c. In respect to the members of the trunk, the turnip is perhaps the
genuine root, the tap-root the stalk-root, the so-called aerial roots, the leaf-roots.
2. EARTH-ORGAN.--STALK.
1096. The stalk is the idea of the whole plant, posited under the import of the organ of nutrition, of the vessels. The structure of the stalk is therefore accordant with that of the root. The anatomical systems are alike in both, bark, liber and tracheæ being in the same envelope.
1097. In the stalk, however, the opposition of tissues and systems emerges more strongly, and therefore they all become individualized also to a higher degree. The spiral vessels become more freed from the cellular tissue; the bark is more distinctly divided from the liber; this again from the wood, and in its centre the cellular tissue dries up into pith. As, however, the stalk is the first product of the light-influence, the tracheal system cannot attain as yet entire freedom. The cells have nevertheless been extended, and the intercellular passages are formed into regular tubes. The _Stalk_ is the trunk of the plant with a preponderating system of tubes.
1098. This separation of the tissues and systems, with the endeavour upon the part of each to become individually perfect and isolated from the other, is effected by the air and the light. As root imbibes the mucous or slimy water, and sustains the chemical process in the plant, so does the stalk set the water in motion, since it exposes it to the air and light, whereby the chemical earth-process becomes separated into different saps and elemental bodies.
1099. Through the influence of the air, light and heat upon the stalk, as well as by its antagonism with the root, its elongation is determined. The greater amount of energy of the aerial polarity is in the higher regions; it is thus more excited by these than by the lower, and the bud being lighter grows more rapidly in the upper parts, and obtains an elongated form. The excitation is stronger also upon the side exposed to the light. It grows also more strongly in that direction, and thus the stalk stands indeed upwards, but somewhat inclined from the perpendicular line towards the sun.
1100. The perpendicular direction of the stalk is, however, as mechanically determined by gravity as the root. Paradoxical as this assertion may appear, it is still correct; for if we think of a moist globe, which is superiorly affected by air, light and heat, the upper aqueous or mucous parts are thus lighter, and necessarily ascend upwards through the pressure of the heavy or unheated parts, just as the air-bubbles in beer ascend to the surface. The light it is true can draw them somewhat sideways, chiefly because this side is more heated, more decomposed, and also undergoes a greater amount of evaporation; but the proper cause invariably resides in the ascent of the light parts between the heavy or immoveable. The stalk therefore grows upwards also in the darkness, and then indeed quite perpendicularly, because it has not been diverted by light. Were the light merely the Dirigent, it could not be comprehended, why towards the poles trees still stand tolerably perpendicular, and do not lie completely upon the earth. Were, however, the air that which determines the direction, plants could not thus have been inclined towards the sun, nor could we comprehend why the flowers and also the leaves obey the sun's course. Finally, were neither air nor light the imparters of direction, then the plant could shoot in no other direction than quite perpendicularly upwards, as is done too for the most part by the fungi.
1101. The _winding_ of the stalk appears to originate from the rotation of the sun. The next cause is probably the greater heating and decomposition that occurs upon one side. In accordance with this assumption the plants upon the northern hemisphere of the earth must wind spirally upwards from left to right, or from morning towards evening, if our gaze be directed towards the meridian, but inversely upon the southern hemisphere. But this is not the case. May we therefore conjecture that plants, twisted contrary to rule, have been transported from their native soil?
1102. The kinds of stalk follow also, without doubt, the direction of developmental stages of the vegetable trunk. There are therefore cellular stalks in the fungi, vascular stalks in the mosses, tracheal stalks in the ferns. A bark-stalk is the culm, a liber-stalk the scape, a wood-stalk probably the trunk of a palm. A root-stalk is the rhizoma, a perfect-stalk the stem, a leaf-stalk being probably the shrub.
_Ramification._
1103. The differencing, severing character of air and light must never be lost from our thoughts, nor also that at a height this character manifests itself more powerfully than on the surface of the earth, where the stem abandons its androgynous position in relation to the root. Through the constant process of differencing the tracheal fascicles of the plant may finally become so independent that they no longer stand in need of the others, and do not merely represent a particular plant, but are perfected also as such.
1104. This dispersion of the tracheal fasciculi will not easily take place upon the earth's surface, on account of its lesser degree of aerial polarity, but at a certain height. If several stalks emerge from the root, the bush or _shrub_ originates, but if only one, the tree. If the subdivision first commence at a certain distance from the earth, then do _Branches_ originate.
1105. The formation of branches demonstrates in part a great store of tracheal fasciculi, and part by an easy differential capacity in the plant. Both come to the same thing.
1106. Plants devoid of branches are similar or analogous to roots.
1107. The branches again ramify from the same cause by which the stem ramifies. A branch or twig must be regarded as an entire fascicle of tracheæ, which forms superiorly a closed vesicle or bud, that raises the bark, bursts it open into scales and then opens itself. The opening of the external tracheal tubes or vesicle becomes a leaf; a twig can therefore occupy no other position than in the sheath or angle of a leaf. Every succeeding leaf upon the twig is in the same manner a ruptured vesicle of tracheæ. As many leaves therefore originate as there are tracheal zones present.
1108. Every branch is an entire plant. All the tissues and systems are found in it. Tracheæ free themselves from the stalk, pass towards the circumference, break through the bark and carry with them the liber, whose external layer again becomes bark. The branch is only an elongated bud. The stalk is the soil or the root of the branches. Branches, that have been cut off and stuck in the earth, grow. There is nothing contradictory, in the tracheal fascicles of the branch growing downwards into the stalk. A branched tree is a complete wood.
1109. From the same cause, or by the influence of water on different situations in the earth, the root has branches. But as this influence is weaker than that of air and light, so the number, density, and length of the roots is less.
1110. The polarization of the tracheal fascicles into branches takes place all around the stalk in one situation. The influence is equal upon all sides. The idea of the formation of branches is the star. All branches have a radiate position around the stalk--all form a verticillum or whorl.
1111. Every other position of branches is only an alteration of the verticillate arrangement.
1112. In most plants the arrangement of the branches admits of being reduced to the spiral line. This position is only the verticil drawn out.
1113. This drawing out takes place through the continued growth of the stalk, in which the bundles of tracheæ, doubtless by the varied operation of light, develop themselves in order, become individualized, and issue from the stalk as branches.
1114. The crucial position of the branches depends upon the same growth of the stalk, but one in which transverse polarities are present. The irregular or dispersed position is most probably the last completion of the spiriform.
1115. The spiriform arrangement stands in relation with the formation of the tracheæ.
1116. The branches of the root observe no such regularity, partly on account of weaker polarity, partly on account of the obstacles placed in their way, at one time by the impenetrable earth, at another by want of water.
1117. The more the stem is differenced, so much the higher is it developed. Thus the richer in number the branches, by so much the more perfect is the stem. The stelliform branches belong to the first development. The plants, that have this arrangement, stand lower in the scale. To this the crucial appears to succeed, as a mediate position between the former and that which follows. The spiriform ranks higher. In it the stem is manifestly differenced more multilaterally. The dispersed arrangement appears to be the highest, because in it the greatest freedom prevails, because the poles have acted on every part of the plant, because they are everywhere in the air and in light. Plants with dispersed branches are organized air; without branches they are organized water as well as earth. Those plants only which have circles of tracheæ or rings of wood, as e. g. the Dicotyledons, ramify; these alone are, properly speaking, a conjunction by growth of many plants, and one that is truly persistent, or that bears fruit several times. The Monocotyledons being devoid of woody rings, do not, or but very rarely, ramify. Most of them therefore die off after they have once produced fruit. The ramification is a multiplication of the plant, in which the buds continue to stand upon the old stem.
_Formation of Nodes._
1118. The formation of nodes, as in the grasses, is an attempt at differentialization, that has not, however, attained to perfection. A node is a branch-whorl which has continued to adhere to the stalk. Therefore the tracheæ also terminate in the circumference of the node.
1119. The formation of nodes consequently ranks directly under the stelliform formation of branches. Taken in a strict sense the formation of nodes occurs only in plants with sheathing leaves or in Monocotyledons.
3. AIR-ORGAN, FOLIAGE.
1120. If in the progressive separation of the tissues the tracheæ finally obtain the preponderance, so that they issue forth free from the envelope of cellular tissue, the _leaves_, or _Foliage_, then originate.
1121. The ribs of the leaf are the fascicles of tracheæ that have become free, and are still only connected together laterally by a thin layer of cellular tissue.
1122. The leaves can be regarded as gigantic and unrolled spiral vessels, and these again as microscopic and involuted leaves.
1123. As through the root the water-process enters the plant, and through the stalk the earth-process, so does the air-process through the leaves.
1124. It is probably the stomata through which the air is conducted into the tracheæ; the connexion, however, has not yet been proved.
_Gemmæ or Buds._
1125. With the formation of branches there is at the same time a diminution of the cellular tissue, and an increase of the tracheæ. Entirely new spiral vessels commence in the branches, and are not continued or prolonged into the stalk. The further indeed the extent of the ramification, by so much less is the quantity of cellular tissue, and by so much greater the number of the tracheæ. It comes at last to this, that the tracheal fascicles, which were from all sides surrounded by dense cellular tissue, are only then loosely connected by a thin layer of such substance. This ramuscule is still therefore only a hollow stalk, consisting of fascicles of tracheæ disposed in circles, and so united by a thin cellular integument or membrane, that the whole forms a vesicle.
1126. This vesicle is a _Bud_. A bud is at bottom none other than the end of a twig that has become hollow.
1127. Several buds are usually involved in each other, i. e. many vesicles of tracheal rings have been encased in each other. They issue gradually forth and become shoots or buds. Buds are bulbs at the end of the branches.
_Leaves._
1128. If the bud or the external vesicle ruptures, while the cellular substance becomes consumed at the apex or between two or more tracheal fascicles, it is then manifested as a _leaf_ or _Leaves_.
1129. Then the second vesicle grows forth, becomes petiolated, ruptures and becomes leaf or leaves. In this manner a twig is formed, surrounded in a spiral manner by leaves.
1130. The younger leaves have been originally inclosed in the older, as in their sheaths.
1131. Every perfect leaf, i. e. every leaf-vesicle, must be regarded as the terminal extremity of an entire twig, from or out of the angle of which a new twig grows forth, that again as a bud ruptures, and from which again a twig grows forth.
1132. All leaves therefore range directly opposite to each other. A branch with many leaves is a system of branches, which grow out of each other, like the articular pieces of the grass-culm or straw.
1133. A leaf is a whole plant with all its tissues and systems; with cells, ducts, tracheæ; bark, liber, wood, stalk and branches. The leaf is a tree of special form, a tree, whose branches or tracheal fascicles all lie in one plane and are held together by parenchyma. It is the bodily expression of the position of the tracheal circle in the stem, only ruptured and to the greatest degree attenuated.
1134. In the division of the ribs of the leaf the internal arrangement of the woody fasciculi in the stem has been placed before our eyes, as by the scalpel of an anatomist.
1135. From the arrangement of the ribs of the leaf the structure of the whole plant can be recognized and its character determined. The leaf is the table of contents or index of the stem.
1136. Plants, which have no tracheæ, have also no leaf-ribs--Mosses.
1137. Plants, which have only isolated or non-ramifying fascicles of tracheæ, have parallel leaf-ribs that do not ramify--Monocotyledons.
1138. Plants, which have a circle of tracheæ, or rings of wood, have leaves with ramified ribs--reticular leaves or true foliage--Dicotyledons.
1139. The stronger indeed the ramification of the leaf-ribs, by so much the higher is the perfection of the leaf. The lowest leaf is that devoid of ribs, the higher that with parallel ribs, the highest being the reticular-veined leaf.
1140. The number and forms of the leaves that proceed from a bud, depend partly upon the number of tracheal fascicles, which pass out of the ramule into the leaf, partly upon the form of the leaf-bud.
1141. If the bud be simply ruptured at its apex or only between two fascicles of tracheæ, there then originates the spathiform leaf.
1142. If the cellular substance between several tracheal fascicles be consumed by the severing action of light and by the air, then the bud divides into several leaves.
1143. The fundamental form of the leaf is the oviform, because the bud is to be thought of as being round. Through the elongation or compression of the bud, lanciform, cordiform leaves, &c. originate. Besides it appears, that the leaf-buds, at least those of the reticular-veined leaves, burst in a circinate manner, like the ferns, and unroll themselves. Therefore the leaves are unilateral and clasp the stem inferiorly; or the petiole displays its spathiform origin. In this case the leaf is not a part of a hollow globe, but the entire globe, that has ruptured in the transverse direction at the extremity of the petiole. Such is the case at least in the demispathiform leaves of the Umbelliferous plants.
1144. The primary position of the leaves like that of the branches is thus verticillate, yet always, however, under the idea of one being encased within the other. The leaves are here only the ultimate branches.
1145. If the leaves upon the ramules issue from each other, then this happens in the same manner as with the branches.
1146. The verticillate position of the leaves is therefore the lowest, next succeeds the crucial, then the spiriform, and lastly the scattered.
1147. The spathose leaves are only to be regarded as a single bud, and therefore differ from the whorl, in so far as we regard this as the development of several leaves, that have not, however, been extruded from each other.
1148. Divided leaves originate through a higher operation of light. In them the formation of ribs preponderates, and therefore they rank higher than the undivided.
1149. From this cause the pinnate leaves are the highest.
1150. From the same cause the radical must be worse developed than the ramular leaves. They are usually non-pinnated, undivided, because they have more cellular substance in their composition than the upper leaves. In the leaf-system, consequently, the whole idea of the plant has been recontained; on the earth resides the chemical character, as is evidenced by cellular, dense and misshapen leaves; above in the air, on the contrary, the leaves are more delicate and are divided--indicating an electric character.
1151. The division and pinnation of leaves can only progress according to the odd numbers, 3, 5, 7, because the midrib determines the odd leaflet.
1152. Leaflets occurring in pairs, or equally pinnated, are arrests of development.
1153. The even number or the symmetrical form is unnatural in the vegetable kingdom.
1154. The leaves are, like the young bark, and thus the whole trunk of the plant, green, because the vegetable kingdom represents the lower totality of the earth, the planet, whose synthesis is the water.
1155. From the same cause, the chief colour of the animal kingdom is red, the colour of fire. Thus, plant is to animal, as green is to red.
1156. The division of leaves passes also parallel to the stages of rank in plants. _Cellular leaves_ are the scales of mosses and ferns; _vascular leaves_, the long riband-like leaves of Monocotyledons; _tracheal_ leaves the reticular leaf of Dicotyledons. The _cortical leaf_ is the sheath; the _liber leaf_ is probably the fat leaf; the _wood leaf_ the acicular leaf. The _radical leaf_ is the undivided reticular leaf; the _stalk-leaf_ the free or ragged reticular leaf; the _perfect leaf_ the pinnate. The _bracteal leaves_ repeat all forms in the thyrsus, since they are floral leaves.
1157. The accessory leaves or _stipules_ are none other than the remnant of the sheath-formation, out of which all the leaves, and therefore the wings of the leaf-petioles or phyllodia, have issued forth.
1158. The thyrsus has also its series of leaves; the scale-like or radical leaf is involucre and bractea; the vascular or spathe-leaf is calyx; the tracheal or reticular leaf is corolla.
1159. The vegetable trunk, namely, root, stalk, and leaf, is a perfect organism, which can exercise all the functions which belong to its individual life. If it therefore produces anything, that can be nothing new, but only itself repeated. This repetition of itself is called propagation. The organs of propagation are thus none other than a repetition of the organs of the vegetable trunk. The plant thereby steps forth out of its individuality into the province of the genus.
B. _Æther Organs._
THYRSUS OR FLOWER.
1160. Hitherto we have regarded the plant as simply a planetary organism, namely, as a trunk with water-, earth-, and air-organ. But the primary vesicle does not lie wholly in the dark, but is illuminated upon its apex by the sun. Every operation, however, produces its like; thus in the plant a light-organ must also be developed. As the light evokes heat in the æther, so also does it evoke a heat-organ in the plant. As the heavenly bodies corevolve in æther by means of light and motion, or the gravitation condenses the æther into matter, so also must an organ of gravity originate also in the plant. These organs are not, however, predominant in the plant, because it is essentially planet or vegetable trunk. They can be therefore nothing else than the parts of the trunk itself with the properties of the æther or fire. They are thus a repetition of the trunk, wherein, in place of the material processes of growth, those of the light, heat, and gravity occur. The light-organ excites the heat-organ by polarization unto motion and thereby originates the organ of gravity.
1161. The process in which through polar tension the trunk has been reproduced, as the Whole in miniature, is called sexual process. The æther-organs are thus sexual organs. These sexual organs can only be a leaf-formation, because the last development of the stem is the leaf. The leaf-formations which, by means of polar tension, reproduce the trunk, are the _Flower_. The light-organ is the _corolla_. The heat-organ is the _pistil_. The organ of gravity the _seed_. The pollen upon the stigma sets the pistil in a state of tension with the trunk, whereby the sap, out of which the seed attains to perfection, ascends. Without this tension the pistil would not have had strength sufficient to perfect the seed. It would wither ere the latter had obtained sufficient nourishment.
1162. The development of the flower takes place through differentialization, individualization, or complete separation of the trunk-organs. The trunk indeed summons up all power in the leaves, to separate the three vegetable tissues and represent each as a particular organ; only in this formation it does not wholly succeed; for in a leaf the ribs or tracheal fascicles are still held together by cellular tissue. In the first place with the perfect separation of the tissues or, properly speaking, with the ex-organization of each of them unto an independent Whole, the limit of the vegetation is attained and the growth completed. This was the course of the whole of nature; in every system she has proceeded to individual consummation of the factors, to their liberation from chaos; and the developments of the systems were concluded, so soon as all factors became independent, or so soon as every factor had itself become an entire nature. Such was the case in the genesis of elements, and such in the metamorphosis of the earth-element into earths, salts, Inflammables, metals.
1163. This complete severance and individualization can no longer be effected by the air, but must be achieved by the light. The air is itself not the wholly differencing element, but derives its potency only from the light. All ultimate separation and individualization is reserved for the light.
1164. Root and stem are the water-and earth-plant, the leaf is the air-plant, the flower is the light-or rather fire-plant.
1165. In the flower the problem has been solved, of producing an entire plant simply by the light without earth, water, and air, or as it were in a merely spiritual manner.
1166. The plant is a flower that has been posited under three ideas, under the idea of earth, water, and air. As in æther or fire all the elements are dissolved; so are all the elements of the plant in the flower.
1167. The flower is truly, not merely in idea, the whole plant with all systems and formations, posited under a _single_ idea, under that of the æther, namely, of the gravity, light and heat, or the fire.
1168. The flower as the æther-organ of the plant is not so independent as an animal, but subordinated to the planetary systems, being only a separation of the parts of the trunk, not a new formation, as in the animal kingdom.
1169. The flower as the highest formative consummation of the plant, or as the highest vegetable organ, is the extremity of the ramules. (Ed. 1st. 1810. § 1176.)
1170. Although the flower is a repetition of the whole plant, namely, of the root, stalk, and leaf; it can still only be a direct transmutation of the leaf. For every Superior proceeds from that which is immediately subjacent. The light-organ can only be developed out of the air-organ, not out of the water-organ. The transition from the water-organ into the light-organ is necessarily indicated through the aerial form. Preparations are necessary, gradual dismemberments must precede, before the isolated consummation can result. The air purifies the organs, in order that they may become participant of light.
1171. The flower is the totality of the leaves of a ramule placed upon the extremity of the latter. For a flower is the whole plant, and is the Ultimate of the plant.
1172. The flower is a terminal leaf-bud. A leaf-bud, after whose rupture the ramule cannot grow any more.
1173. The flower necessarily stands in a whorl-shaped manner, because it is the end of the branch; it is the terminal whorl of the plant.
1174. With the flower the ramule or the plant dies off, partly because it is the extremity, partly because wholly separated tissues cannot live. In the flower therefore the plant reverts to its origin. It is a ramule, whose buds have continued stationary.
1175. The idea of the vesicle lies at the basis of the flower. It is an entire vegetable vesicle, a leaf-bud that has not been drawn asunder. The flower is the last vesicle into which the stalk swells out.
1176. The flower-vesicle agrees with the leaf-vesicle. The form of the flower must pass parallel to the form of the leaves. This has reference principally to the position and number of parts.
_Division._
1177. The flower is the synthesis of the entire plant with complete analysis of the organs. Flower, pistil, and seed are the leaves, stalk, and root separated, and yet all combined to form a common organ. This flower regarded in its analysis is the flower proper; in its dissolution it is called _fruit_.
1178. The flower-vesicle is according to its essence a threefold vesicle. In it the leaf-system or the air-plant has been represented, but in like manner and of necessity also the earth-and water-plant, or the vesicles in which stalk and root have been taken up into the kingdom of light. Thus there is the _leaf-_, _stalk-_, and _root-flower_.
