportion as they open, the lips of the new vorticella begin to display themselves. The motion before spoken of may then also be perceived. Indeed it is the best time of observing it; it is at first slow, but more rapid in proportion as the mouth opens, when it is as swift as that of the vorticella before it began to divide, and we may now look upon it as completely formed. A vorticella is generally about one hour in dividing itself.

The lower of the three drawings, Fig. 30, represents two vorticellæ joined by their posterior extremity to one pedicle; soon after the division, each vorticella begins to shew a pedicle of its own.

Fig. 30 represents a cluster of eight vorticellæ; by this figure we may form some idea in what manner the pedicles are disposed as their number increases. There were at first only two at b, whose branches lengthened to d, and then each of them was divided into two, now forming four; these again lengthened and reached i; when they were again subdivided, as in the figure.

The reader will join with Bonnet in admiring the group of wonders afforded by a single spot of mouldiness. What unforeseen, varied, and interesting scenes are presented within so small a compass! what a theatre is exhibited to a thinking mind! But our abode is so recluse, that we have but a glimmering view of it: how great would our astonishment be, if the whole spectacle was disclosed to us at once, and we were enabled to penetrate into the interior structure of this wonderful assemblage of living atoms! Our eyes see only the gross parts of the decorations, whilst the machines that execute them remain in impenetrable darkness! Who shall enlighten this profound obscurity, or dive into an abyss where reason is lost; or draw from thence the treasures of wisdom concealed within it? Let us learn to be content with the small portion that is communicated to us, and contemplate with gratitude the first traces of human understanding that are imparted to us in these discoveries.

VORTICELLA BERBERINA.

Vorticella composita, floribus ovalibus muticis, stirpe ramosa. Compound, with oval beardless florets.

This is a species of the vorticellæ, which resembles the preceding one in many respects, particularly in being multiplied in the same manner, that is, by dividing or splitting, according to its length.

They are more slender than the vorticella umbellaria; the branches of the clusters are transparent. When many of them are together, they appear of a changeable violet colour; the clusters are not unlike a sprig of spun glass. The motion of the lips is not so easily distinguished as in the foregoing species, though it may be observed in these whilst they are opening and completing their formation. For at these times the motion is but slow, whereas it becomes afterwards very quick in those that are arrived at a state of perfection.

All the cluster vorticellæ detach themselves from time to time from the stem, and from these they swim about till they fix again upon some convenient substance; the branches, when deserted, bear no more vorticellæ.

VORTICELLA DIGITALIS.

Plate XXII. Fig. 31.

Vorticella composita, floribus cylindricis, unisulcatis semiclausis, stirpe ramosa. Compound, with cylindrical florets.

Vorticella composita, cylindrica, crystallina, apice truncata et fissa, pedunculo fistuloso ramosa. Müller Animal. Infus. p. 327.

This species of the vorticella is very scarce, it seems only to have been seen by Rösel, who found it on the monoculus quadricornis, till it was discovered in the year 1784 by Müller, who had sought for it several years before, but in vain.

The body is cylindrical, crystalline, and appears almost empty; it has three pellucid points disposed lengthways, the apex is truncated in an oblique direction, the margin bent back. The upper part contracts itself, and the margin then assumes a conical shape, with a convex surface; there are in general but few branches from the principal stem, and these are short and thick. It excites an undulatory motion, but no hairs, nor any rotatory motion, have been discovered. Fig. 31, o and n, represents the vorticella adhering to the monoculus quadricornis.

VORTICELLA CONVALLARIA.

Plate XXII. Fig. 39.

Vorticella simplex, gregaria, flore campanulata mutico; tentaculis bigeminis, stirpe fixa. Simple, but gregarious, the florets bell-shaped, with two pair of little arms, and a fixed stem.

Vorticella simplex, campanulata, pedunculo rotortili. Müller Animal. Infus.

These vorticellæ, or bell-animals, as they are termed by Baker, are generally found adhering to some substance in the water; they are represented here as found by Rösel, fixed to a curious cornu ammonis, with points projecting from the back. To the naked eye they appear only as so many little white points, but under the microscope, as little bells, agitating the water to a considerable distance. The stems of these have a particular motion, they draw themselves up and shorten all at once, taking the form of a spiral wire or screw; in a moment after they again resume their former shape, stretching themselves out straight as before. Many of them may be seen at times adhering to each other by their tails; the cilia, which are two on each side of the mouths, are very seldom to be perceived.