1179. The leaf-flower is in the periphery, the stalk- and root-flower in the centre of the vesicle. For the former is the metatype or copy of the leaves, the latter of the stalk and the root.
1180. The leaf-flower is the highest and the very first to be developed. It is that, which chiefly corresponds to light; the trunk-flower is, however, the lowest, the last developed, because it is only the trunk that has been prolonged with difficulty to form a flower. It is the child of the heat and gravity.
1181. It may be also said that the leaf-flower is the electrical, the stem-flower, however, the chemical. In the latter the chemical process must still act visibly, it must still be produced mucus; in the former, however, this must disappear and resolve itself into purely electrical bodies.
1182. The flower consists of three leaf-buds. The leaf-bud is the corolla or blossom. The stalk-bud the pistil. The root-bud the seed.
1183. The corolla is the external whorl of leaves, is first developed, has the form of a leaf, is a vesicle, secretes in itself electrical, inflammable bodies, and is directed towards the sun.
1184. The difference between corolla and pistil is that of the two principal tissues, the tracheal and cellular tissue. By the light the tracheal fasciculi become finally separated from the cellular substance, evolved to a higher degree as the child or product of light, and planted outwardly. The corolla is the tracheal circle, which has forcibly gained its freedom.
1185. The pistil is the vascular substance that has become freely evolved, yet to the highest stage; in a similar relation likewise does the seed stand to the cellular substance. In the fruit therefore the flower again reverts to the primary condition of the plant.
1186. The flower and pistil are therefore those very organs which have been most antagonized in the plant. They are in the highest state of polar tension, and stand opposite each other like electrism and chemism, or as light and matter. This antagonism in the Organic is called _sex_.
1. FLORAL ENVELOPES.
1187. The blossom is the leaf-formation, in which the separation of the principal tissues is completely attained, where the tracheal fascicles entirely separate from the cellular substance, and become a leaf with free ribs. The cellular substance becomes the _corolla-petal_, the rib the stamen.
1188. The blossom passes through the three stages of leaf-varieties before it attains its completion, and divides therefore into three whorls, which correspond to the root-leaves, stalk-, and ramular or perfect leaves. The radical or squamose leaf appears in its repetition as _involucrum_ or _spatha_. The stalk or vascular leaf as _calyx_. The perfect or tracheal leaf as _corolla_.
a. _Involucrum._
1189. As the root puts forth numerous branches, so the involucrum or spatha frequently incloses many blossoms that constitute the thyrsus or synthetic form of inflorescence.
1190. The thyrsus or dense panicle is the whole ramage or branch-fabric repeated in the involucrum; it is therefore just as manifold as the former.
1191. The inflorescence is still as complex as the arrangement of the branches, because with every flower the ramule dies off or ceases to grow, whereby very numerous and strange relations are brought to light.
1192. The involucrum corresponds to the scale-leaves or bracts, and is therefore as a rule polypetalous. The involucral leaves stand upon a lower stage of development, are for the most part only squamose or spathiform, rarely divided or pinnate.
1193. The involucral leaves necessarily stand, as being the radical leaves of the flower, at the bottom of the peduncles or flower-stalks; each flower-stalk therefore has usually an involucral or bracteal leaf also.
b. _Calyx._
1194. The repetition of the cauline or spathe-leaf in the flower is the _Calyx_. It does not stand therefore at the base of the peduncle, but towards its summit, and is the external bud of the blossom; it is mostly spathiform, rarely or but slightly cloven, and very rarely polysepalous; it is generally green like the leaves. Although the calyx is no essential organ in itself, still it is rarely wanting and is often itself the supporter of the corolla and stamina.
1195. As the calyx is more incomplete than the corolla, so it has usually but three lobes, and if it has five, then they are frequently placed irregularly.
1196. The developmental stages of the calyx are also three. Corresponding to the scale-like leaf or bract, it is only squamiform, as in the catkins of the hazel and fir-cones; corresponding to the spathe-leaf, it is tubular or unisepalous; corresponding to the reticular leaf, it is multisepalous, and mostly deciduous.
1197. The tubular calyx exhibits likewise three stages of development. At first it is scale-like, confluent with the ovary or epigynous--_ovarial corollæ_. It is next spathe-like, simply connected with the corolla or perigynous--_calycine corollæ_. Lastly, it resembles the reticular-veined leaf, being free from both the above organs, or hypogynous--_pedunculate corollæ_.
c. _Corolla._
1198. From the character of the corolla or crown as being a ramular leaf all its properties admit of being deduced. It is the upper leaf-bud, inasmuch as the ramular leaves stand above those of the stalk. It is, on this account too, the internal, inasmuch as the lower leaves admit within them the upper.
1199. The corolla is homologous with the whole mass of the petiolated or ramular leaves. Thus their arrangement, form, and number taken together are not worth more than the characters of the corolla, nor are they worth less, because they are relations of deeper organs.
1200. The corolla stands also in a whorl, because it is the totality of leaves.
1201. The laws of the leaf-formation are also the laws of the corolla-formation. The corolla will therefore represent at one time a greater, at another a less, ramified leaf-system.
1202. The corolla is the last verticil in the series of leaf-whorls; for it represents the last leaf-form, and must fade, because the tissues have completely separated from each other, namely, the tracheæ as stamina, from the cells that constitute corolla leaflets or petals. No part can carry on life by or for itself.
1203. The parts of the corolla range in an alternating manner with those of the calyx, because they are the next supra-consecutive bud.
1204. In accordance with the three stages of the leaf-formation the corolla is also resolvable into three forms. The scale-or root-leaf returns in the several squamiform micropetals of what have been called the Apetalous plants; it corresponds to the involucrum--squamoid corolla. The spathe-or stalk-leaf appears in the tubular-shaped or Monopetalous corolla. It is a leaf-bud, which has only ruptured at the apex, like most calyces, unto which it corresponds--spathoid corolla. Lastly, in the polypetalous corolla the reticular or petiolated leaf appears upon its highest stage--reticular or phylloid corolla. The rank of the leaves consequently determines also that of corollæ.
1205. Although we regard the corolla as a whorl of leaves, namely, as several leaf-buds approximated, a yet clearer insight into its numbers and positional relations is attained, by viewing it only as the lobes of a single leaf.
1206. If we regard the petals of a polypetalous corolla in the import or light of _pinnate leaflets_, the legitimate regularity of their numerical relations, and the so-called irregularity of their arrangement is easily explained. The papilionaceous corollæ have been manifestly constructed according to the plan of the pinnate leaf; the vexillum or vane corresponds to the odd leaflet, the alæ or wings to the two anterior, the carina or keel to the two posterior pinnate leaflets; and hence their consecutive ratio of decrease in respect to size.
1207. All irregular corollæ admit of being referred to the papilionaceous form. One petal is always found, which separates from the others, is either larger or smaller, or entirely ablated, and consequently corresponds to the odd pinnate leaflet or vane.
1208. The irregular monopetalous or tubular corollæ admit also of being referred to the papilionaceous form. They need only be regarded as a confluence of petals. The labiate corollæ are one of the same character; but, on the contrary, the lower lip is trifid, the upper, and thus the keel, bifid. Here the papilionaceous corolla stands, properly speaking, reversed.
NUMERICAL LAW.
1209. The number of petals in the corolla ranges parallel with the mode of development of the pinnate leaves. But at first the odd terminal leaf necessarily originates, because a leaf must have several pairs of ribs, before it splits in a ptiloidal or pinnate manner. The primary number of petals is therefore one. This number is found in those plants called apetalous, where the small lateral petals are abortive, and frequently the terminal petal itself, as in the catkin, where only the calyx has been left remaining.
1210. The normal ratio, in which the numbers of the petals progress, is the odd or uneven. For a leaf is to be regarded as a single fascicle of fibres with cellular substance. This packet of fibres grows straight out. It is solicited by the light to give off bundles of fibres; there is thus no existing cause, why it should only give off such upon one, and not upon the other, side of the bundle. At the first division therefore it must be an _odd_ leaflet, which is the principal bundle of fibres, while two _even_ pinnate leaflets take their origin upon its sides.
1211. The second number of petals is therefore _three_; for this is the first number in which a pinnate leaf can appear. This number is much more frequently met with than the former, because all division of a terminal bud necessarily strives after the representation of the whorl. Under this head come most Monocotyledons, as grasses, rushes, lilies.
1212. Of the three petals of a corolla all three are not of equal import, two only are alike as being lateral pinnate leaflets; the third, however, is present as the odd or uneven leaflet.
1213. Upon this inequality of import depends the irregularity of many ternary corollæ, e. g. of the Orchideæ, Apiaceæ, and even many Irides and lilies.
1214. If the trinity or ternary division depend upon the pinnated leaf, so must also the next number depend thereupon.
1215. The third prevalent number in the vegetable world is the quinary. The quinary division originates, when to the two pinnate leaflets two others are superadded. The pentapetalous corolla is also an _odd_ pinnate leaf in a whorl-shaped position with four pinnate leaflets.
1216. The pentapetalous is naturally higher in rank than the tripetalous, and this than the monopetalous, corolla.
1217. In the former four petals are of equal rank, but the fifth or odd petal differs from them. This difference is shown in its position, size, form, design, and colouring. The odd leaf is usually larger, of a rounded form, having more ribs and other spots or markings.
1218. It appears as if, with this second liberation of fibrous fascicles from the main bundle, the differentialization were closed. For most corollæ are only quinarily divided, or admit at least of being referred back to that number.
1219. It is readily comprehended why there are only three breaks present in the number of petals, that these are denoted by one, three, and five, and that they rarely ascend to seven, nine, and so on. For the posterior pinnate leaflets are wont also to abort or be arrested in the leaves.
1220. There can be no plant with _originally_ two corolla-lobes or petals; for the fibrous bundle does not thus divide itself; one main fasciculus is always left. The cause why the principal bundle does not divide into two equal fasciculi resides in the very nature or essence of the stalk. The idea of the stalk operates throughout the whole plant. The odd or uneven leaflet is only the last expansion of the stalk. The even leaflets are its branches.
1221. For the same reason there can be no _originally_ quaternary corolla.
1222. There can be none that is originally sexanary.
1223. But an originally septenary corolla is conceivable, if forsooth the energy of light is still in a condition to separate two bundles of fibres. How rarely this occurs is well known, and thus it still continues doubtful whether this form is not to be explained by an arrest of development.
1224. Nature can produce no originally octo-, deka-, or dodeka-petalous, &c. corolla.
1225. Corollæ with originally nine, eleven, and more numbers are not impossible. The last seem only to exist.
1226. All corollas having even numbered petals, originate through arrest of the odd leaflet.
1227. The number two originates most usually and in the simplest manner from the arrest of the number three. If it originates from the quinary quantity, two pinnate leaflets are then coarrested.
1228. The aberration is recognized either from the position of the lobes or petals that have been left remaining, or from comparison of the number in other parts, in the calyx and capsule.
1229. The quaternary corolla is a quinary without the uneven leaflet.
1230. The hexapetalous is a doubling of the tripetalous, corolla. There are two whorls of petals, provided the calyx has not become corolla-like in character. Both cases are demonstrable through the alternating arrangement of the parts.
1231. The number eight is a doubled four.
1232. Nine is indeed in most instances a product of three multiplied by three.
1233. The number ten is a doubled five.
1234. In every number of petals the law of unequal development consequently prevails.
1235. The original arrangement of the parts of the corolla is bilateral, and therefore symmetrical. In the papilionaceous corollæ this originally symmetrical arrangement is most perfectly maintained. They repeat the position of their pinnate leaves.
1236. This symmetrical arrangement is shown even in many tubular, as in the labiate and personate, corollæ. The trifid lower lip is the standard and wings; the bifid upper lip, on the contrary, the keel.
1237. The small liguliform petal of the lettuce's corolla is a tubular corolla entirely slit up; it is therefore mostly quinque-dentate.
1238. Corollæ which have only a single petal (the tubular-shaped corollæ should not be styled monopetalous, but those which actually have a single petal to the corolla) are indeed to be regarded for the most part as an odd leaflet; yet still much variety appears to take place in these developmental arrests. Thus here no division of the fibrous bundle was attained, or the lateral leaflets have wholly disappeared.
1239. In many this one leaflet also is arrested, and the corolla is wholly wanting. Such a corolla is to be viewed as a stem with radical, but without ramular leaves.
1240. It is not a matter of indifference whether the single involucre that is left remaining be called calyx or corolla; the distinction between both is philosophically correct, though at the same time also it may be frequently difficult to determine. Colour and relation to the stamina and fruit determine much; but respect must be also paid to the whole idea of the plant, whether it has radical leaves or not, whether the leaf-ribs do or do not ramify. Alternating stamina afford evidence of its being the corolla.
COLORATION.
1241. As the colour ranges parallel with the import or quality of the matter, or since the matter and colour are of one and the same kind, so also must this hold good of the colour of the light-flower.
1242. As the corolla only, and not the calyx, is the proper light-organ, so also will it only obey the light in the coloration.
1243. The corolla can no longer be coloured green, for it is no longer a leaf. Now that which obtains another signification, which passes over into another element, must, with the function, lay aside also the old colour. The corolla is besides the perishing, fading leaf; as this begins to turn yellow or red in the autumn, so does the corolla immediately at its origin. It is a born autumnal leaf.
1244. The whole plant must be regarded as a green synthetic colour; the corolla as the analysis of the Green.
1245. The first division of the Green is Yellow and Blue. These two colours are the first which make their appearance in the corolla.
1246. Yellow is the earth-colour, corresponds to the root, and consequently indicates the lowest colour. Yellow corollæ are less developed than those which are otherwise coloured. The corollæ of spring flowers are therefore yellow; so likewise is the middle of the corolla, as is specially exemplified in the disk of the Syngenesious plants.
1247. Blue is the second colour of corollæ in the scale of dignity, or rank. Blue is displayed on the better developed corolla, and is frequently the colour in the rays of the Syngenesiæ. Blue belongs to the temperate zones.
1248. If Yellow and Blue be the divided Green of the leaves, the complementary colour to that of the corollæ must thus remain in the trunk. The trunks of plants having blue corollæ should therefore be yellow; those with yellow corollæ should furnish blue colouring matters, like the Woad.
1249. Red is the third corolla-colour, the true light-colour, in the which properly all corollæ have been immersed; and if they exhibit any other colours, these should be viewed only as instances of aberration from Red. The Reds are the splendid colours, which develop themselves in the middle of summer; in flaming red mantles are the flowers of the torrid zone veiled.
1250. Finally, the form triumphs over the colour. The light has in Red done everything which it could do for colour, having allured as it were all the colours out of the plant; it now, on the contrary, bestows its attention upon the form and delicacy of the substance. The white colour makes its appearance in antithesis to the red, and is mostly associated with very delicate structure.
1251. The cells of the red corollæ are replete with starch-granules, but those of the white are quite empty. The yellow and blue corollæ range in the middle. Red is a superabundance, White a deficiency, of nutriment. The most noble and beautiful corollæ, as well as the lowest also, may therefore be white. White and red are the general colours for all families of plants, but yellow and blue are special colours. In general the trunk is green, the corolla white, the seed black. The mediate stages are red, yellow, and blue.
STAMEN-FILAMENTS.
1252. At length we come to the last work achieved by the light in the corolla, or to complete separation of the systems or tissues. If ever bundles of fibres may entirely separate from the cellular substance, this is possible only in the corolla, as being the final light-organ. Separation must, however, be attained: for thus far do the claims of light extend. But no development remains stationary before it has corresponded with the operations of the developing agent.
1253. In the corolla, as being the highest kind of leaf, the ribs, as the fibrous fascicles, must finally separate themselves from the leaf-substance as cellular tissue. The corolla is a double organ.
1254. In conformity with the whole structure of the plant, the ribs are placed internally, the membranes externally.
1255. The leaf-ribs, isolated and perfected as a particular organ, are the _Stamen-filaments_.
1256. The leaf-membranes, or probably the phyllodia, isolated and evolved into a particular organ, are the corolla-petals. These compose in the strongest sense the corolla. The filaments consist for the greatest part of spiral fibres, and the corolla-petals of the finest cellular tissue, which may be almost designated as granular. This then would be the rational import of the corolla and its stamen-filaments. Both are of similar production; they exhibit like substance, colour, and delicacy, with cotemporaneous development and cotemporaneous death.
1257. Not only are the ribs of the corolla, but those also of the calyx, liberated to become filaments. There are calyx-and corolla-filaments.
1258. As ribs, the filaments must stand in the middle of their petals, i. e. opposite to them.
1259. Filaments, which alternate with the parts of the corolla, are consequently calyx-filaments; such as alternate with the lobes of the calyx, or stand opposite to the petals of the corolla, are corolla-filaments.
1260. Most filaments, and consequently the calycine, are alternating in their arrangement. Most corollæ therefore have no longer strength sufficient to produce filaments.
1261. Flowers provided with opposite and alternating filaments have consequently two circles of them, as is the case in many Pinks. With the determination of the number of filaments the race has been therefore specified.
1262. The number of filaments stands in relation to the parts of the corolla; therefore three and five are the prevailing numbers.
1263. There is no absolute number in the filaments, but only one of relation. Corollæ with three petals have invariably also three filaments, and those with five of the former, the same number too of the latter.
1264. The number of filaments is always the simple or multiple of the parts of the corolla. Three calyx-or corolla-parts have 3 × 1 or 3 × n, filaments. 6 is not 6, but 3 × 2; 9 is 3 × 3; 10 is 5 × 2; 20 is 5 × 4; or 5 × 3 + 5 × 1, and so on.
1265. The filaments do not simply follow the number, but also the conjunction, position and arrest, of the corolla. They are epigynous, peri-or hypogynous.
1266. In irregular corollæ the filaments are usually abortive; as in those of the orchideous, labiate and papilionaceous plants.
1267. The arrest of the filaments usually stands in inverse relation with the corolla. In the case of a larger-sized petal the filament is small, and, on the contrary, larger in opposite parts of the corolla.
ANTHERS.
1268. The corolla obtains its last function in the production of the highest electrical bodies, which it exhales as sweet odours. Ætherial oils ascend out of the corolla into the air.
1269. The filament, as a leaf-rib that has become free, is a moribund ramuscular extremity, which still strives, according to the law of pinnation, to produce three buds, whereof, however, the terminal one is in general arrested, and the two lateral scarce attain unto apertion.
1270. The two lateral buds of the filaments are the _Anthers_. They mostly open in a spathose manner, because they have not strength enough to develop themselves as perfect gemmæ or buds.
1271. The anthers are to be regarded as follicles, which mostly rupture upon the dorsal or external aspect.
1272. The starch-flour, which forms the precipitate termed albumen in the seed, here obtains in the light-organ electrical properties, and is called _pollen_.
1273. The pollen has a light-function in the plant.
1274. The function of the pollen must be differencing, thus vivifying and secernent.
1275. The principal antagonism of the pollen is with the pistil, upon which it must therefore act in a properly differencing manner. The pollen does not hang, like the seed, by a stalk or pedicle to the wall of the anther, but is exudated from it like chemical bodies. It is nevertheless a vesicle, like all organic parts. This vesicle consists of two membranes, and contains yet smaller vesicles, which are called _fovilla_ or pollen-viscus. When the pollen comes into contact with the moist surface of the stigma, its external membrane ruptures, and the internal with its contained fovilla protruding in the form of a tube, penetrates the style in many instances down to the seed, whereby the germ first becomes developed or self-subsistent.
2. PISTIL OR OVARY.
1276. If the corolla be the light-flower, the stalk-flower is heat-flower.
1277. The stalk-flower, as being a repetition of the stem, must be developed later than the leaf-flower. It consequently stands superiorly upon, and so far within, this. The corolla is related to the stalk-flower like circumference to centre.
1278. The stalk repeated in the flower is the _Ovarium_, _germen_ or _pistillum_. It is frequently converted into wood in the nut and becomes hardened into stone.
1279. Nevertheless, the pistil like the corolla is a leaf-formation, because everything that originates subsequently to, can be none other than, the leaf. It is a leaf-bud under the idea of the stalk. The pistil is thus a leaf-whorl like the corolla, and one which is subject to the same fatalities, only with this difference, that its leaves are wont to open first after it has withered, and consequently through physical forces.
1280. Every leaf that has been closed in a vesicular or tubular form is a _follicle_ or _carpel_. There are therefore uni-, bi-, and tri-locular ovaria, &c. The loculi or cells of the ovarium are none other than closed carpels. As many therefore as there are of the former, so many are there of the latter, and vice versâ. The septa or partition-walls are none other than the involuted edges of the closed and confluent carpels.
1281. Uni-locular ovaria consist therefore only of one leaf. The legume is only a compressed carpel.
1282. Every carpel or every cell has its raphé or suture directed inwards or along the axis of the flower. For the leaves are always so conjoined that the two halves of the upper or inner side stand counter to each other.