VORTICELLA URCEOLARIS.

Plate XXII. Fig. 33, 34, 35, 36, 37, 38.

Vorticella simplex, pedunculata, ore dentato. Single, with a short tail, and toothed mouth.

Brachionus capsularis testa ovata apice sexdentata basi incisa, cauda longa bicuspi. Müller Animal. Infus. p. 356.

To the naked eye it appears as a white moveable point; but when examined by the microscope, a tail projecting from the lower part is discovered, and a double rotatory instrument is seen, which it can conceal or expose at pleasure. It has been seen and described by most microscopical writers; but as Baker’s seems to be the most perfect description, I shall principally follow his account of it.

He discovered three species of them, two of which are included under the vorticella urceolaris. Fig. 33, 34, 35, are of the first species; Fig. 36, 37, 38, are of the second kind. The first sort, when extended, is about twice as long as it is broad. It is contained in a shell; the fore part of this is armed with four sharp teeth or points; the opposite side has no teeth, but is waved or bent in two places, like the form of a Turkish bow. At the bottom there is a hole, through which it pushes the tail. It fastens itself by this tail to any convenient substance when it intends to use its rotatory organs; but when it is floating in the water, and at all other times when not adhering to any body, it wags the tail backwards and forwards something like a dog.

We may consider it as divided into a head, thorax, and abdomen; each of which may be extended and contracted considerably: it can, by dilating all three, protrude the head beyond the shell, or by contracting them, draw the whole body within the same.

The head, when extended, divides itself into two branches, between which, another part, a kind of proboscis, is pushed out; at the end of this are two fibrils, that appear when they are at rest like a broad point, but which can be moved to and from each other very briskly with a vibratory motion, see Fig. 33.

The form and situation of the two branches are sometimes changed, the ends thereof becoming more round, and the vibratory motion is altered to a rotatory one: this alteration is represented at Fig. 34: the head also appears in this figure. The thorax is annexed to the lower part of the head; it is muscular: within it there is a moving intestine, which has been supposed to be either the lungs or the heart of the little creature, see b, Fig. 33 and 34.

A communication is formed between the thorax and the abdomen by means of a short vessel c, whose alternate contractions and dilatations occasion the abdomen to rise and fall alternately, having at the same time a sort of peristaltic motion. The food is conveyed through this vessel into the abdomen, where it is digested; it is then discharged by the anus, which is placed near the tail.

The tail has three joints, and is cleft or divided at the extremity, by which means it can better fasten itself to suitable objects. It is in general projected from the lower end of the shell, moving nimbly to and fro, serving the animal as a rudder when it is swimming, to direct its course.

When the water in which the little animal is placed is nearly dried away, or when it has a mind to compose itself to rest, it contracts the head and fore-part of the body, brings them down into the shell, and pulls the tail upwards, so that the whole of this minute creature is contained within the shell, see Fig. 35. The shell is so transparent that the terminations cannot be easily distinguished when the animal is extended; but whatever is transacted within the shell, is as plain as if there was no substance between the eye and the interior parts.

Fig. 36, 37, 38, exhibit the appearance of another species of these animals, which differs from the foregoing kind. This has also a head, a thorax, and abdomen, but then they are not separated by a gut or intermediate vessel, as in the former, but are joined immediately together, and at the place where in the first kind a moveable intestine was seen; in this a muscle, most probably the heart, may be discovered; it has a regular systole and diastole: this part is intended to be shewn at a, Fig. 36, 37, 38. Like the other, it draws the head and tail within the shell, which then appears to have six teeth or spikes on one side, and two on the other. It very seldom protrudes its head so far out as the other; sometimes the fibrillæ may be seen within the margin of the shell.

Both species carry their young in an oval integument or bag, fastened externally to the lower part of the shell, somewhere about the tail; these bags are sometimes opake at one end, and seemingly empty at the other, see d, Fig. 34: sometimes the middle is opake, with a transparent margin, see b, Fig. 36.

It is highly entertaining to see a young one burst its integument, and gradually force its way out; in performing this operation, it is much assisted by the motion of the tail of the parent. The head part comes out first, it then sets its rotatory organ in motion, by which it is completely disengaged, leaving the integument behind, which the vorticella freed itself from by repeated strokes with its tail. A young one almost disengaged is seen at b, Fig. 38; another embryo, c, was left adhering to the shell.