1283. All other sutures are adventitious and, by their mode of dehiscence, determine those parts of the ovarium which are called valves. These sutures are either _dorsal_ or on the back of the carpel, e. g. _capsula loculicida_; _conjunctive_, where two carpels abut against each other, e. g. _capsula septicida_; or finally, between both by the side of the dorsal suture, so that the valve springing up resembles a shutter, as in many siliquæ or pods.
1284. The columella of the ovary is none other than the internal edge of the carpels from which the leaf-wall has been freed.
1285. Each carpel-leaf is to be regarded as the common petiole of a pinnate leaf, upon whose lateral petioles the seeds depend. The seeds always hang therefore upon the inner angle of the cells.
1286. As the parts of the corolla alternate with the calyx, so do the carpels or cells of the ovary with the corolla; they stand therefore opposite the parts of the calyx or are situated in front of them.
1287. The parts of the ovary follow also the uneven series of numbers, one, three, five. The number two is usually found in irregular, e. g. the labiate, corollæ.
1288. If few cells be present as parts of the flower, the carpels are then to be regarded as arrested. In the personate corollæ three are arrested, but in the papilionaceous, four. The legumen is only a fifth part of the ovarium.
1289. The development of the carpels stands usually in an inverse relation to the size of the parts of the corolla. Thus the legume is situated between the two insignificant petals of the keel, opposite to the large vexillum; in the Personatæ a carpel is situated in the fissure of the upper lip; upon the lower lip, consisting of three lobes, only one carpel is situated, which consequently supplies the place of four, and is therefore also larger.
1290. The stages of leaf-formation are also displayed in the matured state of the ovarium. The _radical_ or _scale-leaf_ is repeated in the _cariopsis_; as in the grasses, orachs, nettles, and such like plants.
1291. The _spathe-leaf_ becomes a _siliqua_, or hollow capsule, in which forsooth the septa are arrested, and the seeds stand upon the conjoined edges of the carpels, upon their walls, or also upon a mediate replum and placenta, as in the proper Siliquose plants with the poppies, Resedæ, Primulæ and pinks.
1292. The reticular-leaf is perfected into a _capsule_, where the carpels are so confluent with each other, that they form septa and bear the seeds upon the internal angle or upon the axis, as in the Rues. If these carpels separate, then polycarpal plants, as the Ranunculaceæ, Malvaceæ, Magnoliaceæ, originate. In these the median columella is the elongated floral peduncle. If they separate, without leaving a median columella between them; they are then simply called _follicles_, as in the larkspur and celandine. If this capsule be compressed flat; it is called _legumen_, as in the beans.
1293. The cariopsis generally contains one and that a large seed; the siliqua or hollow capsule is many-or small-seeded; the follicle few-or moderately-seeded; the capsule many-and also few-seeded.
1294. In the cariopsis the seed is attached to the base or apex; in the follicle in a row upon the inner suture; in the siliqua upon the carpellar edges, or their walls, and on a meso-replum or frame; in the capsule upon the inner angle or on a middle axis or column. Seed-bearing alæ to the columella are only the carpellar edges prolonged into the loculi or cells.
STYLE.
1295. What the stamen-filament is for the petal of the corolla, that is the style for the ovarial petal or the carpel, viz. the rib that has become free. As, however, the leaf-formation in the ovary is generally imperfect, so also is the separation of the tissues or systems. The style is not therefore freed at once from its root, but projects only above the leaf-substance.
1296. But as the singular circumstance occurs in the ovary that the midrib is arrested while only the marginal ribs shoot out, so is the style the elongation and coalescence of the two marginal ribs. Every stigma is therefore biacuminate.
1297. There must always be as many styles as the ovary has carpels or cells. If only one style appear, in this case it is then made up of several mid-ribs. In most instances the number of styles is recognized in the number of the stigmata.
1298. As being the rib of the ovary the style is the last ramuscular extremity of the stalk, which is resolved into mucus upon the stigma.
1299. Stamen-filament is related to style, as leaf to stalk, thus as air to earth, as Differencing to Differencizable, as electrism to chemism or rather nutrition. This is the lower comparison; in a true sense they are related as light to heat.
1300. The light is the Active, the heat the Passive; light the Moving, heat the Moveable; light the Vitalizing, heat the Inactive, or that which becomes vitalized; light the spirit, heat the matter--male and female principle. Thus are corolla and pistil related to each other.
3. SEED.
1301. The root repeats itself in the interior of the ovary under the æther-form. The root ascends out of the earth to become an organ of gravity.
1302. After the leaves have been made independent in the corolla, and the stalk in the pistil, the root also separates and appears as a free organ, as _Seed_.
1303. The seeds are necessarily in the interior of the ovary; for the cellular organ can first appear, after the leaf-and stalk-buds have opened as corolla and ovarium. The blossom is a bulb, the external testa or covering of which is the leaf-, the mediate the stalk-, and lastly, the internal the root-vesicle. The stalk is adherent to the leaves, the root to the stalk; so also the seeds to the ovary, and this to the corolla.
1304. The seeds are developed in the ovary under the same relations in which the root is developed in the earth, namely, in the dark.
1305. The darkness does not allow the chemical body to attain unto difference; therefore the sap within the capsule must, instead of separating itself into spiral vessels and leaf-substance, continue to remain there undivided and without form, i. e. in the condition of simple granules, or germs of future cells.
1306. The seeds, like the root, are a mass of cells; like it they contain an accumulation of mucus, but of course more highly-formed, being separated into flour, starch, acid matter, oil, and such like substances.
1307. These seminal substances are deposited upon the alkaline, in opposition to the acid, side of the ovary; just as the root also represents the alkaline factor in reference to the stalk, in which appears the formation of acids.
1308. The seeds are the pinnate leaflets of the ovarian leaves, which continue in the condition of buds. They stand therefore as unclosed vesicles upon both edges of the carpel, as is particularly distinct in the legumens.
1309. As both edges are similar to each other, so there can be no ovary that has fewer than two seeds. In all one-seeded ovaries therefore one seed has been arrested, a fact that admits too of being demonstrated in the majority of cases.
1310. Every seed is placed at the extremity of a lateral rib of the carpel. These lateral ribs are called seed-bearers or _placentæ_. If such lateral ribs terminate before reaching the edge of the carpel, then the seeds stand upon the juncture or wall of the carpel. This does not, however, occur frequently, but only in the Siliquosæ, poppies, and some others. The elongated lateral rib, whereupon the seed hangs, is called umbilical cord. It is no peculiar organ, but only the seed-petiole.
1311. The direction of the seeds is possible in five ways, either upright and horizontal, transverse, or rising obliquely upwards and downwards in relation to the axis of the ovary.
1312. Every perfect seed (of Dicotyledons) is none other than a pentifoliar, involuted, pinnate leaf. The shell of the seed is the leaf-spathe or _phyllodium_, the two seed-lobes are the two posterior pinnate leaflets, while the germinal leaflets, or _plumula_, are the two anterior pinnate leaflets, together with the odd leaflet. The seed-rib or vascular cord (_raphe_) is continued into the seed-rootlet or radicle, and this into the petiole of the cotyledons.
1313. Every seed-coat must consist of three integuments; for every leaf consists of the lower and upper membrane, and of the interjacent parenchyma, in which the vessels are dispersed. The external leaf-membrane forms the most hard and coloured covering of the seed (_testa_), the internal the brown seed-tunic or _pellicula_; between the two lies the brown fibrous tissue, or desiccated parenchyma with the vessels.
1314. The hilum is the basis of the bud or of the seed-leaf; the seed-hole or micropyle is on the apex of the involuted bud, or rather of the phyllodium, in which the germ lies rolled up.
1315. Umbilicus and micropyle are united with each other by means of seed or leaf-rib (raphe). Both rarely stand opposite to each other, so as that the one should be below, the other above; but the apex of the leaf is usually so involuted that it again reaches the bottom of the leaf, whereby umbilicus and micropyle come into close and mutual approximation, as in the beans. The seed-petiole elongates itself into the seed-rib; this is continued upon the back of the phyllodium or testa, bends round, and returns again to the umbilicus, so as to describe a complete circle. The shell of the seed has consequently the form of the young fern or fern-capsule.
1316. The radicle is the continuation of the seed-rib, which is, however, dismembered itself, moves off, and thereby causes the micropyle to be, or, properly speaking, only renders it, free. The testa or seed-shell is consequently a phyllodium thrown over the germ, but the micropyle is the upper opening of the bud.
1317. The germ of the seed or embryo, namely, radicle, cotyledons and plumule, is therefore only the quinary pinnate leaf without the spathe or testa.
1318. Thus the whole seed, shell and embryo, completely resembles a pinnate leaf with a phyllodium, as we see it in the umbellate plants, only it must be thought of as one so involuted, that the fine leaves adhere reversed in the phyllodium. Seeds may consequently change into leaves. A seed has therefore been formed also in all its parts like a papilionaceous corolla. This resemblance speaks moreover retrogressively, for the petals of the papilionaceous corolla being viewed only as a single leaf-bud. Seeds may therefore change also into corollæ. All parts of the seed are thus an unity, a single pinnate leaf, and it is consequently impossible for them to have been patched up out of what has been called the seed-ovum, namely, the testa and embryo, which would come from some other source or out of the pollen.
1319. The seed is the whole plant in miniature: the root being portrayed in the umbilical cord, and radical leaf in the phyllodium; stalk in the radicle; caudal leaves in the seed-lobes; ramuscule in the cotyledonal petiole; ramular leaves in the cotyledons. Seeds may thus change into an entire plant. The seed is consequently nothing new in the plant, but the repetition of the same under the relations and forms of the root.
1320. It is plain that the seeds must always change into the same plant; for they are indeed nothing else but this. The identity ensuing upon propagation is accordingly nothing singular and incomprehensible; it would be so were it otherwise. With the seed the plant has but reverted again to its primary condition, to the galvanic, mucous vesicle, out of which in a secondary manner the young plant is developed like as is the first plant out of the primary vesicle.
1321. The radicle is not therefore root itself, but only emits rootlets.
1322. The germ or the radicle must observe different positions towards the umbilicus, according as the seed-leaf or the testa has been more or less involuted, and according as the germ is removed from the micropyle.
1323. The albumen or perisperm is no particular organ, but only the deposit from the sap, which the inner wall of the testa secretes. The albumen stands in no organic connexion with the parts of the seed. That therefore which has become connate with the kernel or nucleus cannot be albumen.
1324. The arillus can be none other than bud-seeds of the testa, because it is placed under the phyllodium. It corresponds to the floral involucrum or bracteal scales.
1325. As the seeds are none other than leaves that have remained stationary in the condition of root, so must they pass through the three leaf-stages. There can be therefore only three principal differences in the seed-formation.
1326. The seeds of plants with reticular leaves consist of several leaves arranged symmetrically or in pairs. They have necessarily _two_ seed-lobes--Dicotyledones.
1327. The seeds of plants with spathose leaves consist too only of the latter, i. e. the seed-leaves remain encased in each other. They have consequently only one seed-lobe, which also incloses only one plumule--Monocotyledones.
1328. This seed-lobe is a phylloidal leaf, whose parenchyma has been superabundantly filled with farinaceous matter or flour.
1329. That which is named vitellus can be none other than the succeeding counter-leaf.
1330. What in the Monocotyledons, at least in most of them, and in the grasses, is called albumen, is not so, but only the flour of the seed-lobe.
1331. The germination of these seeds is nothing else than an elongating of the spathiform seed-lobe into a culm, from the bottom of which radicles spring forth, as out of a bulb. A monocotyledonous seed is in its structure none other than a small bulb with undivided coverings.
1332. Lastly, the third form of seeds makes its appearance in those plants which have only scale-like leaves. The seed-lobe is wanting in them, and they elongate themselves directly into the plumule or little stalk--Acotyledones.
1333. They are devoid of the distinction of testa and embryo, because, on account of the deficiency of genuine leaf-formation, they are none other than the first. For the embryo is only a small leaf. They therefore include only albumen or _germinal powder_, as it is correctly named.
1334. Here belong not merely fungi, fuci, lichens and mosses, but also the ferns, as being those which have only cells or squamoid leaves. For the frond with the spiral vessels does not rank in the category of leaf, but of stalk.
FRUIT.
1335. The fruit is the coalescence or blending of the three parts of the flower, or the seed, ovarium and corolla. In the flower the individual perfection of every part of the trunk was completely attained; the leaves being separated quite freely from the stalk became corolla; the stalk separated from the leaves and root became pistil; lastly, the root separated from all, became the seed. In this manner indeed each organ attained its ratio of perfection; the perfection, however, of the Whole does not alone consist in the perfection of the several parts for themselves, but in the union or combination of these individual perfections. The vegetable trunk, as being a Partial, has been represented in the parts of the flower, but as a Whole, in the fruit.
1336. The fruit is therefore the last and most perfect consummation of the plant, or the whole vegetable trunk repeated as unity.
1337. In the fruit not merely the sum of all vegetable forms is united, but also that of all vegetable matters. It is the whole vegetable body repeated mathematically, physically and chemically.
1338. The fruit therefore is also that part of the plant in which all vegetable bodies have been concentrated into flesh. Now, as the highest vegetable bodies pass over into the next, and consequently into the animal kingdom, and are therefore palatable as food, so is the fruit also in essence the sarcocarp. For this is the directly edible part of the plant, that e. g. which does not need being cooked.
1339. The nutritious substance of the fruit can be none other than a highly elevated and analysed kind of mucus, such as starch and gum, sugar and acids. Flour is that which resembles the root, sugar the stalk, acids the foliage; therefore flour is in the seed, sugar in the pistil, and acid in the calyx.
1340. There can be only three kinds of fruit, that constantly accord with the preponderance of the three parts of the flower, as seed-fruit, pistil-and corolla-fruit.
1341. The fruit that has the preponderance of the seed, or where the edible substance resides in the seed, and the ovary itself has become seed-shaped, is the _Nut_. The nut is the cariopsis that has become sarcocarp; it is therefore one-seeded--farinaceous sarcocarp.
1342. In the ovarian fruit the ovarium has become demi nut-like, half corolla-like or fleshy, as in the _Plum_. It is the carpel that has become sarcocarp--acid sarcocarp.
1343. The fruit, in which the whole ovarium together with the calyx is edible, is the corolla-fruit, the _Berry_. Only those perfectly soft fruits are true berries which are inclosed by the calyx, as being forsooth a part of the blossom. The berry is the siliqua or hollow capsule that has become sarcocarp. Therefore it has numerous and small seeds--saccharine sarcocarp.
1344. Finally, these fruits combine to form a common fruit, which represents the proper synthesis of all parts of the blossom, or in which seed, ovarium and flower, along with the calyx, have become sarcocarp. This is the _Apple_, a syncarpus. The apple is the calyx become sarcocarp, and as it usually incloses several carpels, it is therefore polycarpal and contains few seeds. It consists of seed, ovarium and calyx, which have become flesh. The apple, as being an unopen calyx-fruit, may probably be regarded as the fruit of the trunk. It furnishes properly drink and food, is the fruit against thirst and hunger--the universal, alimentary sarcocarp. The apple contains all the bodies that have been named, viz. flour, acids and sugar. It is thus chemically also the synthetic fruit, which may be converted into the whole animal flesh, and thus become a true medium of nutrition. The nut is only garbage, the plum and berry, cherry and grape, only drinks or delicacies.
1345. Other vegetable substances, which range lower in chemical development, as mucus, bitter and colouring matters, with resins, are associated for the greatest part in root, stalk and leaves.
FRUIT OF THE FLOWERLESS PLANTS.
1346. The flowerless or asexual plants can have no genuine seed or no embryo. For the genuine seed is the repetition of the blossom under the idea of root. (Ed. 1st, 1810. § 1564.)
1347. That which has been called germinal powder is no seed or germ, but only albumen or perisperm. It has no seed-petiole, has only been exuded out of what has been called the wall of the capsule, and exhibits in its composition no seed-lobes. (Ed. 1st. § 1586.)
1348. What is termed capsule in the Acotyledones is none other than the seed-shell, whereupon it follows of itself, that the so-called seeds can have no umbilical cords or seed-petioles. (Ed. 1st; 1810. § 1573.)
1349. The capsules of ferns are involuted like most dicotyledonous seeds. The ring corresponds to the seed-rib or raphe, the fissure to the seed-aperture or micropyle. The involuted fern-capsule is a repetition of the involuted fern-frond. The little heaps of capsules or sori are consequently not pollen, but a nest of seeds surrounded by an indusium or veil, which, probably, corresponds to the ovary.
1350. The capsule of mosses is an antetype of monocotyledonous seeds; it is a spathe-leaf with the lateral suture; it springs up in a tubular manner similar to grass-leaves, that free themselves from the nodes of the culm.
1351. The hollow columella, which likewise contains germinal powder, is an internal spathe, which corresponds to the germinal leaf of grasses.
1352. The oral teeth are the dissevered parallel strips of vessels in the culm and leaf of the Monocotyledons.
1353. The urn-supporting pedicel is the seed-petiole or umbilical cord.
1354. The calyptra probably corresponds to the arillus and thus to the bud-scales; or possibly to the indusium of the ferns, and thus to the ovary.
1355. The leaf-roses would consequently be the involucral leaves of the moss-stalk; the moss-stalk itself a peduncle or flower-stalk; so that in the upper involucral leaves rudiments might indeed appear of stamina.
1356. In the lichens and fuci the whole trunk is none other than seed-shell.
1357. In the fungi, the antetype of the Acotyledons, it may be almost said that the whole stem is nothing else but albumen, the external layers of which only cling together in a membranaceous manner, and represent a kind of seed-shell. The fungus is an albumen-body, which has coagulated out of vegetable juices. In the fungus, seed, seed-vessel, ovary, blossom, foliage and trunk have become blended into one.
1358. In a perfect blossom the albumen is therefore the repetition of a fungus; the acotyledonous seed that of lichen; the monocotyledonous seed-vessel that of a moss; the dicotyledonous, however, is the repeated fern. It may be also said that the albumen were fungus; the germ, lichen; the seed-vessel, moss; the ovary perhaps, a fern, namely, its indusium.
II.--_PHYTO-PHYSIOLOGY._
1359. The life of the plant consists in the co-operation of its functions. The representation of these functions is the vegetable physiology or the theory of vegetation.
1360. _Vegetation depends first of all upon the two principal antagonisms of the plant, or those between the tracheal and cellular systems, or between the stem-and rootsystems, sun and planet, air and water with earth, light and matter, electrism and chemism._
1361. The functions divide into those of the _æther-organs_--_blossom_, and the planetary organs--_stem_.
I.--FUNCTIONS OF THE TRUNK.
1362. The functions of the trunk are those of the tissues, systems and members; thus first of the cells, vessels or ducts and tracheæ; secondly, of the bark, liber and wood; lastly, of the root, stalk and foliage.
1. _Facts._
1363. The phenomena to be regarded in plants bear relation to their constituent parts, and the changes or preliminary incidents which they undergo.
A. _Constituent Parts._
The chemical constituent parts of vegetables are inorganic and organic.
a. INORGANIC BODIES.
_Elements._
1364. The plant contains all the _primitive bodies_; carbon, oxygen, hydrogen and nitrogen. The carbon forms the principal mass, and almost alone constitutes its solid parts. The nitrogen is only present in small quantity, being, as it were, a trace only of the future animal kingdom.
1365. In the plant all the _elements_ are also active, as the æther, which, through the gravitation of the root, strives towards the middle point or centre of the earth. The light, which imparts the general polarity and decomposition, as well as produces colours. The latter appear to reside in the starch-flour. The heat, that sustains the indifference, promotes the evaporation and the course of the sap, as also protects the plant from being killed by frost or cold. The air that penetrates through the spiral vessels to all parts, and is also met with occasionally in the hollow stalks, the interstices and cells of the pith and cuticle. It imparts the process of oxydation. The water is the proper mother of the plant, being the medium by which the nutrition is imparted. It contains in a state of absorption some hydrogen and nitrogen, a larger amount of oxygen, and abundance of carbonic acid; besides different salts, mucus, sugar, and acids. The earth, as element, bestows upon the plant a firm station, so that the water-and air-organs continue separate from each other.
_Minerals or Earths._
1367. The plant is also a totality in reference to the _earths_. It contains all the mineral classes, and from each of these indeed the principal or fundamental minerals. It can therefore only thrive in a soil which represents the whole mineral kingdom. Among the earths the siliceous earth is very frequently found in plants, and especially in the graminaceous kinds. This, having been dissolved in the earth by potash and the rich supply even of carbonic acid, appears to be absorbed or imbibed by the plant. The argillaceous earth is scarcely met with in the plant itself; but from imbibing and storing up water for the consumption of the plant, it is without doubt its best and most necessary soil. The talcose earth is rarely found contained in plants; it, however, keeps the soil slacker, by dividing into laminæ, and being present for the most part as mica in sand. The calcareous earth is a more essential constituent part of plants, and is found therein in tolerable quantity, usually combined with phosphoric or carbonic acid. Of the salts, all plants contain a fair proportion of common salt and potash, combined too with carbonic acid; soda with saccharic or oxalic acid; probably also ammonia. Of the acids, carbonic acid appears to be alone contained in a free state in vegetable sap; the other elemental and mineral acids are united to alkalies, talcose and calcareous earths. As regards the Inflammables, almost the whole plant consists of carbon, but contains also some sulphur. The metals are represented by iron, which occurs in all plants.
b. ORGANIC VEGETABLE BODIES.