There are four more species of the vorticellæ mentioned by Linnæus, which are, the vorticella encrinus, the vorticella polypina, the vorticella stellata, and the vorticella ovifera; which, being marine animals, do not come properly within our plan. The vorticella polypina will be described hereafter.

TUBULARIA CAMPANULATA.

Plate XXII. Fig. 32.

Tubularia reptans, tubis campanulatis. Creeping, with campanulated tubes.

It is called by Baker the bell-flowered, or plumed animal.

These little creatures dwell in colonies together, from ten to fifteen in number, living in a kind of slimy mucilaginous case, which, when expanded in the water, has some resemblance to a bell with its mouth upwards. These bells or colonies are to be found adhering to the large leaves of duck-weed and other aquatic plants.

The bell or case which these animals inhabit, being very transparent, all the motions of its inhabitants may be discerned distinctly through it. There are several ramifications or smaller bells proceeding from the larger one; in each of these there is an inhabitant. The opening at the top of these bells is just large enough for the creature’s head, and a small part of its body to be thrust out from it, the rest remaining in the case, into which it also draws the head on the least alarm.

Besides the particular and separate motions which each of these creatures is able to exert within its case, and independent of the rest, the whole colony has a power of altering the position of the bell, and removing it from one place to another. These animalcula seem not to like to dwell in societies, whose number exceeds fifteen; when the colony happens to increase in number, the bell may be observed to split gradually, beginning from about the middle of the upper extremity, and proceeding downwards towards the bottom, till they at last separate and become two colonies, independent of each other.

The arms are very near each other; sixty may often be counted in one plume, having each the figure of an Italic _ʃ_, one of whose hooked ends is fastened to the head; and altogether, when expanded, compose a figure somewhat like a horseshoe, convex on the side next the body, but gradually opening and turning outwards, so as to leave a considerable distance within the outer extremities of the arms.

The plumed polype is of a very voracious disposition, devouring a great number of small animals. If the arms, when extended, be observed attentively with the microscope, they will be found to have a constant vibratory motion; alternately bending withinside of the plume, and then rising up again. When one arm ceases its motion, the same is performed by another; thus by the perpetual agitation of the several arms, such a strong current is produced in the water, as brings the animalcula, and other minute bodies, that are floating near the polype, into its mouth, which is situated between the arms. The food, if agreeable to the creature, is swallowed; if otherwise, it is rejected by a contrary motion.

The animal may be seen very plain when it has retired within the tube. The body is about one-eighth of an inch long, without reckoning the plume, which is about the same length. It is cylindrical, and the skin is very transparent. The plume is only a continuation of this transparent skin, it is very broad in proportion to the body, and of a remarkable figure; the base is of the shape of a horseshoe; from this base the arms project, they bend rather outwards. The plume which they form, gives them a resemblance to some flowers. The arms may be compared, from their fineness and transparency, to very fine threads of glass. The base of the plume is grooved, and is fixed to the animal by the middle of the horseshoe which it forms, and it is here that there is an opening which serves as a mouth to the animal. The intestines are easily distinguished through its transparent skin; when it has just been eating, they are of a deep brown colour. Three principal parts are very visible, the oesophagus, the stomach, and the rectum.

In the inside of these animals a small oblong whitish body is formed, which is carried to the outside, and remains fixed in a perpendicular direction to the body; many of these are formed daily, and of these oval bodies new animals are produced, exactly similar to the parent.

If these minute bodies be eggs, they are of a singular kind, being destitute of any covering, and are neither membranaceous nor crustaceous; we cannot with propriety say the young ones are hatched from them; we can, however, perceive these oviform bodies to unfold themselves gradually. The developement is accomplished in a few minutes, and an animalculum appears like the parent.

Trembley amassed a great number of these eggs, and carried them from England with him, keeping them quite dry; on putting them into water, they gradually developed, and became as perfect as the tubularia from which they proceeded.

There is a very great similarity in the construction of this little creature and many of the marine polypes, who, like it, exist in tubes of the same growth with themselves.