1368. These must be regarded as the repetition of the inorganic bodies. The alcohol, which does not indeed occur ready formed in the plant, but is developed out of the sugar, certainly corresponds to the æther. The ætherial or volatile oils, and the balsams and resins that are thence formed, correspond with the air. The mucus, gelatine, albumen, and sugar correspond to water; the wood, gum, starch and vegetable mould, to the earth. Of the organic salts, plants contain tannin, with azetic, benzoic, mucic, gelatic, saccharic, tartaric, citric, malic, oxalic, tannic, oleic, isatic, and hydrocyanic acids. The alkaline bodies are pungent, bitter, stupefying, and saponaceous; the fixed or greasy oils, the wax and the vegetable butters, are to be regarded as organic Inflammables; the colouring matters, as the organic ores.
1369. These bodies intermixed form the compound vegetable matters. What has been called vegetable sap is for plants, what the blood is for animals. It consists for the greatest part of water and mucus, starch, sugar, acids, and salts. It passes over into vinous and then into acetous fermentation. The starch-granules appear to form in the cells.
1370. To the secreted saps belong the coloured milky juices present in particular vessels, and consisting for the most part of water with resins, as in the celandine and spurge. The particular saps, especially those of the fruits, are very composite, consisting for the most part either of mucus, sugar and acids, or occasionally of gelatine and albumen. Solid compound matters are almost universally made up of flour, which consists principally of starch and gum; or they are furthermore mucus in the roots and seeds. The excreted or separated matters, which no longer interpose in the vegetable process, are the etherial oils, resins, fixed oils, colouring matters, poisonous substances, gum, tannin, nectar-juices, and even water.
B. _Preliminary Events._
a. WROUGHT BY EXTERNAL INFLUENCE.
1371. The influence of the elements produces different phenomena in the plant. I have felt constantly more inclined to consider, that not merely the descent of the root, but even the ascent of the stalk was simply to be viewed as a mechanical event, or one forsooth effected by gravity. The roots obey under all circumstances the gravity and would grow as far as the centre of the earth, were they to meet with no impediment; and there they would follow the revolution of the earth, and consequently become spirally convoluted upon themselves. It is almost beyond doubt that the water, which sinks downwards and, as it were like that in stalactites, invariably rigidifies or hardens at the radical capillaries, is heavier in the root. The cause of this greater weight depends upon the mucus not being decomposed.
1372. The straight ascent of the stalk also depends upon nothing else than gravity. The upper drops of mucus become lighter by means of greater heat and by decomposition in light and air, and they are therefore compressed by the heavier in the upward direction. It is always such a small drop upon the summit, which hardens into its uppermost cell. The stalk, therefore, grows upwards through the same forces and in the same manner, as the air-bubbles ascend in a glass of beer. The cause of their becoming lighter resides certainly in the vital process, which nevertheless effects in this respect nothing else than the extension or increase; but yet the cause of the ascent is naught else than the gravity.
1373. The _light_ likewise acts upon the direction of plants and especially that of their leaves; not simply from its promoting growth by elevation of temperature and by decomposition, but obviously in a mechanical manner also; for not only do the branches of plants in a green-house grow towards the window, but most leaves turn themselves the whole day in obedience to the course of the sun. This turning must, nevertheless, have one and the same kind of cause with the growing towards light; it also is only a conatus or effort unto growth. The upper leaf-cells, being illuminated by the sun, become lighter, and are therefore directed at once, like the apices of the branches, toward the influence of light. The cells that stand perpendicularly upon the surface of the leaf are to be regarded as branches conjoined by growth.
1374. The _sleep of plants_ depends also upon the same influence of the light. The upper leaf-cells sleep during the night, while the lower cells, especially those of the petiole, fill and consequently bend the latter upwards. The sleep of the flower must have the same cause. As likewise the alternating motion of many leaves, as in the Mimosæ.
1375. The motion also performed by the staminal filaments towards the pistil must finally depend upon this unequal replenishment of the external and internal cells.
1376. The coloration of the parts of plants is a result of the decomposition of the starch-granules in the cells by the agency of light.
1377. The operation of the _heat_ is more intelligible than any other. That which is to move and separate itself, must have a certain degree of extension, or must be fluid, namely, aquiform. In a cold temperature the upper saps, not becoming warmer than the lower, are consequently not lighter, and on that account also do not ascend upwards. The mortal freezing of trees descends from above downwards. In other respects plants have, like animals, a self-inherent, though very feeble, process of heat. Germination proves this, in cases where many seeds lie upon each other.
1378. The air acts also mechanically and physically upon plants, by causing motion of the solid parts and by promoting evaporation. Electricity is without doubt active in the spring of the year, and evokes the antagonism between the fabric of the stem and root.
1379. The physical operation of water consists indeed, for the greatest part, in its preserving the solid parts in a supple or pliant state. Its principal office is, however, to convey nourishment to the plant.
1380. The earths act beneficially only upon plants, if they have been all mixed with each other. Mineral salts occurring in moderate quantity in the soil promote growth; alkalies and acids are injurious thereunto. The same holds good of Inflammables and metallic limes.
b. BY INTERNAL ACTIVITY.
1381. That the plant imbibes water, and this indeed in great quantity, by its whole surface is a well-ascertained fact; but it has been by no means equally determined whether it obtains its nourishment simply through the water, or directly also from the air, e. g. the carbon, as well as the nitrogen, from the carbonic acid. The principal imbibition, however, takes place through the root; but experiments that have been made upon this subject leave it doubtful, whether in this case it is simply mucus, extract from the humus or vegetable mould, or simply carbonic acid that has been absorbed.
1382. It is moreover a fact that the green parts of plants exposed to direct sun-light consume or take in carbonic acid, and develop or give out oxygen; on the contrary, during the night, and even in cloudy or gloomy days, they absorb oxygen and exhale or develop carbonic acid. Now, as there are far more gloomy or at least cloudy than clear days, it thus becomes evident that far more oxygen has been taken up, than separated, from the air. During germination oxygen gas is consumed and, on the other hand, carbonic acid developed.
1383. The saps ascend upwards, and chiefly indeed, in the liber; on its passage different substances forming from it, which appear especially in the fruit in greatest proportion and variety.
2. _Processes._
1384. The tissues of plants form three formations, which must be similar in their functions, and can only exhibit subordinate differences. The cell-formation is displayed in the cellular tissue, in the bark and root. The vascular formation in the vascular tissue, in the liber and stalk. The tracheal formation in the tracheal tissue, the wood and the leaves. There can accordingly be only three principal functions in the vegetable stem, and of these each will display minor differences.
A. _Cellular Processes._
a. ROOT-PROCESS--ABSORPTION.
1385. As the root is the cellular organ proper, so in it principally resides the water-process or the commencement of chemical elaboration and analysis. Now the chemism in an organic body is called digestion.
1386. The root is the mouth or pharynx of the plant, and is therefore principally concerned with absorption. Its process is therefore the formation of mucus, or as it were of salivation. The root cannot, however, create mucus, as it was created at the conclusion of the earth-metamorphosis in the sea; it can absorb it or in the highest degree compound it out of the constituent parts.
1387. The process of the formation of mucus is a process of putrefaction; the function of the root consists accordingly in supporting a constant process of putrefaction. The soil in which the root stands must contain substances susceptible of, and the conditions necessary to, putrefaction. These substances are organic matters and water; the conditions heat and access of air. Such a soil is called humus or mould. In a pure, dry earth, no root can thrive.
1388. Carbon, from its being the earthy body, is the principal one in the formation of mucus, and the basis also of the vegetable bodies. A root can develop itself, if it stand only in a soil such as the calcareous, which contains carbon and water. The calcareous soil is as it were an original mould. It is probable that the calcareous earth is constantly decomposed by the root and its carbon absorbed. The calcareous earth is again neutralized by the carbonic acid of the water and air.
1389. There can be no doubt that the root also abstracts carbon from these elements, and converts it into mucus, or probably separates it from carbonic acid. The mucus approximates the animal nature, so that the root in its constituent parts, in its smell, and even in its structure, exhibits animal properties; animal substances therefore are also the best nutritive media of plants.
1390. That which putrefies most easily is the best manure.
1391. Through the process of putrefaction many kinds of antagonisms and attractions, by which the absorption takes place through the root-filaments, are aroused.
1392. The root has not merely one orifice for absorbing, but it imbibes upon the whole surface, from its being still immersed in the chemical menstruum. The integument of animals does the same.
b. BARK-PROCESS--EVAPORATION.
1393. The bark, as an organ of cellular tissue, which is placed wholly in the outward direction, must principally exercise the process of absorption and evaporation. Now, as there are two kinds of bark, a root-and a stalk-bark, or a water-and air-bark, so upon the former will the business of absorption chiefly devolve, on the latter that of evaporation.
1394. As the bark of the stalk possesses stomata, which are wanting in that of the root, so is this a probable reason for these apertures being organs of evaporation. This opinion is corroborated also by aquatic leaves being without stomata, while they occur in the leaves exposed to air.
1395. Meanwhile the stalk is of a twofold character; it is only the root that has ascended into the air. As an aerial root it absorbs. Without doubt the stalk absorbs the same as the root, namely, moisture from the air and carbonic acid. Experiments prove it.
c. CELL-PROCESS--DIGESTION.
1396. The cells are the crystallized drops of mucus, the fundamental mass of the vegetable and consequently the water, which converts itself into the Earthy, or wherein the Solid has been elaborated and precipitated. They construct the Solid that has been absorbed into new cells. But the Solid can only assume other forms by means of water. The solution, however, with mixture of bodies and formation into globules is digestion. The cells are thus the stomachs of which the plant has millions like mouths.
1397. The bodies absorbed must move in the cells; for chemical solution and mixture, being itself nothing else than separation and union of atoms, is consequently motion. In a single cell the motion must be upon all sides, because the atoms are attracted and repelled from all points of the cell-wall. In cells, however, which are united with others and therefore subjected to longitudinal polarity, this motion must be performed in accordance with the axis of the cells.
1398. This motion proceeds to and fro, because the extremities of the cells have different polarities, and therefore repel the same atoms, which they have before attracted. In the cells the mucus appears to be converted into starch-granules.
B. _Vascular Processes._
a. _Vessel-process--Conveyance of Sap._
1399. The vessels or intercellular passages conduct the sap, or the water of the plant. Their function is therefore the continued conveyance of the sap that has been absorbed from the root, and rendered solid or consistent by the evaporation going on in the bark and elaborated by the cells.
1400. The vessels of plants are notwithstanding to be compared with the lymphatic vessels of animals, in so far also as these are distributed throughout the whole body, and convey the sap simply in one direction not in a circle.
1401. As the passages between all the cells are in all directions, so the vegetable saps or fluids flow in all directions, and not to one centre as in the animal. Plants have no heart. The sap pursues a tolerably rapid course in the vessels. A fading or drooping cabbage, two feet in length, can gradually become erect in a few minutes after being put to soak in water. In other respects the course of the sap in the vessels may be seen in many plants under the microscope.
b. _Liber-process--Mixture of Sap._
1402. In the liber, as being the mass of intercellular passages, the sap contained in the vessels principally accumulates, as in the thoracic duct of animals; in it the matters have not been simply conveyed and dissolved, but also mixed and converted into true vegetable sap, into blood.
1403. The tubes of the liber are those by which the chemical life is sustained.
c. _Stalk-process--Secretion._
1404. The stalk is the root planted in air, and consequently its process is the differenced process of putrefaction, in which the mucus becomes further evolved.
1405. The analysis chiefly occurs in the stalk; the mucus, or rather the starch, becoming converted into sugar and acids.
1406. Sugar is the mucus of the stalk, and is found in every vegetable sap, especially that of such plants as are characterized by the systems of the stalk, and have not yet attained the formation of the reticular leaf, as the Monocotyledones, e. g. the grasses.
1407. The sugar originates from a process of fermentation; the process of the stalk must consequently be regarded as a vital _process of fermentation_.
1408. The process of fermentation is that of putrefaction carried on in the air, or the polar process of fermentation. Both processes consequently observe a polar relation towards each other.
1409. The sugar-process passes over finally into acidification.
1410. The _Inflammables_, as the ætherial oils, balsams, and resins, are formed in the antagonism of the sugar or of the _acids_. Here also belong most of the peculiar vegetable matters, as the milky saps, colouring matters, medicinally active bodies, poisons, and the alkaloids.
C. _Tracheal-processes._
a. _Leaf-process--Inspiration._
1411. In the foliage the woody rings have issued freely into the air, in order that they may offer their whole surface to its influence, and thus become electrified and oxydized.
1412. The leaf is the free, external organ of respiration to the plant; it is its _lung_. Through the leaf the air, and chiefly its oxygen, is transferred into the plant, just as it is through the lungs into the animal.
1413. The leaves take in oxygen gas; this is their _essential_ function, and not that of exhaling it.
1414. The leaves _only_ exhale oxygen gas when exposed to light. The development of oxygen in the plant is accordingly a light-and not an air-process. In consequence of this the leaves give out oxygen gas only during the day, but during the night and even upon gloomy days, where not the light but only the air is active, they take in oxygen and give out carbonic acid.
1415. The light develops the oxygen gas out of the plant in a perfectly inorganic manner, like as from every water, that can be set in a process of tension. _Rumford_ has developed by simple glass tubes oxygen gas out of water. The oxygen gas of plants is therefore a result only of the decomposition of water in an inorganic manner by the agency of light, or virtually the separation only of the oxygen that is clinging to the water.
1416. Through the process of respiration in the plant carbonic acid has been formed and excreted. For the mucus becomes oxydized, and thereby the process also of fermentation, the product of which is carbonic acid, is promoted.
1417. The respiratory process of the leaves is the perfected process of fermentation in the stalk, in which finally, namely, in the fruit-saps, the separation of both the products of fermentation, the vinous and acetic, is prepared.
1418. Just as acids and sugar originate in the stalk, so in the foliage does their electrical antagonism, or the ætherial oils and perfumes. Sweet scents or perfumes are properties of the air, and therefore originate also with the aerial process. This is retrospectively a proof that the leaf-process is the respiratory process.
1419. Through the leaves, with which the whole surface of the earth is covered, the planet respires, and thereby the surface of the earth principally obtains its electricity.
1420. Vegetation must therefore effect an important change in the earth's electricity. The earth must be differently polarized after, to what it was before, the fall of the leaf.
1421. Thereby the northern hemisphere is differently polarized to the southern, because the latter has less soil than the former.
b. _Wood-process--Nutrition._
1422. As most of the spiral vessels are collected together in the body of wood, and finally in the leaves issue forth quite free and naked into the air, so must the wood conduct for the most part air into the plant. The polarization of the other systems, of the liber and the bark, must therefore proceed from the body of wood.
1423. The greatest amount of induration must originate in the body of the spiral vessel, because in it the process of oxydation takes place in the most active manner. From the same cause the process of nutrition must also be supported by it in the most powerful manner. The wood is the chief seat of nutrition.
c. _Tracheal-process--Oxydation._
1424. The structure of the spiral vessels, their resemblance to the tracheæ or air-tubes of insects, their distribution throughout the whole trunk, the air they contain which is found decidedly free in the plant, leave no doubt that the tracheæ are air-conveying organs, and consequently have, like the arteries in animals, the process of respiration directly intrusted to them.
1425. Now through the process of respiration the general polarity, and consequently the cause or fundamental principle of all life, enters the plant.
1426. The tracheæ penetrate or traverse the whole plant from the apex of the root to that of the flower. Their operation must therefore also extend through the whole plant.
1427. The tracheal system must also govern the plant by polarity, and thus in an immaterial manner.
1428. This polarity acts simply in the direction of the plant's longitude, not transversely, like the material fundamental processes.
1429. The tracheæ impart in a spiritual manner the antagonism between the root and fabric of the stem.
1430. As the tracheæ are, or constitute, the highest system of the plant, so must it be them upon which the light principally acts. The material processes of plants are kept in activity by the antagonism of light.
1431. By this only are the instantaneous changes, which follow upon the influence of light or section of the spiral vessels, to be explained. Upon this, therefore, depends the instantaneous elevation of the processes under the influence of a ray of light, and their depression, if only a cloud pass in front of and obscure the sun; hence too does the plant die, so to speak, upon the very spot, if the spiral fibres within the liber be cut through, but the latter left uninjured.
1432. The liber no longer conveys any sap to the divided tracheæ, solely because it has lost the condition to be affected by the light-polarity. On the contrary, a plant does not die so soon, if the liber be cut through, but the spiral vessels preserved. The spiral fibres conditionate consequently the motion and the excitation of the organic processes.
1433. _The spiral fibres are therefore, apart even from their function of respiration, or rather because this is the highest vegetable function, that for the plant, which the nerves are for the animal._
1434. The tracheæ of plants do not ramify like the nerves of animals; but if they divide, they separate only as fascicles, which have been liberated from their origin. The tracheæ commence too directly in the mass of cells, wherever that may happen to be, and thus become what governs an organ, exactly like the animal nerves. Their analogy is greatest with the sympathetic nerves. The tracheæ, just as in the animal kingdom, are the mediators, not the founders, of vegetable life.
1435. The principle of motion must reside in the tracheæ, provided that higher, and not merely chemical movements, occur in the plant.
1436. These movements must and can only exist in those organs which consist almost entirely of spiral vessels, and thus only in the highest organs.
1437. Such are the leaves and the corollæ. Is it wished to compare the corolla, apart from its sexual relation, with an organ in animals, it can only be contrasted with the highest nervous organ. The corolla is the brain of plants, that which corresponds to light, but which here remains stationary upon the sexual stage. It may be said that what is sex in the plant, becomes brain in the animal, or the brain is only the animal sex.
1438. The most general function of the brain is, however, feeling or touch combined with motion. If the corolla could attain to a sensorial function, it would be to that of touch.
1439. It is conducted thereto; but at the instant, when it is indulged in feeling the mental capacity of the animal, it sinks down exhausted and dies. It is punished for the risk that has been run in wishing to attain unto self-cognition.
1440. Motion and touch are revealed only in the highest organs of the plant, or in the stamina. The filament moves upon the pistil and touches it with the pollen, which at that instant, however, is scattered in small particles, and leaves behind the filament in a withered state.
1441. The motion performed by the filaments appears to be a simple operation of the irritability in the tracheæ that have become soft, without undergoing chemical decomposition, but probably by sudden influx of sap, induced by the tension of air in the spiral vessels.
1442. In the highest, or the pinnate, leaves, movements, which are probably a result of the tracheal irritability, also occur, but are devoid the object of coming into contact with, or of touching, anything. The sensitive plants, as Hedysarum gyrans, move their leaves, not from any intrinsic determination upon their part, but in accordance with an antecedent stimulus, and thus not voluntarily, but probably through the influence merely of polar tension. The movements of leaves are convulsions of plants, although too an afflux of sap be caused or induced by the stimulus.
SAP-MOTION.
_Galvanic Process._
1443. _The motion of the sap is imparted through the antagonism of the respiratory and digestive processes._ For these two processes are the combination of the Chemical with the Electric, which is the galvanism.
1444. The galvanic poles attract and repel the fluidity; thus the vegetable sap is attracted by the root and by the stalk. But the differencing or the oxygen pole is the stronger of the two. The determining principle of the movement of the sap resides consequently in the stalk, and the chief direction of the sap-motion tends upwards.
1445. At times, when the air-polarity is elevated, the sap also ascends more rapidly. As in summer, upon clear warm days. It ascends slowly upon gloomy and chill days. That in this also light and heat are playing their parts, is self-intelligible. Thereby the upper particles of sap become lighter and ascend, being pressed upwards by the lower and colder particles. As they are nevertheless by no means changed, this is a proof that, during the time so employed, polar forces also act upon them.
1446. But the root has also the endeavour to attract the sap; but as its pole is feebler in character, the stalk draws the sap from the ultimate extremities of the root into itself. If accordingly the polarity of the air becomes weaker, while the plant is losing its leaves or the organs of polarization; so is it easy to imagine why the motion of the sap becomes slower. As, however, the aerial polarity is always stronger than that of the earth, the sap must thus in winter also take the same, or upward, direction.
1447. A fall or descent of the sap can therefore never take place abstractedly forsooth from the root, in which it sinks by its own gravity. How a part of a plant, e. g. a twig, could continue alive, were the sap to have fallen or receded from it, is not to be conceived. It does not follow from what has been just stated, that movements of sap should not take place in all directions, and consequently too downwards; they must indeed occur rather than otherwise, and that indeed upon all sides; only the principal track or course of the sap must always pass in the direction upwards.
1448. The movement of the sap consists simply in an ascent and impulsion of its particles upon all sides, but without any _circulation_. A circulation would only be possible if the plant were an organism disengaged from the elements; but as the earth and air belong to its organization; it thus necessarily oscillates between both, and its movements also can only be oscillations of a similar intervening character.
1449. There are consequently no arteries and veins, and still less a heart in plants, as some have striven to make out.
1450. The vessels of plants are most properly to be compared with the lymphatic vessels of animals, the fluid or sap of which also tends from all parts towards one summit, namely, the lungs, while still at times retrogressive movements also seem to occur.
1451. The vegetable sap does not move in a straight line upwards, but in all directions, to the right and left, in a zigzag manner, and so on. This is proved by two incisions being made in a branch opposite to each other. The motion of the sap in the plant is more an impulsion of the sap toward all sides, with a predominance in the direction upwards, than a rapid current as in the blood. If we reflect that the motion of the sap when seen under the microscope and then magnified several hundred times, still only resembles a gentle rippling of small drops, it thus becomes clear that the true current or flow occurs only in a very tardy manner. Wherever therefore in the plant the process of differencialization may be brought into play, there the sap is impelled.
1452. Through the polarization of the sap the cells also become polar towards each other, and then even the cell-walls, whereby the cellular sap with its mucous granules is kept in constant motion. The theory of the motion of sap has not consequently been based upon the theory of capillary tubules; nor is heat alone the cause of its ascent; nor the empty space, which originates superiorly by evaporation; nor electricity in an inorganic sense.
II. FUNCTIONS OF THE FLORAL-ORGANS.
1453. These functions correspond to those of the light, heat, and gravity in the corolla, pistil and seed. The corolla irradiates, the pistil gives out heat, the seed sinks like the earth towards the centre.
1. _Function of the Corolla._
FERTILIZATION.
1454. As in the vegetable trunk the principle function has been the antagonism between the aerial and the terrestriaqueous plant, so must the same function be repeated in the corresponding organs of the blossom. It oscillates in the principal antagonism between the corolla and pistil, which is the antagonism of leaf and stem, that of electrism and chemism, of light and body, of spirit and matter.
1455. The pollen electrifies, animates, or inspirits the ovarium, by which means it becomes stimulated to the development of seeds. Without this animating influence the seed had not been developed.
1456. This relation, whereby through the balance of an antagonism, a whole organism has been evoked into life, is called the _sexual relation_.
1457. The sex is consequently the antagonism between spirit and matter, light and mass, æther and the terrestrial elements, sun and planet, between electrism and chemism, that has been represented in an organism as totality. In sex consequently the primary antagonism of the world, or that of spirit and matter, centre and periphery, has been organically represented.
1458. The sex has hence from the beginning been established and prophesied; manifests itself also under diverse forms in the Organic, but becomes first individualized in an organic body. This is the lofty sense or signification of the sexual relation, that in it the Spiritual and Material pair together, and thereby sprout forth or germinate into a whole world. In sex the mystery of creation lies concealed.
1459. What is producing the fruit is called the _female_, that which awakes production, the _male_.
1460. Mascularity is the spirit of the world, feminality the matter which becomes animated by the former; mascularity is the light of the word, which illuminates the feminality, and it is pregnant; mascularity is the electricity of the world, which arouses the female chemism unto galvanic circulation. By the male the female becomes animated; before this it is dead and devoid the differentialization, which is necessary to every action.
1461. Impregnation is a simple act of light upon the matter, an _irradiation_, as it was termed, with such an exalted appreciation of its significance, by the ancients. The male imparts nothing in impregnation but the _solar ray_, or _fluid nervous mass, in its semen_, which awakes, animates, and inspirits the quiescent female. The female supplies or furnishes all the Material or, as in the plant, the fruit. In other respects it is not to be understood, as if no material whatever had been imparted by the male, but only that it is not the matter as such, which the male gives the female, that becomes fruit; but that the tension which resides in male semen, evokes at the same time, as by a process of contagion or fermentation, a similar tension in the female.
1462. The process of tension resides originally in the male, because he is related or akin to the light; but the female first obtains the light through the male.
1463. Impregnation is an excitation by the electrical, of the slumbering chemical, process. The pregnancy is consequently an uninterrupted chemical process.
1464. The female is the first and lies deeper in the developmental history of the planet (but not in the creation), just as the digestive process is prior or antecedent to the respiratory.
1465. In the truest sense is feminality co-ordinated with the digestive, the mascularity with the respiratory, system. The female is (organically considered) abdomen, the male, thorax. Pregnancy is a sexual process of digestion, impregnation a sexual process of respiration. In impregnation the female respires the male, whereby it receives into itself a thoracic function, becomes itself male, i. e. is then capable of producing something out of itself. Now, the female produces a fruit, which is synonymous with both principles.
1466. The semen is the fruit of the male. The male is always pregnant, and that indeed by virtue of his own power. This power is deficient in the female, which does not possess the light in itself, but only the elemental bodies that are ready and susceptible of form.
1467. The anthers are the male organs, the pollen is the semen. The pistils are the female organs, the seed-granules are properly speaking, the _germ_.
1468. The pollen is a most highly differenced, electrical product; the seed-granule a wholly indifferent, and tranquil mucous mass. The pollen falls upon the stigma of the pistil, and irradiation has taken place; the material fruit-capsule gains thereby so much polarity, that saps enough ascend, in order to develop the germless seed-vesicles.
1469. It is quite unnecessary for the pollen, with its sap or gas, to be materially conveyed through the style to the seed. It is only requisite for the style to be excited, dualized, electrified, and then it has life enough of its own. But it does not follow, because it is unnecessary for the sap of the floral dust or pollen to reach the seed-granules, that it cannot or ought not to reach thereunto. In many plants the pollen-tube does actually reach there and penetrate through the micropyle. In many styles it is still held as impossible, for the pollen-tube to penetrate through them to the seeds. The pollen-sap indeed simply evokes on the apex of the seed (upon the summit of the lorical rib, through whose liberation the micropyle originates) the vital process, which, without this stimulus, would perish. Thereby a new cell is secreted, from whence the germ is developed.
IRRITABILITY--MOTION.
1470. In impregnation the heaven is married to the earth; for then the spirit descends, and does not esteem itself too highly to become flesh. Impregnation is the highest immaterial action of the plant.
1471. If, therefore, the irritability of the plant at any time, or but once only, makes its appearance independently, it must be in the sexual organs, and in the moments of impregnation. Impregnation ensues, when the two mundane principles of the plant, light and matter, have attained, as corolla and fruit, to the highest pitch of perfection; then the tension of the spiral vessels ranks so high, that they exercise their function independently of what is terrestrial in the plant, _move_ themselves in the male filaments, touch the female organ, and die in this their highest effort.
1472. Thus has it only been conceded to the plant to be, in the instant of impregnation, an animal and enjoy animal passion.
2. _Function of the Ovarium._
1473. The ovary, by its own power, is in a condition to draw towards it the chemical saps from out the stem, and as it were by its own heat to thrust new buds from its leaf-ribs, namely, the seed-pellicles or testæ. It has not, however, strength sufficient to put forth also the leaf-work, namely, the embryo, upon the apex of the seed-shell. It requires for this purpose the stimulus of the floral pollen. If the plant is very rich in sap, the ovary is so likewise, and converts itself into fruit or sarcocarp. As a rule, therefore, trees only bear a crop of fruit. If also the impregnation is less perfect, the force of the sap continues to remain inherent in the ovarian leaves; they become rich in sap, fleshy, and likewise fruity in character; trees, therefore, with imperfect or separated blossoms, as the Amentaceæ, Urticaceæ, Euphorbiaceæ, Papilionaceæ, Terebinthaceæ, and Rosaceæ, usually bring forth a crop of fruit.
1474. A stronger degree of refinement appears in these fruit-saps than in the saps of the stem, because corolla and seeds range closer to each other. They are therefore more varied and richer in substance. The fruit-substances range usually upon the side of the water or the salts, while those of the seed range upon the side of the earth or the Inflammables. The substances of the seed are flour and oil, those of the fruit sugar and acids; the former supplies food, the latter drink.
1475. Seed and ovary stand therefore in antagonism, like earth and water.
3. _Function of the Seed_.
GERMINATION.
1476. The seed is the plant contracted upon its centre, it is the heavy mineral mass, which can only undergo changes by the operation of the other elements, like as it arrived only at completion by the operation of the floral pollen. This acts upon it when within the dry ovary, like the water and oxygen in the dry earth. These changes are its development or germination.
1477. All the planetary elements belong to germination, and to growth the Cosmical also with all its actions. To germination belongs earth, water, and air; to growth, light, heat, and gravity; with all the four mineral classes also, such as earth, salt, Inflammable, and metal. The plant contains silicious and calcareous earth, salts, coal with sulphur, and lastly, iron.
1478. Germination is the disjunctive emergence wrought by means of moisture, heat and oxydation in the processes of decomposition and fermentation. No seed germinates in irrespirable kinds of air.
1479. The cotyledons or seed-lobes are the synthesis of the two processes; they are at once root and leaf, therefore resolvable into mucus, and may yet become green.
1480. In germination the elemental bodies of the root-and stalk-polarity directly emerge; the mucus or the flour separates into alcaline gum, that seeks the darkness, and into acid sugar, which elevates itself into the illuminated air.
GROWTH.
1481. Growth is none other than continued germination. The sap being polarized by the air becomes of necessity decomposed. One part evaporates as carbonic acid and water, the other coagulates into oxydized mucus or into cell-walls.
1482. Growth proceeds directly from the process of digestion and respiration, while its polar organs constantly remove further from each other.
1483. Properly speaking the digestive and respiratory processes are none other than growth, since both separate from each other. That, which originates between them, is the process of nutrition, the vascular system.
1484. Growth oscillates between the process of decomposition and that of fermentation; it is an uninterrupted fermentation.
FALL OF THE LEAF.
1485. If every pole of the plant has been perfected in an isolated manner, it has thus become identical with the air, and the aerial process ceases.
1486. With the cessation of the aerial process, the respiratory organ must also die off or perish.
1487. The decadence, or falling off, of the leaves is the result of the tension having been abrogated between them and the stem; it is a death by suffocation.
1488. The fall of the leaf therefore occurs in the autumn, or after the fruit is matured.
DURATION OF LIFE.
1489. The age of a plant is included between the the limits of the sap's impulse, and that which has been called its fall or descent.
1490. The actual fall of the sap is the death of the plant.
1491. If with the cessation of the influence of light, the polarity ceases entirely in the plant; it is then one year old or an _annual_. Every part of it dies off.
1492. In biennial plants the aerial polarity indeed disappears, but the polarity of the root remains. Flower, leaf, and stalk die.
1493. Perennial plants, also, do not entirely lose the stem-polarity, but only while they develop a new plant about the old. Flower and leaf only perish, while the water-and earth-organs remain alive.
1494. The old liber dies with every maturation of the fruit, because there the difference attains solution. But a new life develops itself in the parenchyma of the plant, and forms new liber or, properly speaking, a new plant about the old.
1495. Persistent plants consist of numerous plants, which gradually grow round about each other.
1496. In accordance with the idea of the plant, each one perishes with the maturation of the fruit.
1497. On account of the addition of the new plant about the old, the plant has also been confined to no definite magnitude and to no definite number in its mode of ramification.
1498. Indefiniteness in form, size and number, is the character of the plant, although a law lies at the basis of all this. The animal has a definite size, because several animals do not grow around each other.
III.--_PHYTOLOGY_.
1499. Hitherto the organs of the plant have been considered in a general point of view or as to their idea in time; to this now follows the development of the plant in a special sense, or its representation in space.
1500. The vegetable tissues, systems, and organs have only by degrees been disengaged from each other and independently perfected. The independent or self-substantial development of the organs constitutes definite or _individual_ plants.
1501. A plant, in which all the organs are present, separately or self-substantially developed and yet combined, is without doubt the highest in point of rank.
1502. Before it attains to this separation, nature can only produce lower forms, in which fewer organs have attained to independence. These forms constitute the diversity of plants and their plurality, for nature establishes every principal form as a finished organization.
1503. There are as many plants different from each other as there are organs, namely, tissues, anatomical systems and members.
1504. The sum of all plants is called the vegetable kingdom; _this is the self-substantial representation of all vegetable organs_. (Ed. 1st, 1810, p. 123.)
1505. The vegetable kingdom is consequently the expression of the vegetable idea, or of the perfect plant represented _in the multiplicity of individuals_; it is the plant disintegrated, or anatomised, by nature herself.
1506. Were we therefore acquainted with all vegetable organs, we should know their rank and developmental series; and thus also recognize the character, rank, and developmental series of the plants themselves, or their divisions. There can be no doubt that the lowest organs, e. g. the tissues, have been first developed and independently perfected as plants; later on they separate into anatomical systems and finally into members, whereby perfect plants must originate. _The division or classification of the vegetable kingdom is consequently that of the vegetable organs._ The Systematic of plants is a copy of that of their organs, or a plastic representation of the philosophical vegetable anatomy. With this every thing has been granted, which is requisite for the building or erection of the vegetable system. All principles, together with the methods, rest in the proposition that has been expressed.
1507. The artificial systems of plants are related to the vegetable kingdom, as the lexicon or dictionary is to language. Those systems which have hitherto been termed natural, but which should properly be called _methodical_, are related to the vegetable kingdom, as the ordinary grammar is to a language. The vegetable system must, however, be related to the vegetable kingdom, as the philosophical or genetic grammar is to language. This only agrees with the essence of the language, or is _natural_. The vegetable system is necessarily a philosophical or genetic one, that alone being truly or legitimately natural. (This system was first propounded by Oken, in the Ed. 1st of the Naturphilosophie, 1810; further developed in _Dietrich's_ Garten Journal, 1813; carried out in his Naturgeschichte fur Schülen, 1821, and in his Lehrbuch der Nat. Gesch. _Botanik_. Weimar, 1825.) The _artificial_ system collects the materials for the edifice, but leaves them to lie without order and in confusion; the _methodical_ or what has been called the natural system separates these materials and arranges them in homogeneous groups; the _genetic_, philosophical or truly natural, system, again mixes them amongst each other, but thereby actually erects the edifice. All three systems are therefore necessary and good, and no one of them merits being despised by the other; it is only when one of them imagines that it is the other, or can render the others unnecessary, that it trespasses from out its circle, and deserves reproach. Thus for _Floras_, whose ultimate object is to find out rapidly the names of plants upon botanical excursions, as also for the labelling of specimens in botanic gardens, the artificial system is the best; for the description, however, of foreign plants the methodical; but for insight into the whole vegetable world the philosophical or natural system. Would we compare Floras with each other, the latter system must certainly come into play; but then the matter to be dealt with is not about an excursion-book.
_VEGETABLE SYSTEM._
1508. Taken in a strict sense all the diversity of vegetable structure of vegetables has reference first of all to the difference in the tissues; these being either unseparated, or separated, into special systems and members. At first the tissues lie confusedly, or without order, amongst themselves. They then _separate_ in a concentric and tubular form into systems, that are encased _within_ each other, like the bark, liber, and wood, which form the _shaft_. Furthermore they separate into members, and appear one _above_ the other, as root, stalk, and foliage, which collectively may be called the _stem_; these are repeated as seed, pistil, and corolla, which together are called _flower_, and combined, _fruit_, namely, nut, plum, berry and apple. I designate by the term _stock_ or trunk all the parts as far as the blossom; and this together with the fruit I name thyrsus. The vegetable stock, whose tissues have not yet separated into members, I style, from want of a better word, _thallus_. As we divide political kingdoms into _provinces_ and _circles_, so also may these titles be suitably applied here. It is evident, that the plants which simply consist of tissues and have as yet no sheaths and members, are the _Acotyledones_; those, however, provided with sheaths, but devoid of true roots, stalk, and foliage, _Monocotyledones_; those with true foliage or reticular-veined leaves are, on the contrary, _Dicotyledones_. The natural system of vegetables stands accordingly in the following manner.
A.--STOCK-PLANTS.
Province I. HISTOPHYTA, OR TISSUE-PLANTS--Acotyledones. Class 1. Cell-plants. 2. Duct- 3. Trachea-
II. THECOPHYTA, OR SHEATH-PLANTS--Monocotyledones.
4. Bark-plants. 5. Liber- 6. Wood-
III. ARTHROPHYTA, OR MEMBER-PLANTS--Dicotyledones.
Circle 1. _Axis-plants_--Tubulifloræ. 7. Root-plants. 8. Stalk- 9. Leaf-
B.--BLOSSOM-PLANTS.
Circle 2. _Flower-plants_--Thalamopetalæ. 10. Seed-plants. 11. Ovarium- 12. Corolla-
Circle 3. _Fruit-plants_--Calycopetalæ. 13. Nut-plants. 14. Plum- 15. Berry- 16. Apple-
1509. A slight glance at the above table shows us the procedure of Nature. The higher she ascends, the more and more she separates, and thereby increases, the organs. There may therefore be plants which have only a single organ or tissue, as well as others, which possess all.
1510. There cannot, however, be any plant which could simply possess the higher without the lower organs. Higher organized plants are not such therefore, by virtue of their having some one organ more perfectly developed, or separated into several parts; but through this, that they actually possess several different organs. The higher grade of organization depends accordingly not upon the perfection of the _Singular_, but the _number_ of the _Different_. The Perfected consists in the _multiplicity_ combined to constitute _unity_, but by no means in the simply homogeneous multitude of the _parts_. Numerous stamina may render a _corolla_ higher, but not on that account the whole plant; many digits may make a hand nobler, but not on that account, the animal. But with many digits also that hand is nobler, in which the digits are dissimilar.
_First Province._
HISTOPHYTA--ACOTYLEDONES.
_Devoid of, or without true spiral-vessels, leaves, corollæ, and pistil._
The vegetable kingdom ascends, in accordance with the five main positions of the organs, by five stages; these are again separable into larger groups, which may be called _asexual_ and _sexual_ plants.
1511. The tissues are a something internal, being, as it were, the viscera of plants or their parenchyma, which does not meet the light, and can therefore have no light-organs, which are developed only out of the foliage or leaves. The anatomical systems and organs are tissues that have become external, have attained to air and light, and are hence developed into air-and light-organs. Now, the light-organs are sexual organs. The Tissue-plants can therefore have no sexual organs; and plants divide accordingly into asexual and sexual plants. The asexual are female plants, and are consequently the first or lowest. Thus there can be thus no sexual or male plants, without the female being found that belong to them.
1512. Male or androgynous plants are only possible, if spiral vessels or tracheæ be present. They first, however, originate when the tracheæ become external, or form a circle in the stalk, i. e. are accessible by light; as in the Mono-and Dicotyledones.
1513. The asexual plants are not cryptogamic, but agamic. They do not perform self-impregnation clandestinely, but not at all; for they do not attain light-difference, and consequently not male organs. Analogues of stamina may make their appearance in the mosses, but they invariably fail to attain the development of pollen. What have been called male parts in other cryptogamia, do not merit consideration. Such projections or prefigurations are besides to be found everywhere.
1514. The asexual plants are simply formations of the tissues, of the galvanic vesicle, and are thus of a female nature. They are nothing more than a great utricle full of small vesicles, which by desiccation subdivide into _germinal dust_ or sporules, each granule whereof attracts other mucous vesicles out of the moisture, in order to form again a large utricle.
1515. The asexuals cease in the process of vegetation, where the other plants begin. With the rupture of the gemmal-or bud-vesicle in the higher plants a new world for the first time emerges into view, such as stem, leaves, blossom, and then the ultimate bud ruptures for the first time as the pericarp, and scatters its higher organized germinal powder as true seeds.
1516. An asexual plant is one, which, without all the intermediate organs of the stem, at once represents the capsule or ovary. It consists only of the beginning and end of the plant.
1517. The higher plants differ from the lower by the interposition of new organs between the two terminal organs, namely, the primary vesicle and the true seed. It may be said, that the asexual plant is naught but seed, and that the seed of the higher plants is a fungus upon a leafy peduncle, a fungus more highly organized by light.
1518. The asexual plants have no true root, stalk, and leaf; they have not even a true bark, liber, and wood, in so far as these first make their appearance through separation. Tracheæ are first exhibited in the higher ferns, and then only as constituting a single string, which occupies the middle of the plant, and consequently forms no circle or zone.
1519. As again the true seed is a leaf-formation, and possesses therefore cotyledons or seed-lobes, such seeds must be wanting in the asexual plants; they are therefore Acotyledones. From the same cause, however, the germinal leaves or plumula must be also wanting; they are therefore germless, or anembryonic.