Fig. 32 represents three tubulariæ campanulatæ or plumed polypes very much magnified, namely, one, b f a c d d e h g i, which is out of its cell; e h, the oesophagus; f g, the stomach; a f, the rectum; a c d d e, the plume, consisting of the base a e, which is but little seen, and the arms c d d, which proceed from the edges of this base; a second polype, _A B I_, which is within its cell, and in which the skin containing the plume is reversed. The third polype, s t u u, is a young one exhibited out of its cell; g o o, threads which are fixed at one end to the intestines of the animal, by the other to the bottom of the cell, l k.

CHAP. VIII.

OF THE ANIMALCULA INFUSORIA.

Our knowledge of the microscopic world is at present very contracted, but we know enough to give us high conceptions of its concealed wonders, and to fill us with profound astonishment at the infinite variety of forms that are made recipient of life. A few of the inhabitants of this minute world have been discovered. The figure and apparent habits of life of these, resemble so little those with which we are more acquainted, that it is often difficult to find terms to express what is represented to the eye.

Animalculum signifies a little animal, and therefore the term might be applied to every animal which is considerably inferior in size to ourselves. It has been customary, however, to distinguish by the name of animalcula only such animals as are of a size so diminutive, that their true figure cannot be discerned without the assistance of glasses; and more especially it is applied to such as are altogether invisible to the naked eye, and cannot even be perceived to exist, but by the aid of microscopes.

By the help of magnifying glasses we are brought into a kind of new world; and numberless animals are discovered, which, from their minuteness, must otherwise for ever have escaped our observation: and how many kinds of these invisibles there may be, is yet unknown; as they are observed of all sizes, from those which are barely invisible to the naked eye, to such as resist the action of the microscope, as the fixed stars do that of the telescope, and with the best magnifiers hitherto invented, appear only as so many moving points.

The smallest living creatures our instruments can shew, are those that inhabit the waters; for, though possibly animalcula equally minute, or perhaps more so, may fly in the air, or creep upon the earth, it is scarce possible to obtain a view of them; whereas, water being transparent, and confining the creatures within it, we are enabled, by applying a drop of it to our glasses, to discover with ease a great part of its contents, and in a space barely visible to the naked eye, often perceive a thousand little creatures, all full of life and vigour.

By the animalcula infusoria are meant, not the larvæ of those insects which in their first state are inhabitants of water, and afterwards become winged insects, as the gnat, &c. Baker, and many other writers on the subject, have often confounded these, and hence entered into a train of reasoning contrary to fact and experience. The animalcula infusoria take their name from their being found in all kinds either of vegetable or animal infusions; if seeds, herbs, or other vegetable substances, be infused in water, it will soon be filled with an indefinite number of these minute beings. There is a prodigious variety in their forms; some perfectly resemble the bell-polype; others are round or oblong, without any, at least apparent, members; some resemble a bulb with a long taper tail; some are nearly spherical; the greater part are vesicular and transparent. Those most generally found in every drop of ditch water are mere inflated bladders, with a small trace of intestines in the center; the next are a flat kind, with a number of legs under the belly.

Motion seems to be their great delight; they pervade with equal ease and rapidity, and in all forms and directions, the whole dimensions of the drop, in which they find ample space for their various progressions, sometimes darting straight forward, at other times moving obliquely, then again circularly: they know how to avoid with dexterity any obstacles that might obstruct their progress. Hundreds may be seen in a drop of water in constant action, yet never striking against each other. If at any time the clusters prove so thick as to impede any of their motions, they roll and tumble themselves over head, creep under the whole range, force their way through the midst, or wheel round the cluster, with surprizing swiftness; sometimes they will suddenly change the direction in which they are moving, and take one diametrically opposite thereto. By inclining the glass on which the drop of water is laid, it may be made to move in any direction; the animalcula in the drop will swim as easily against the stream as with it.

If the water begin to evaporate, and the drop to grow smaller, they flock impetuously towards the remaining part of the fluid; an anxious desire of attaining this momentary respite of life is very visible, as well as an uncommon agitation of the organs by which they imbibe the water. These motions grow more languid as the water fails, till at last they entirely cease.

Animalcula and insects will support a great degree of cold, but both one and the other perish when it is carried beyond a certain point. The same degree of heat that destroys the existence of insects, is fatal to animalcula; as there are animalcula produced in water at the freezing point, so there are insects which live in snow.