1520. The farinaceous or granular matter, lying next to the germ when within the shell of the true seeds, is called the _albumen_ or perisperm; the seeds of the asexual plants are therefore nothing else but albumen. They are therefore devoid of the funiculus or, what has been called, umbilical cord.
1521. The involucre, wherein, in true seeds, the germ and albumen are found, is the seed-coat or testa; consequently what has been called the capsule of the asexual plants (of mosses and ferns) corresponds simply to this spermoderm or seed-covering, and is no true ovarium. The capsules of mosses and ferns are therefore seeds full of albuminous dust.
1522. If any of these be regarded as a capsule, it can be the _calyptra_ of the mosses. This, too, is probably nothing else than the external testa; the proper capsule being its internal coat.
1523. The _indusium_ of ferns incloses several capsules as they have been called, or properly seeds, and might therefore, if considered alone, be compared with an ovarium, but it is probably none other than the covering corresponding to the perichoetium that surrounds the base of the setæ in mosses. The sorus is an accumulation of seeds with pulverulent albumen contained in a membranaceous covering, the indusium.
1524. The life of the asexual plants consists simply in the galvanic process. They are the primary organisms, planted in air.
1525. As being simply galvanic process, they would require but little light and air; they therefore seek the darkness, like the roots, and thrive also in a corrupt atmosphere, in caves, mines, cellars, and such like situations. They can from the same cause thrive only in moisture, in water, upon marshy meadow lands, after rain, copious dew, and so on.
1526. They are devoid the process of fermentation, as being that which is imparted by the oxydation of air, and they therefore yield neither sugar nor acids. They are simply the organized process of putrefaction; their ultimate product is therefore germinal powder, infusorial matter. Their remaining secretions are alkaline bodies; to which belong the pungent, fetid, and nauseous excretions, as the hydrogen gas and ammonia of the fungi, the mucus of the fuci, the carbonate of lime in the lichens, the cell-threads of the mosses, the fetid principle of ferns.
1527. Very few of these plants require the course of a summer in order to perfect or complete the vital course; a single ray of light of one day's, aye, of one hour's duration, is sufficient with most of them to evoke the feeble difference, to rouse the swell of the sap, and precipitate the infusorial powder.
1528. Automatic movements, as in the leaves and stamina of the higher plants, scarcely occur in them, or at most in the ferns, from their possessing spiral vessels. They divide, according to the tissues, into three classes, into Cell-, Vessel-, and Trachea-plants.
_CLASS I._
_Cell-plants--Fungi._
Here belong those plants which consist simply of cellular tissue, having no sap-tubes and tracheæ. Such plants, too, possess no regular or hexagonal cellular tissue.
1529. The cellular tissue, in which there is only a single active process, cannot essentially alter its primary form. It is therefore an accumulation of round or cylindrical mucus-vesicles.
1530. Mucus-vesicles, in which the air-process is not as yet active, cannot be coloured green; but must have the colour of the earth.
1531. Plants, composed of amorphous and earth-coloured cellular tissue, are _Fungi_. The fungi are simply clusters of mucus-vesicles joined together in a more or less regular manner, their union being effected in dark, hollow and wet situations.
1532. They may therefore originate wherever mucous juices are evolved by the potential agency of a higher organization, and thus by putrefaction. The fungi originate by _æquivocal generation_. They are the anal-organizations of the higher plants and animals; the corrupted and luxuriating juices.
1533. Nevertheless the fungus is propagated by division of its vesicles, which again, in accordance with their peculiar laws of polarity, attract mucus-vesicles, and thus obtain the form of the earlier or parent fungus. This is only a more regulated kind of _æquivocal generation_.
1534. The origin of the fungi may therefore happen in a twofold manner, namely, by formation from other juices, and by that of their own, which is called propagation. Still at bottom both are one in kind.
1535. Their granules or vesicles are seeds, properly sporules, which are self-developed without male polarization.
DIVISION.
1536. The fungi pass moreover through stages of development, which range parallel to the vegetable classes; since it is impossible for any other organs to originate in them but such as belong to the idea of the plant. The lowest fungus can therefore change only, by endeavouring to develop in itself ducts, tracheæ, roots, and such like parts.
1537. There are accordingly as many developmental stages of the fungi as there are vegetable classes. These divisions are called _families_.
1538. The vegetable families range parallel to the classes. This law must hold good of all the classes. There are therefore in each class 16 families. An association of families upon each stage may be called an order.
1539. At first the fungus is none other than a mucus-vesicle or a small cluster of vesicles, e. g. an _uredo_ or _mildew_. Such a vesicle next becomes longitudinally extended, and includes within itself other vesicles or granules, e. g. _mould_. These mould-filaments or threads unite again so as to form a common mass, which is surrounded by an external membrane, and is then called _puff-ball_. The pulverulent granules, which were irregularly accumulated in the puff-balls, unite at length in a regular manner to constitute a trunk of varied form, as in the _ascomycetes_, e. g. _sphæriæ_. Finally, the mould-filaments with their sporules are regularly collected together in an investing membrane, which, like a puff-ball, is supported upon a stipes or stem, e. g. the sarcomycetes or _agarics_. There are therefore 5 developmental stages of the fungi, which correspond to those of the classes; viz. the parenchyma, shaft, stem, flower, and fruit; and constitute orders.
1540. Each order is resolvable again into three divisions or families, which correspond to the organs. Thus there are in each class 16 tribes or families, which obviously range parallel to the vegetable organs or classes. (Vid. Tab. B.)
1541. A tribe or family is consequently the representation of a vegetable organ within a class.
1542. The genera obey the same law; for essential differences are only conceivable through the presence of different organs.
1543. Species is in the animal kingdom that which copulates without necessity and compulsion. The same definition is applicable to plants. The species range, without doubt, according to the diversities in the individual organs themselves, which admit of a great multitude of combinations, the number of which is not as yet to be determined.
1544. The component parts of the fungi are either perfectly indifferent, mucous or gelatinoid matter; or they are of an alcaline nature, being acrid, poisonous, and such like. Their odour is usually dead, disagreeable, and loathsome, or analogous to their essential process of decomposition.
_CLASS II._
_Vessel-or Duct-plants--Mosses._
1545. The intercellular passages or succigerent vessels of plants make their first appearance in a state of perfection, when, the cells being extended lengthways, have become hexagonal and are placed in regular juxtaposition. In these plants therefore we meet with regular cellular tissue, but still without spiral vessels or tracheæ.
1546. As the vessels or ducts constitute the fundamental tissue of the liber, while this is the principal system of the stalk; so now does the stem begin to be manifested and separated from the fruit. The seeds are no longer therefore distributed in the present class throughout the whole trunk, but developed in a special involucrum or theca, which corresponds to the puff-ball, or to the pileus of the higher organized fungi.
1547. Plants with vessels, and consequently a cauliform formation, have at once also the commencement of a bark, and next the green colour. The vascular are the first green plants, and differ chiefly through that character from the fungi. They are the _Fucacæ_ or _Sea-wracks_.
1548. They have the colour of the water, because the course of the sap corresponds to the aqueous process; they are aquatic, just as the brown fungi are terrestrial, plants. Their component parts are aqueous, indifferent, mucous, and filose. Their habitation is the water itself or bogs. If they occupy dry situations, they live only when it rains.
1549. They likewise pass through the five stages of vegetation, and form therefore five orders.
1550. Order 1. The lowest or _Tissue-mosses_, corresponding to the Uredines; are again naught but cells or mucous pellicles, but, from growing in water, and being consequently exposed to light and a stronger oxydation, they are green--_Tremellini_.
1551. They multiply by subdivision, since new vesicles or granules are developed in their interior, which become separated, and subsist or continue to grow for themselves. They therefore originate also by æquivocal generation, but by such an one as constantly occurs in water and light.
1552. The second order, or the _Vascular mosses_ corresponds to the sheaths, or to the Hyphomycetes. They are long filaments replete with granules, growing in water, and therefore green--_Confervaceæ_. These plants begin to ramify, and either increase by this means or by effusion of granular matter.
1553. The third order, that of the _Tracheal mosses_, corresponds to the stem, or to the Gasteromycetes. A membranous trunk originates in water, which in certain places secretes the seeds in special vesicles or cysts--_Fucales_ or connate Confervæ. The fuci have at once the form of a stalk with root and leaves, because they correspond to these three organs of the axis.
1554. The fourth order, the _Floral mosses_, endeavours to obtain the _blossom_, and therefore elevates itself out of the water, but loses on that account the trunk-like character, and exhibits for the most part only membranous expansions, upon which seeds are secreted, which being usually of a beautiful colour, thus assume the appearance of corollæ--_Lichenales_. The lichens are fuci in dry situations. They correspond to the Pyrenomycetes or Sphæriaceæ.
1555. As the variegated colours appear in the blossom, so do they also in the lichens; but here they are for the first time chemically developed, distributed throughout the whole substance, and concealed. Most lichens yield colouring matters.
1556. As the stem is, in accordance with their signification, wanting to the lichens, they thus require a foreign trunk for their nutrition. They are therefore developed for the most part upon other plants, and principally upon the bark.
1557. Lastly, the fifth order, _Fruit-mosses_, originates through the development of a self-substantial _fruit_ upon a cauliform stem--the _Mosses_ proper.
1558. As these are the highest plants of this class, and those that directly precede the tracheal formation, so the bark already resolves itself into individual leaves, which are, however, still destitute of spiral vessels.
1559. What have been called the seeds or sporules are accumulated in a capsule-like fruit upon the summit of the stalk. This fruit corresponds to the pileated fungus, and therefore springs up in an opercular manner like the latter.
1560. But this capsule is only a spermoderm, which incloses albuminous granules that have no proper germ or seed-lobes; they are plants with seed-vessels or pyxidia, upon an open-leaved stalk.
1561. They divide likewise into sixteen families. (Vid. Tab. B.)
_CLASS III._
_Trachea-plants--Ferns._
1562. At first a fascicle only of spiral vessels can originate, which is necessarily surrounded by cellular tissue, and therefore lies in the middle of the plant. Such plants are the _Filices_ or ferns.
1563. As the spiral vessels are the antetype of the leaves, so does the trunk here obtain the form of the leaf, without itself producing true leaves. For, in the ferns, the fruits lie upon the back of the apparent leaf, which can only be the trunk.
1564. The fruits, being further removed from the fungi, no longer spring up in an opercular, but in a valvular, manner like the higher capsules.
1565. Green plants with imperfect spiral vessels and blossoms, and also with naked seeds devoid of true capsules, belong to the class of Ferns.
1566. I therefore place in this class the _Coniferæ_ or trees with acicular foliage, because they have no ovarium, but naked seeds; and besides these, some other plants, though doubtfully, on account of their very abortive blossoms, as the Naiadaceæ. There are therefore Tracheal plants without and with stamina. The first portray the stock or trunk, the second, the thyrsus or blossom; they live mostly in dry situations, and produce resins or fetid matters.
1567. First order. _Parenchymatous ferns--Aquatic ferns._ I here place the aquatic ferns, because, as water-plants, they occupy a lower situation, because they support the fruits upon a radical trunk, and finally, because these fruit-vesicles have two kinds of contents, all of which seems to remind us of the fuci and lichens; they correspond to the Tremellini.
1568. Second order. _Sheath-ferns--Club-ferns._ Here commence the land ferns, and those kinds indeed whose so-called capsules open in a valvular manner, just as in the liverworts; or almost after the fashion of a pyxidium by an orifice, somewhat as in the mosses; the trunk is provided with squamose leaves or lobes, e. g. Lycopodiaceæ and Osmundaceæ; they correspond to the Confervaceæ.
1569. Third order. _Stem-ferns--Annular or Ring-ferns._ Here we meet with phylloidal involuted capsules or seeds upon the back of a stem that is likewise leaf-like; e. g. the typical or true ferns.
1570. They have rudiments of roots and a stem, together with foliage, because they are the prototypes of these three organs.
1571. The ring of their capsules corresponds to the midrib of the leaf. In the preceding order the capsule was only an upsprung stalk; but it is here an upsprung and unfolded leaf, the prototype of the bud-development.
1572. The fern-capsules, namely, the true seeds, are an accumulation of leaf-buds at the extremity of the fascicle of spiral vessels. The indusium is the upraised epidermis, which opens in a spathose manner; it consequently stands in the signification perhaps of the floral spathe or involucrum. They correspond to the Fuci.
1573. Fourth order. _Floral ferns--Fluviales._ If the tracheal plants be exalted unto the flower, male organs cannot fail in at once beginning to develop. I place therefore in this order the Naiadaceæ with very arrested blossoms, and simply stamina without calyx and corolla. The spiral vessels are rather doubtful.
1574. Fifth order. _Fruit ferns--Coniferæ._ Tree with imperfectly-formed spiral vessels, stamina without corollæ, seeds without ovarium; thus agreeing with the Cryptogamia even to the stamina; they form likewise sixteen families. (Vid. Tab. B.)
SEXUAL PLANTS.
1575. So soon as the three tissues separate completely from each other into bark, liber, and wood, while the tracheæ are arranged circularly into several clusters or groups, does the antagonism of these organs also make its appearance, and exhibit itself as sex in the floral organs.
1576. This separation can only be attained through the influence of the air and light, whereby the sexual organs are conditionated. These plants have therefore the several organs of the trunk and blossom.
1577. Anthers can be wanting to none of the following plants. Now, the anthers are leaf-buds; the leaf-formation must be therefore developed also in their antagonism, or in the seed. The leaves of the seed, however, are called seed-lobes; consequently all seeds of sexual plants have seed-lobes or cotyledons.
1578. Now the perfect seed is the whole plant in miniature with root, stalk, and leaf. This formation is, however, only possible where there is a sex, or where the vegetable tissues have emerged self-substantially from each other.
1579. But the anthers are buds upon a floral rib; consequently all sexual plants must have a part of the blossom, which ranks in the signification of leaf, and thus either the calyx or, with this also, the corolla.
1580. All sexual plants must have a shaft or scape, in which its three parts, bark, liber, and wood, are to be distinguished; even so must the three parts of the stem, the root, stalk, and leaf, have the parts of the blossom which correspond to them, viz. seed, ovarium, and calyx or corolla.
1581. The sexual plants next divide into Stock-and Blossom-plants, the former of these parting into Shaft-and Stem-plants.
_Second Province._
SHAFT-PLANTS--MONOCOTYLEDONES.
1582. In the Shaft-plants the invaginate character is predominant, the wood being surrounded by liber, and this by bark; they are therefore tubular in form--_tubular plants_.
1583. The cellular tissue preponderates in them, and the tracheal fascicles form therefore no closed circle, but are dispersed; scattered fasciculi cannot, however, ramify but only pursue a straight course and form streaks--_striated plants_.
1584. Ramification is wanting unto these plants, partly on account of their tubular form, partly on account of the smaller quantity of tracheal fascicles--_aramose plants_. A vegetable stem without branches is called shaft or scape; thus they are _shaft-plants_.
1585. The blossoms do not stand upon ramules, but upon the stem itself; thus, if ever ramification originates, it can first occur in the floral peduncles.
1586. As they are deficient in branches, so also are they in buds--_agemmal_ or _budless plants_.
1587. The tendency to produce branches forms nodes, which are only imperfect ramal zones--_nodose plants_.
1588. Where the formation of branches is wanting, the minor ramification of ribs must be also wanting in the leaves; they are parallel-ribbed.
1589. The leaf is only a ruptured and expanded tube or spathe, which surrounds the shaft--_spathose plants_.
1590. Where branches are wanting, there can be also no petiolated leaves; they have therefore only radical leaves, from the midst of which the shaft or scape sprouts forth, only as a floral-peduncle--_radical-leaved plants_.
1591. If all these leaves be still sheathed or encased within each other, they are called bulbs--_bulbose plants_.
1592. Since the corolla is the repetition of the leaves, so also must its structure resemble theirs--_spathose corollæ_.
1593. But as there are here only radical leaves, so also is the corolla only a radical leaf-corolla. In the blossom, however, the radical leaf has become spathe, the cauline leaf, calyx, the ramuscular leaf, corolla. Such blossoms consist therefore either of a spathe only, as in the Aroideæ, or of a calyx--_calycine plants_.
1594. The stamina originate from the calyx. In calycine flowers therefore all the stamina must range opposite to the corolla-petals. These plants have only _opposite_, no _alternate stamina_. It has been generally assumed that the Monocotyledones have no true corollæ, but only coloured calyces. Philosophy agrees with this opinion; but adds thereunto that the corollæ of the Monocotyledones may be also merely spathes or involucra.
1595. The separation of the bud-formation here takes place for the first time in the blossom, namely, in the stamina. The number of the floral parts must be limited to three; for this is the first number, in which the leaf-ribs may divide. Corollæ, stamina, and capsules are ternary--_trinity-plants_.
1596. As the seed is only an undeveloped leaf-bud, so in it only can the number of the leaf be found. Since, however, the leaf does not ramify in the present group of plants, but is only a single spathose leaf; so also does the seed consist only of one such leaf or of one seed-lobe--_Monocotyledones_. The wheat-seed or grain is none other than a grass-leaf with a short spathe and very dense, highly farinaceous, lamellæ. In germinating a new leaf emerges from the short spathe; it is the germinal leaf.
1597. Thus plants having a tubular stem, and such kind of foliage, corollæ and seeds, are Monocotyledones.
1598. The chemical bodies are more diversified in these than in the remaining members of the vegetable kingdom. In the roots, as in the tubers of the Orchideæ we meet with distinct mucilage; in the bulbs with alkalies or acrid matter; and with sugar, as being a feeble conversion of the starch in the stalk. The mucilage of the root becomes, when repeated in the seed, flour. Oily matters or acids seldom occur, and fleshy fruits scarcely ever.
1599. The ovary is almost throughout this region of plants either a single spathose leaf or caryopsis; or three spathe-leaves are united together, which thus, as follicles or carpels, usually burst open upon the internal edge.
DIVISION.
1600. If we now proceed to survey the Monocotyledons, in the order just set before us, we shall recognize among them three typical groups, to which the others are allied; they are the Gramineæ, Liliaceæ, and Palmaceæ.
1601. The Grasses obviously rank the lowest as well in respect of their root, stalk, and foliage, as also of their stunted blossoms, ovarium, and seeds.
1602. To them succeed the Lilies, which have a well-marked root, a more perfect, though still always herbaceous, stalk, and a few spathoid, more numerously ribbed leaves; lastly, their corollæ, ovaria, and seeds are perfect; but still they invariably have no genuine fruits.
1603. Finally, the Palms are elevated above all by their stem being rich in tracheæ and wooded, as also by the perfection of their fruit. Scientifically, the scapose or shaft-plants must also resolve themselves into three classes; into Bark-, Liber-, and Wood-plants.
_CLASS IV._
_Bark-plants--Gramineæ._
1604. In these plants the whole stalk must have assumed the form of the cortex or bark, and consequently be hollow--_tubular_, or _culmaceous plants_.
1605. A mere bark cannot ramify. But the tendency unto ramification is manifested as nodes--_nodose plants_.
1606. The leaf which still represents the bark, is only imperfectly slit up, and therefore still forms a tube. Such leaves are called tubular or spathe-leaves proper.
1607. Such tubular leaves being only half slit up can only shoot forth gradually from each other, and that indeed in such a manner that they stand actually by twos, encased in, or opposite to, each other.
1608. Since the blossom is, as it were, an impression, or copy of the leaves, so will it here also consist only of spathiform, involucral, or calycine leaves, and only of two, one of which, though opposite to, is surrounded by the other. Such floral parts are called glumes--_glumaceous plants_. If four glumes are present, then the external pair of them corresponds to the involucrum or spatha, the internal to the calyx.
1609. The corolla-petals are of necessity arrested in plants such as these, where no true leaf is as yet developed; frequently two only are left persistent as pellicles or _lodiculæ_.
1610. This is still more the case with the ovary and seed; in each only one leaf attains development, and the seed has entirely coalesced with the ovarium. This kind of fruit is called _caryopsis_.
1611. Nodose plants with hollow scape or shaft, tubular leaves, glumose blossoms, and cariopsidal fruits are _Grasses_.
DIVISION.
1612. The Bark-plants pass moreover, together with their subdivisions, through the five stages of vegetable organs, and they will therefore produce also a more perfect stalk, leaves, and flowers. The whole calyx will, however, never be coloured or corolla-like in its character. Cortical plants are thus herbaceous plants with hollow stalk, and with an arrested or green calyx, without sarcose or fleshy fruit.
1613. Those, which have simply glumose flowers, are without doubt the lowest in rank, as corresponding to the tissues and stock, but not yet to the flower.
1614. These again divide into two great groups, whereof the one includes plants with simply uni-seminal cariopsides, the other, on the contrary, capsules containing a free seed--_Grasses_ and _Reed-grasses_. The grasses which have a cariopsis or grain-fruit reascend by two stages; the most inferior in rank do not attain to a ramification, but the flowers stand crowded together in spikes; the others, on the contrary, are pedunculated and ramify in panicles.
1615. Among the higher kinds instead of glumes there are regular flowers, of which, however, the calyx is still glumose or at least green. The cariopsis is converted into a multilocular capsule, as in the Restiaceæ, Commelineæ, &c.
1616. The first order, or the _Parenchymatous grasses_, have glumose flowers with a cariopsis for fruit, borne upon culms or nodose straws, and do not attain to any ramification--_spicate grasses_. In their seeds we meet with the greatest amount of starch developed, though doubtless at the cost of the trunk.
1617. The second order of _Spathose grasses_ are similar, and support ramified flowers--_paniculate grasses_. In this division occur grasses of a dendroidal character, and having occasionally fruit-like ovaria, as in the bamboos.
1618. The third order consists of the _Cauline grasses_. Here the leaves at once separate completely from the shaft, which is therefore free from nodes. Nut-like capsules, though still inclosed in glumes, also make their appearance--_reed-grasses_.
1619. The fourth order, _Floral grasses_. The leaves are still only radical; the shaft is anodal; the flowers are separated into green calyx and coloured corolla, with three or six stamina, and mostly with a trilocular capsule--Junceæ, to which the Commelinaceæ are allied.
1620. The fifth order, _Carpal grasses_. Hollow anodal shafts, with scarcely spathiform, mostly broad leaves, the ribs of which begin to ramify; with similar calyces and corollæ, and numerous capsules--_Seeroseæ_, to which the _Alismaceæ_ and _Hydrocharideæ_ are related.
1621. The highest kind of fruit attained by this class is nut-like, never fleshy in character. The stalk is nowhere woody as in some grasses. The grasses divide, as do all other orders of plants, into sixteen families. (Vid. Tab. B.)
_CLASS V._
_Liber-plants_--_Liliaceæ._
1622. The substance of the stalk is soft and succulent; its structure devoid of nodes; the leaves are tolerably free and ribbed; calyx and corolla coloured, both being perfectly formed and invariably tripartite, as is likewise the capsule, which has many seeds upon its inner angle. These plants are the _Lilies_. In the present class all parts have been developed in conformity to the liber, are rich in sap, and have become dense and fleshy.
1623. The roots are mostly tubers or bulbs, containing a superior kind of mucilage, or aromatic principles. The shaft is not hollow but, though herbaceous, filled up; the leaves are elevated upon the stalk. One division of these plants has irregular corollæ with stunted stamina and capsules, the latter containing mostly dust-like seeds, as in the Orchideæ and aromatic plants.
1624. The other division has regular 2 × 3ary corollæ, with, perfect glume-capsules and middling-sized seeds, as the Irideæ and Liliaceæ proper.
1625. The first order, _Parenchymatose lilies_. The corollæ are irregular, bilabiate, and stand upon the calyx and a membranous sexlocular capsule with very small seeds; are divided according to the pollen--e. g. _pulverulent Orchideæ_.
1626. The second order, _Sheath-lilies_. Characters similar to those of the preceding order, but the pollen is agglomerated into waxy granules--_granular orchideæ_.
1627. The third order, _Stem-lilies_. The corollæ are likewise bilabiate and situated above the calyx, but the ovarium contains few seeds, and the capsule is mostly nut-like--_aromatic plants_, such as the _Scitamineæ_ and _Musaceæ_.
1628. The fourth order _Floral-lilies_, have regular blossoms, separated into calyx and corolla, placed _above_ the capsule, and mostly furnished with three stamina. Here belong the _Hypoxideæ_, _Hæmodoraceæ_ and _Irideæ_.
1629. The fifth order, _Fruit-lilies_. The leaves have not yet completely separated from each other, but still form bulbs; the corollæ are regular, have six stamina, and are placed _beneath_ the capsule, as in the _true lilies_. To this order belong the _Colchicaceæ_, _Aloinæ_ and _Liliaceæ_. Their bulbs contain mostly acrid matters. They divide into sixteen families. (Vid. Tab. B.)
_CLASS VI._
_Wood-plants--Palmaceæ._
_Plants with woody shaft and with fruits, mostly enclosed in spadices._
1630. The desiccation of the cells and fibres is promoted by the increased process of oxydation. Where therefore the tracheæ attain the preponderance, there the conversion into wood originates.
1631. The stalks of these plants are not hollow, but have a dense interior, because the fasciculi of tracheæ lie within the liber, and there increase.
1632. The main bulk of the stalk will consist of tracheæ.
1633. As the tracheæ are longitudinal organs, and the other tissues also extend lengthways, so does the stalk or stem in these plants predominate over the other parts.
1634. In this class the most perfect leaves as regards their present stage are developed; for they are only expansions of the tracheæ, which are here present in superabundance. As regards the form also, these leaves must rank higher than those of the preceding class; the spathe is shorter, the leaf itself usually broad, many-ribbed, and frequently pinnate. The leaves are also perfect as to position, being no longer mere radical leaves, but situated upon the stalk, even on its apical extremity.
1635. The ramification gradually emerges into view, where forsooth its occurrence is possible in the shaft-plants, or in the inflorescence. It is always multiple in character, mostly spadici-, muscari-, and paniculiform.
1636. As regards the blossom the ovarium is most perfectly evolved, because it is developed out of the stalk; it is ternary and becomes elevated into a fruit with few seeds.
1637. The corollæ are frequently stunted, yet regular and 2 × 3ary, though insignificant on account of the preponderance in size of the fruit.
1638. In this class we meet with the first true or genuine fruits; because in it for the first time the three anatomical systems are completely separated.
1639. Plants having a ligneous stalk, free and many-ribbed leaves, ramified inflorescence and ternary fleshy fruits, are the _Palmaceæ_. The Palmaceæ have a woody, very hollow stem, with many-ribbed, divided, and often pinnate leaves; a muscariform inflorescence lodged in spathes, sexanary corollæ with nuts, berries, or drupes. With the palms are associated the Typhaceæ, Aroideæ, Piperales, Pandanales, Dioscoreæ, Smilaceæ, Asparagi, Convallariæ, and Bromeliæ; for their stalks are mostly woody, the leaves broad, and placed upon the stalk, the corollæ stunted, while, on the contrary, the ovarium is carpoidal or fruit-like. The five orders may be disposed as follows:
1640. Order I. _Palmaceæ parenchymatosæ._ _Cynomoridæ_, _Typhaceæ_, _Aroideæ_.
1641. Order II. _P. thecales._ _Saurureæ_, _Piperaceæ_, _Pandanaceæ_.
1642. Order III. _P. axonales._ _Dioscoreæ_, _Smilaceæ_, _Parideæ_.
1643. Order IV. _P. florales._ _Asparagoideæ_, _Convallariæ_, _Bromeliæ_.
1644. Order V. _P. carpales._ _Palms._
1645. The plants of the first order are very imperfect herbs with spadices. Those of the second have mostly a ligneous, nodose stalk, with one-seeded fruits in spadices, without corollæ. The third have separated corollæ disposed in an open form of inflorescence. The fourth have perfect sexanary corollæ, with a frequently woody stalk and ternary, many-seeded berries. The fifth order consists of trees having large leaves, muscariform spadices, and perfect fruits, nuts, plums, and berries, ternary and one-seeded. They divide into sixteen families. (Vid. Tab. B.)
_Third Province._
RETICULAR-LEAVED PLANTS--DICOTYLEDONES.
1646. With the separation of the stock or trunk into root, stalk and leaf, the latter organ attains its perfection; it becomes a reticular leaf--the plants possessing it being called _reticular_.
1647. The retinerved or reticular leaf is, however, only the result of a modified organization in the stalk, and indicates a ramification or foliiform arrangement of the tracheæ in the stem. The foliiform arrangement of the tracheæ in the stalk is their circular disposition. These plants have _woody zones_. Through this zone or ring of wood first originates the perfect separation into wood, liber and bark, whereof each formerly occupied the whole stalk or stem.
1648. The stalk is no longer a shaft or scape, but it divides into branches and twigs--_ramular plants_.
1649. The reticular-veined are ramular leaves, and are no longer therefore spathiform but petiolated--_petiolated leaves_. It is only at the root that spathose leaves may occur, and this only in the plants of the inferior classes.
1650. With the disappearance of the spathose leaves, and the appearance of the ramules, the nodes and bulbs also disappear.
1651. The blossoms stand no longer upon a radical peduncle or stipes, but upon ramules; in other words, upon a plant, which again stands upon another plant, namely, the stalk.
1652. As all the higher separations of leaves here occur, so also does the flower obtain its higher amount of separation; it becomes quinary--_pentaschematose plants_. The ovarium passes through all numerical conditions, being 1, 2, 3, 4, 5, and polycarpellar. In like manner all the forms of ovaria and fruits occur in the present class, such as caryopsis, follicle, legumen, siliqua, capsule, nut, plum, berry and apple.
1653. As the seed is a leaf-formation, so must it resemble the reticular leaf. But reticular leaves are not spathes or simple tubes, but ramified or separated ribs. The seed has therefore several leaves, and two indeed for the first time, which are called seed-lobes. These plants are therefore styled _Dicotyledones_.
DIVISION.
1654. The Dicotyledones are, in the first place, empirically divisible into apetalous, monopetalous and polypetalous, or into plants with calycine, tubular and petalous corollæ.
1655. It might be believed that the Apetalæ were, without further trouble, the lowest in point of rank; but, when closely considered, they appear as Polypetalæ with stunted corolla-petals, and are obviously allied to the Rosaceæ. Moreover, they all bear nuts, a fact which occurs in no other class, and they must be therefore placed among the fruit-bearing plants. Since, however, they are epigynous and perigynous, so must the other polypetalous Perigynæ enter into proximity with them, and in like manner forsooth come among the Fruit-plants.
1656. Thus the Dicotyledones separate into the Monopetalæ, hypogynous Polypetalæ, and perigynous Polypetalæ, along with the Apetalæ. Viewed in a scientific light, they separate according to the principal members of the plant into three districts, Axis-, Flower-, and Fruit-plants.
FIRST CIRCLE.
_Stem-plants--Monopetalæ._
1657. The Monopetalæ or Tubulifloræ are the lowest in rank, and must therefore take the place here assigned them. They are still spathose corollæ. Among them are found for the most part only cariopsides and membranous capsules, rarely fruits. They are usually, too, merely herbs, rarely bushes, and still more rarely trees.
1658. They divide into Epigynes, Peri-and Hypogynes, of which the former are the lowest, the latter the highest; for in that which is left similar the coalition is an inferior sign.
1659. Their essential or typical character does not, however, reside in the blossom, but in the trunk, and that indeed in the root, stalk, and foliage. The question therefore may now be asked, whether in the Epigynes the root, in the Perigynes the stalk, and in the Hypogynes the foliage, be the principal organ.
_CLASS VII._
_Root-plants._
1660. That the Tubulifloræ with superior blossoms and fleshy root are radical plants, admits of being easily demonstrated. The preponderance of the root is evident from its size, its quantity of contained sap, or special chemical ingredients. A root that is rich in sap, and much denser than the stalk, is called a turnip. These plants are thus _turnip-plants_.
1661. Among the Monopetalæ, however, there are napiform roots only in the _Epigynes_, namely, the Syngenesia or salad-plants, and among some _Perigynes_, namely, the Campanuleæ, as well as the Cucurbitaceæ. The Syngenesia are consequently the radical plants. The roots of the Scorzoneræ, Pastinaceæ, Cichoraceæ, and Tussilago or coltsfoot, &c., belong to this class.
1662. That the Scabiosæ and Valerianeæ are directly related to the Syngenesia is likewise indicated by their roots. Unto these succeed in point of structure the Campanulaceæ and the Cucurbitaceæ, which have frequently, too, napiform or turnip-shaped roots.
1663. The number of the Syngenesia is so great that they fill up all the orders of the trunk. In accordance with their whole structure they are obviously the lowest, the stalk being for the most part herbaceous and placed within a circle of radical leaves, but being itself provided with few, imperfect, and scarcely ever pinnate leaves; they have moreover numerous stunted blossoms that are connate with the single or solitary seed, and crowded together, like spadices, grass-spikes, or the fungal pilei, upon a carpoclinium or receptacle.
1664. They are a repetition of the fungi and grasses; of the former in their fleshy root and inflorescence, of the latter likewise in the inflorescence and in the spathoidal root-leaves; above all, in the single large seed, confluent with the ovarium and calyx. The principle of their division must, where it is possible, and for obvious reasons, be drawn from the organs of the trunk.
1665. First order. _Radicariæ parenchymatosæ._--Syngenesia, having radical leaves and uniform florets, tubular or wholly stunted ligular florets--_Cichoraceæ_ and _Thistles_.
1666. Second order. _P. vaginatæ._--Syngenesia, with opposite leaves and different kinds of florets, such as radiated, partly lingual and partly tubular corollæ--_Sunflowers_, _Silphieæ_.
1667. Third order. _R. axonales._--Syngenesia, with alternate leaves and diversified florets--_Anthemideæ_, _Senecionidæ_, _Astereæ_.
1668. Fourth order. _R. florales._--Here the ovarium is no longer dense and confluent with the calyx, and it begins to become trilocular--_Scabiosæ_, _Valerianeæ_, and _Campanulæ_.
1669. Fifth order. _R. fructuariæ._--Here a perfect fruit is developed, which is connate with the calyx--_Asarideæ_, _Passifloreæ_, and _Cucurbitaceæ_. They have apple-like 3-5 ry fruits, and many of them have napiform roots, e. g. the Gichtrüben. They divide into sixteen families. (Vid. Tab. B.)
_CLASS VIII._
_Stalk-plants._
1670. Plants with a predominating development of stalk, leaves narrow, mostly opposite, quaternary corollæ upon the calyx, ovarium multilocular, and containing few seeds.
1671. Here everything, both root and leaf, must be stalk-like in character; the stalk is therefore woody, the root fibrous, the foliage twig-like or narrow, like needles.
1672. This structure is chiefly found in the _Heaths_ and _Stellatæ_. The stalk is mostly woody; the foliage either aciculiform or leathery in texture, and never pinnated. The leaves are either arranged in whorls or opposite, a position which indicates a lower grade of development. Moreover, they are related to the plants of the preceding class; they are either _Epigynous_ or _Perigynous_. The corolla and ovarium follow the opposite position of the leaves; the former being quadripartite, the latter bi-and quaternary. Most of them grow in hot countries upon dry ground, and their virtues reside in the stalk, e. g. the Peruvian bark.
1673. The Stellatæ or Rubiaceæ are without doubt the lowest, because they are epigynous and have a quadri-petalous corolla, with a binary and frequently only follicular ovarium.
1674. First order. _Cauliariæ parenchymatosæ._ The _Stellatæ_ proper along with the _Coffeaceæ_, all of them being two-seeded.
1675. Second order. _C. vaginatæ._ The Rubiaceæ with bilocular many-seeded capsules--_Rondeletiæ_ and _Cinchoneæ_.
1676. Third order. _C. axonales._ _Rubiaceæ_ with fruits--_Guettardidæ_, _Hameliæ_, _Gardeniæ_.
1677. Fourth order. _C. florales._ Quadripetalar Perigynæ, with similar capsules or berries--_Epacrideæ_, _Vacciniaceæ_, _Ericaceæ_.
1678. Fifth order, _C. fructuariæ._ Quaternary Perigynes with fruits--_Myrobalaneæ_, _Olacineæ_, _Diospyroideæ_ and _Sapoteæ_. (For the arrangement of their families vid. Tab. B.)
_CLASS IX._
_Leaf-plants._
1679. Herbs having broad leaves, quinary hypogynous corollæ, and bilocular capsule. Here the whole stem has become leaf; all the parts are soft; they are _herbs_ in the propermost sense of the word.
1680. Here belong the _hypogynous_ Monopetalæ: _Primulaceæ_, _Personatæ_, _Solaneæ_, _Gentianeæ_, _Asclepiadæ_, _Cariceæ_, _Asperifoliæ_, _Sambuceæ_. The roots are fibrous; the stalk herbaceous, being wholly covered, and that indeed with large leaves; calyx and corolla quinquepartite, frequently bilabiate; the germen a bilocular membranous capsule, which seldom becomes fleshy, and contains few seeds. It is these plants which serve chiefly as food for cattle, and whose whole trunk is officinally known under the name of _herbage_; relations which express the leafy character.
1681. First order, _Foliariæ parenchymatosæ_. Herbs with bilocular capsules, in which there are many seeds placed upon a median cone. The capsule dehisces, while both carpels separate from each other--_Primulaceæ_, _Scrophulariæ_, _Solaneæ_.
1682. Second order, _F. vaginatæ_. For the most part herbaceous, their many-seeded carpels springing open at the dorsal suture--_Orobanchaceæ_, _Rhinanthaceæ_, and _Bignoniaceæ_.
1683. Third order, _F. axonales_. Regular quinary corollæ with seeds borne upon the margins of the two carpellar valves--_Gentianaceæ_, _Asclepiadaceæ_, _Jasmineæ_.
1684. Fourth order, _F. florales_. Few seeds in one capsule; the pistil becomes nut-like or trilocular--_Labiatæ_, _Polemoniaceæ_, _Convolvulaceæ_.
1685. Fifth order, _F. fructuariæ_. Herbs and shrubs with fruits; as nuts, plums and berries. For their sixteen families (vid. Tab. B.)
B. BLOSSOM-PLANTS.
1686. Flowers polypetalous.
SECOND CIRCLE.
_Flower-plants._
1687. Calyx, corolla, stamina and ovarium perfectly separated from each other--_Pedunculate corollæ_, _Stielblumen_ or _Hypogynes_. The blossom must be here developed in the most perfect manner; i. e. all its parts must be complete and separated from each other. This is the case only in the hypogynous Polypetalæ.
1688. The lowest organized kinds must, from being a repetition of them, remind us of the grasses and Syngenesia. They are therefore polycarpellar or multi-ovarial.--_Ranunculaceæ_, _Malvaceæ_, _Magnoliaceæ_.
1689. Unto these are allied those plants whose ovaria consist of several carpels, but that are connate with each other, and mutually separate for the first time with their maturation or decay, as in the _Rutaceæ_, _Polygaleæ_, _Malvaceæ_, _Aurantiacæ_, _Platanaceæ_, _Malpighieæ_, _Sapindeæ_.
1690. The highest are characterized by coalition of the carpels into a single ovarium with stunted dissepiments, and by corollæ that are well developed and distinguished for colour, delicacy, and magnitude--_Carnations_, _Violets_, _Cistaceæ_, _Siliquosæ_, _Poppies_, _Gamboge-trees_.
_CLASS X._
_Seed-plants._
1691. Plants having a preponderance of seed, that draws after it all the floral parts.
1692. The ovaria have become seed-like, have separated from each other, and inclose for the most part only a single seed.
1693. As in the grasses and Syngenesia many flowers are collected into a spike or upon a receptacle, so here are many carpels in a single corolla--_Ranunculaceæ_, _Geraniaceæ_, _Tiliaceæ_, _Malvaceæ_, _Magnoliaceæ_.
1694. The stamina are usually of indefinite number, and mostly connate.
1695. All forms of stalks are here met with; such as herbs, bushes, shrubs, and trees. All forms also of leaves; spathose leaves, petiolated leaves, simple and divided, yet rarely pinnated.
1696. The component parts are usually mucilage, as in the roots of the Syngenesia.
1697. They divide into two great groups, into the quinary and sexanary. Since among the quinary, herbaceous stalks with nodes and spathe-leaves, but capsules only, occur; they must be arranged in the lowest rank. The sexanary bear fruits.
1698. First order, _Seminariæ parenchymatosæ_. Herbs with nodes and spathose leaves, together with numerous, mostly one-seeded, carpels, attached in an irregular manner to a median columella--_Ranunculaceæ_ and _Geraniaceæ_.
1699. Second order, _S. vaginatæ._ Trees having many-seeded carpels, coalesced like the style, _Theaceæ_, _Tiliaceæ_, _Elæocarpaceæ_.
1700. Third order, _S. axonales._ Bushes and shrubs with free, mostly simple, leaves, ovaria, mostly one-seeded, and arranged in a circle around the mediate axis or columella; anthers bilocular--_Hermanniaceæ_, _Dombeyaceæ_, _Sterculiaceæ_, and _Büttneriaceæ_.
1701. Fourth order, _S. florales._ For the most part trees, having frequently divided leaves and similar ovaria, yet mostly many-seeded and connate--_Malvaceæ_ and _Bombaceæ_.
1702. Fifth order, _S. fructuariæ._ Corollæ mostly sexanary, ovaria in a circle without a columella--_Magnoliaceæ_, _Menispermaceæ_, _Dilleniaceæ_, _Anonaceæ_. They divide into sixteen families. (Vid. Tab. B.)
_CLASS XI._
_Ovarium-plants._
1703. Hypogynous Polypetalæ with perfect multilocular ovarium--_Polygaleæ_, _Melieæ_, _Aurantiaceæ_, _Platanaceæ_, _Malpighieæ_, _Sapindeæ_.
1704. While in the preceding class the number of the carpels was usually indeterminate; it is here limited to three and five. In the one, they usually stood around a middle columella or axis, in the other, they form a true capsule with perfect partition-walls or septa, and a single style. The number of the seeds is moderate, i. e. there are more than one, but they are easily counted. They are therefore of mediate size, having no kernels, as is the case in the nuts, but also no granules, as in the berries or poppy capsules. In the preceding class the fruits were rare; here they occur frequently in the upper orders. The number of the floral parts is, throughout the present class, five, that of the stamina five or ten, and they are seldom coalesced; the stalk also passes through all the stages of development from that of the herb through the shrub into the tree. The leaves are seldom spathe-like, but frequently coriaceous and aciculate, as in their predecessors the _Heaths_; many are pinnated.
1705. First order, _Capsulariæ parenchymatosæ_--_Rutaceæ_. Herbs and shrubs frequently provided with aciculate and coriaceous leaves; corollæ regular with ten stamina; ovarium consisting of five carpels, which separate when ripe, and contain few seeds--_Rutaceæ_, _Diosmaceæ_.
1706. Second order, _C. vaginatæ._ Shrubs and trees with similar corollæ and ovaria, which are nevertheless frequently separated and fleshy--_Quassiaceæ_, _Ochnaceæ_.
1707. Third order, _C. axonales._ Mostly shrubs and trees with irregular corollæ and bilocular ovarium--_Polygalaceæ_, _Vochysieæ_, _Pittosporeæ_.
1708. Fourth order, _C. florales._ Trees with a woody or berry-like ovarium, having several cells--_Cedreleæ_, _Melieæ_, _Aurantiaceæ_.
1709. Fifth order, _C. fructuariæ._ Trees; flowers quinary, ovarium mostly ternary, becoming a winged or fleshy fruit. (Their sixteen families probably pursue the order indicated at Tab. B.)
_CLASS XII._
_Corolla-plants._
1710. Corollæ stipaceous, having free stamina; ovaria with stunted septa, and numerous marginal seeds--_Carnations_, _Violets_, _Cistaceæ_, _Siliquosæ_, _Papaveraceæ_, _Guttiferæ_.
1711. The stalk passes through all the stages of formation, from the nodose herb unto the shrub and tree. The leaves occur likewise under all forms, modes, division, and arrangement. The plants of this class are found in all climates, and yield etherial or volatile and fatty oil with resins. They divide first of all into quinary and quaternary; the former being mostly herbs with a hollow capsule; the latter herbs, shrubs, and trees, with siliquæ or berries.
1712. Their strength or virtue resides in the flower, which, is therefore of large size, beautifully coloured, sweet-scented, is frequently appreciated and adopted as an ornament. On the contrary, the ovarium and seed are stunted. The former is a siliqua or hollow capsule, which therefore supports the numerous smaller seeds on the septum.
1713. First order, _Corollariæ parenchymatosæ_. Nodose herbs with spathose leaves, quinary corollæ, and ten stamina; many seeds upon a columellar or median placenta situated within a hollow capsule--_Portulaceæ_, _Carnations_.
1714. Second order, _C. vaginatæ_. Herbs, shrubs, and trees, with similar but mostly multi-staminal corollæ, and seeds on the capsular septa--Droseraceæ, _Hypericineæ_, _Violaceæ_, _Cistaceæ_, and _Bixeæ_.
1715. Third order, _C. axonales_. Herbs with quaternary corollæ and a siliqua--_Siliquosæ_.
1716. Fourth order, _C. florales_. Bushes and shrubs with quaternary corollæ and numerous stamina; ovarium a siliqua or multi-valvular hollow capsule--_Capparidæ_, _Berberideæ_, _Papaveraceæ_.
1717. Fifth order, _C. fructuariæ_. Trees with quaternary and quinary corollæ, numerous stamina, and a fruit--_Guttiferæ_. They divide into the usual sixteen families. (Vid. Tab. B.)
THIRD CIRCLE.
_Fruit-plants--Apetalæ, Perigynes._
1718. Stunted calycine corollæ, with nuts, plums, berries, or apples.
1719. They are perigynous Polypetalæ, and include the Apetalæ and Diclines.
1720. The nut consists of a large seed, connate with the woody ovarium, and frequently with the calyx.
1721. The plum is a legumen, between the coats of which fleshy matter has accumulated, and whose internal tunic or endocarp has become woody. The berry is a many-seeded hollow capsule, which, as well as the calyx, has become soft and succulent.
1722. The apple is an ovarium surrounded by a fleshy calyx.
_CLASS XIII._
_Nut-plants--Apetalæ, Diclines._
1723. Ovarium woody, and inclosing only one seed. Here belong the Apetalous and Diclinous Exogens.
1724. These plants repeat the fungi, grasses and Syngenesia, and have therefore imperfect corollæ, the calyx of which has alone remained, and usually closely surrounds the nut.
1725. The stalk is indeed usually woody; yet is still found to be also herbaceous and nodose with spathiform leaves. The leaves are simple, frequently needle-shaped or else arrested. The principal ingredients are starch, as in the Gramineæ and Syngenesia.
1726. The inflorescence is mostly amentaceous, as in the Agarics, Grasses and Syngenesia.
1727. They divide into androgynous and dioecious plants.
1728. First order, _Nucariæ parenchymatosæ_. Hermaphrodite herbs with nodes and spathose leaves, calyx green, superior, and quinquepartite with five opposite stamina; nut mostly triangular and utricular.
1729. Second order, _N. vaginatæ_. Form of vegetation pretty nearly as in preceding order, but the calyx is corolla-like, and the stamina mostly alternate--_Phytolacceæ_, _Illecebreæ_.
1730. Third order, _N. axonales_. Hermaphrodite, calyx corolla-like, superior and wholly quaternary; herbs and shrubs bearing nuts and plums--_Nyctagineæ_, _Daphneæ_, and _Santalaceæ_.
1731. Fourth order, _N. florales_. Trees, calyx corolla-like but inferior; capsules, plums and berries--_Proteaceæ_.
1732. Fifth order, _N. fructuariæ_. Diclines; herbs, shrubs and trees without corollæ, but with nuts or plums. (Their sixteen families stand in the order exhibited at Tab. B.)
_CLASS XIV._
_Plum-plants--Papilionaceæ._
1733. Polypetalous calycine corollæ, with a drupe or its fundamental form, the legumen. Here belong the papilionaceous plants, Rhamnaceæ and Terebinthaceæ.
1734. The stalk is frequently herbaceous with nodes; but mostly fruticose and arborescent.
1735. The leaves here attain their highest development, and are mostly pinnated, sometimes endowed with the power of independent motion.
1736. The corollæ are mostly irregular, quinary, arranged like pinnate leaves, with ten, rarely more, connate and free stamina.
1737. The ovarium is a single carpel, owing to the four others being arrested; it is usually compressed and bivalved, with few seeds; it is a legumen, frequently converted into a fleshy fruit.
1738. The Papilionaceæ are so rich in number that they include all the orders of the trunk, and even transcend or exceed its limits; their allied families are the Rhamnaceæ and Terebinthaceæ with fleshy fruits.
1739. First order, _Drupariæ parenchymatosæ_. Papilionaceæ with herbaceous, nodose stalk and pinnated leaves; corolla-petals and single stamen free; seed-lobes thin--Hedysareæ, _Astragaleæ_, _Glycineæ_.
1740. Second order, _D. vaginatæ_. Herbs and shrubs with ternary or tendrilless leaves; the corolla-petals or stamina frequently confluent--_Trifoliaceæ_, _Genisteæ_, _Galegeæ_.
1741. Third order, _D. axonales_. Bushes, shrubs, or trees, frequently training with pinnate leaves and tendrils; calyx large, seed-lobes thick--_Vicieæ_, _Beans_, _Dalbergiæ_.
1742. Fourth order, _D. florales_. Shrubs and trees, with tolerably regular flowers and separated stamina; legumens with frequently transverse septa, embryo straight--_Geoffroyæ_, _Swartzieæ_, _Detarium_, _Mimosæ_, and _Cassiæ_.
1743. Fifth order, _D. fructuariæ_. Regular flowers with separate stamina and multilocular plums--_Stackhouseæ_, _Empetreæ_, _Celastrineæ_, _Rhamneæ_, and _Terebintheæ_. (For their sixteen families vid. Tab. B.)
_CLASS XV._
_Berry-plants_--_Umbelliferæ_, &c.
1744. Ovarial and calycine corolla with a single perfectly soft fruit or berry, on which are five corolla-petals, with a moderate number of stamina. This fruit is wholly edible, and has only one or two styles; stalk and leaves pass through all the stages of formation.
1745. They divide first of all into two groups, having few or many stamina; of these the fruit of one is dry, of the other fleshy. The dry fruits are also perfectly edible, as the Caraway-seeds.
1746. First order, _Baccariæ parenchymatosæ_. Epigynous; nodose herbs with two seeds in the calyx; only five stamina--_Umbellatæ_.
1747. Second order, _B. vaginatæ_. Mostly shrubs with quinary corollæ and bi-or quinquelocular berries--_Mistletoes_, _Elders_, _Araliaceæ_, _Vines_.
1748. Third order, _B. axonales_. Bushes and shrubs with quaternary corollæ, only one style and one multilocular many-seeded capsule--_Epilobeæ_, _Salicariæ_.
1749. Fourth order, _B. florales_. Mostly shrubs with quinary corollæ and manifold stamina; capsule or berry multilocular--_Melastomaceæ_.
1750. Fifth order, _B. fructuariæ_. Trees with many, frequently fasciculated, stamina; fruit multilocular and many-seeded--_Myrtaceæ_. These plants likewise divide into sixteen families. (Vid. Tab. B.)
_CLASS XVI._
_Apple-plants_--_Rosaceæ_.
1751. The fruit is an apple, i. e. several carpels, containing but few seeds, adhere in one calyx, upon which there are five corolla-petals, with four to six times as many stamina. Herbs, shrubs, and trees with different kinds of leaves are here met with; the corollæ are mostly small and perigynous, the style separated, and thus polycarpal. Perigynous Polycarpæ. They grow, dispersed over the whole earth, in dry situations; several of them yield edible fruits, and are pretty generally cultivated.
1752. The apples are without doubt the most perfect fruit, as well in reference to their structure as their chemical ingredients. The apple consists of all the parts of the blossom: seed, capsule, and fleshy calyx, and is besides polycarpellar, i. e. composed of separate carpels. Its edible substance or flesh is not simply a sweet-meat, but a true aliment, which admits of being eaten after it has been kept fresh for a year, of being dried, exported, or cooked as a kitchen vegetable; in cases of exigency too it occurs in such abundance that the whole human race might live upon it, which cannot be said of any other fruit. The apple quenches at the same time the thirst, and thus supplies also the place of drink. All the other fruits are either a dainty relish only against thirst, or a simple amylaceous medium of nutrition. However, most of the plants belonging to this class produce only dry capsules and calyces.
1753. They divide into two groups, having few or many stamina.
First order, _Pomariæ parenchymatosæ_. Herbs with few stamina and five or more carpels, as the _Crassulaceæ_ and _Mesembryanthemaceæ_.
Second order, _P. vaginatæ_. Shrubs with few stamina, only two to three carpels and few seeds.--_Tamaricaceæ_, _Bruniaceæ._
VEGETABLE SYSTEM Table B +==========================================================================+ | FIRST PROVINCE. | | | | HISTOPHYTA--ACOTYLEDONES. | +--------------------------------------------------------------------------+ | | +====================+=================+==================+================+ | | | | | | | CLASS I. | CLASS II. | CLASS III. | | | | | | | ORGANS. | Cellulares. | Vasculares. | Tracheales. | | | _Fungi._ | _Mosses._ | _Ferns._ | | | | | | | ORD. I. Parenchyma.| Roste. | Schleim-Moose. | Wasser-Farren. | | | | | | | F. 1. Cells. | Entophyta. | Diatomaceæ. | Marsileaceæ. | | 2. Ducts. | Epiphyta. | Nostochineæ. | Pilulariæ. | | 3. Tracheæ. | Tuberculariæ. | Batrachospermeæ.| Equisetaceæ. | | | | | | | O. II. Sheaths. | Schimmel. | Wasserfäden. | Kugelfarren. | | | | | | | F. 4. Bark. | Mucorini. | Confervaceæ. | Lycopodiaceæ. | | 5. Liber. | Mucedines. | Ulvaceæ. | Ophioglosseæ. | | 6. Wood. | Byssaceæ. | Leptomiteæ. | Danæaceæ. | | | | | | | O. III. Axis. | Balgpilze. | Fuci. | Ringfarren. | | | | | | | F. 7. Root. | Trichodermaceæ.| Cerameæ. | Parkereæ. | | 8. Stalk. | Trichiaceæ. | Floridæ. | Cyatheæ. | | 9. Leaves. | Lycopodineæ. | Fucoideæ. | Polypodiaceæ. | | | | | | | O. IV. Blossom. | Kernpilze. | Lichens. | Fluvialen. | | | | | | | F. 10. Seed. | Cytisporeæ. | Coniothalameæ. | Naiadeæ. | | 11. Ovarium. | Phacidiaceæ. | Gasterothalameæ.| Podostemeæ. | | 12. Corolla. | Sphæriaceæ. | Hymenothalameæ. | Ceratophylleæ.| | | | | | | O. V. Fruit. | Fleischpilze. | Mosses. | Nadelholz. | | | | | | | F. 13. Nut. | Tremellini. | Marchantiaceæ. | Abieteæ. | | 14. Plum. | Cupulati. | Ricciaceæ. | Taxaceæ. | | 15. Berry. | Clavati. | Jungermanniaceæ.| Cupresseæ. | | 16. Apple. | Pileati. | Bryaceæ. | Cycadeæ. | +--------------------+-----------------+------------------+----------------+
+====================+=====================================================+ | | SECOND PROVINCE. | | | | | | THECOPHYTA--MONOCOTYLEDONES. | +--------------------+-----------------------------------------------------+ | | | +====================+==================+===================+==============+ | | | | | | | CLASS IV. | CLASS V. | CLASS VI. | | | | | | | ORGANS. | Corticales. | Liber-plants. | Wood-plants.| | | _Gramineæ._ | _Liliaceæ._ | _Palmaceæ._ | | | | | | | ORD. I. Parenchyma.| Spiked Grasses. | Staub-Orchiden. | Rohrkolben. | | | | | | | F. 1. Cells. | Secale. | Neottiæ. | Cynomoria. | | 2. Ducts. | Phleum. | Arethuseæ. | Typhaceæ. | | 3. Tracheæ. | Festuca. | Ophrydeæ. | Aroideæ. | | | | | | | O. II. Sheaths. | Panicled Grasses.| Granular Orchideæ.| Piperaceæ. | | | | | | | F. 4. Bark. | Bulrushes. | Malaxides. | Saurureæ. | | 5. Liber. | Milium. | Epidendra. | Piperaceæ. | | 6. Wood. | Arundinaceæ. | Vanillæ. | Pandanac | | | | | | | O. III. Axis. | Reed-Grasses. | Aromatic plants. | Sassaparillæ.| | | | | | | F. 7. Root. | Sedges. | Amomeæ. | Dioscoreæ. | | 8. Stalk. | Junceæ. | Canneæ. | Smilaceæ. | | 9. Leaves. | Ruscus. | Musaceæ. | Parideæ. | | | | | | | O. IV. Blossom. | Juncineæ. | Schwerdel. | Asparagoidæ. | | | | | | | F. 10. Seed. | Restiaceæ. | Hæmodoraceæ. | Asparagoidæ.| | 11. Ovarium. | Serpeæ. | Irideæ. | Convallariæ.| | 12. Corolla. | Commelyneæ. | Narcissi. | Bromeliæ. | | | | | | | O. V. Fruit. | Seeroseæ. | Liliaceæ. | Palms. | | | | | | | F. 13. Nut. | Alismaceæ. | Colchicaceæ. | Calamariæ. | | 14. Plum. | Hydrocharideæ. | Asphodeleæ. | Cocoeæ. | | 15. Berry. | Hydropeltideæ. | Alliaceæ. | Phoeniceæ.| | 16. Apple. | Nymphæacæ. | Tulipaceæ. | Borasseæ. | +--------------------+------------------+-------------------+--------------+
VEGETABLE SYSTEM. [TABLE B, p. 316.]
+=====================================================+ | | | THIRD PROVINCE. ARTHROPHYTA--DICOTYLEDONES. | | | +-----------------------------------------------------+ | CIRCLE I. _Stem-plants--Monopetalæ._ | +==================+================+=================+ | | | | | CLASS VII. | CLASS VIII. | CLASS IX. | | | | | | Root-plants. | Stalk-plants. | Leaf-plants. | | _Epigynæ._ | _Perigynæ._ | _Hypogynæ._ | | | | | | Root-leaved. | Stellatæ. | Personatæ. | | | | | | Cichoraceæ. | Galiaceæ. | Primulaceæ. | | Carduraceæ. | Spermacocidæ. | Scrophulariæ. | | Mutisiæ. | Coffeaceæ. | Solanaceæ. | | | | | | Opposite-leaved. | Cinchonaceæ. | Bignoniaceæ. | | | | | | Eupatorium? | Hedyotidæ. | Orobancheæ. | | Helianthus. | Rondeletiæ. | Rhinantheæ. | | Silphieæ. | Cinchoneæ. | Bignoniæ. | | | | | | Alternifoliar. | Hameliaceæ. | Contortæ. | | | | | | Senecionidæ. | Guettardidæ. | Gentianeæ. | | Astereæ. | Hameliæ. | Asclepiadeæ. | | Vernoniæ. | Gardeniæ. | Jasmineæ. | | | | | | Aggregatæ. | Ericaceæ. | Tetraspermeæ. | | | | | | Scabiosæ. | Epacrideæ. | Labiatæ. | | Lobeliæ. | Myrtilli. | Polemoniaceæ. | | Campanulæ. | Heaths. | Convolvulaceæ. | | | | | | Sicuonoidæ. | Diospyraceæ. | Pyrenaceæ. | | | | | | Asaridæ. | Myrobalaneæ. | Asperifoliæ. | | Loaseæ. | Olacineæ. | Verbenaceæ. | | Passifloreæ. | Diospyros. | Sambuceæ. | | Cucurbitaceæ. | Sapotæ. | Myrsineæ. | +------------------+----------------+-----------------+
+===================================================+ | | | THIRD PROVINCE. ARTHROPHYTA--DICOTYLEDONES. | | | +---------------------------------------------------+ | CIRCLE II. _Flower-plants--Hypogynous Polypetalæ._| +===============+=================+=================+ | | | | | CLASS X. | CLASS XI. | CLASS XII. | | | | | | Seed-plants. | Ovarium-plants. | Corolla-plants. | | _Polycarpæ._ | _Capsuliferæ._ | _Siliquosæ._ | | | | | | Ranunculaceæ. | Rutaceæ. | Caryophyllaceæ. | | | | | | Ranunculi. | Ruteæ. | Portulaceæ. | | Helleboreæ. | Diosmeæ. | Spergula. | | Geraniaceæ. | Zygophylleæ. | Carnations. | | | | | | Tiliaceæ. | Ochnaceæ. | Violaceæ. | | | | | | Theaceæ. | Xanthoxyleæ. | Droseraceæ. | | Lime-trees. | Ochneæ. | Violets. | | Elæocarpeæ. | Quassieæ. | Cistaceæ. | | | | | | Sterculiaceæ. | Polygalaceæ. | Cruciferæ. | | | | | | Hermanneæ. | Polygaleæ. | Radishes. | | Dombeyaceæ. | Vochysieæ. | Cresses. | | Sterculeæ. | Pittosporeæ. | Cabbages. | | | | | | Malvaceæ. | Meliaceæ. | Papaveraceæ. | | | | | | Malveæ. | Cedreleæ. | Capers. | | Hibisceæ. | Melieæ. | Berberideæ. | | Bombaceæ. | Aurantiaceæ. | Poppies. | | | | | | Magnoliaceæ. | Malpighiaceæ. | Guttiferæ. | | | | | | Magnolieæ. | Platanaceæ. | Dipterocarpeæ. | | Menispermeæ. | Malpighieæ. | Calophylleæ. | | Dilleneæ. | Hippocrateæ. | Marcgraviaceæ. | | Anoneæ. | Sapindeæ. | Garcinieæ. | +---------------+-----------------+-----------------+
+===================================================================+ | | | THIRD PROVINCE. ARTHROPHYTA--DICOTYLEDONES. | | | +-------------------------------------------------------------------+ | CIRCLE III. _Fruit-plants--Apetalæ_ and _perigynous Polypetalæ_.| +==============+===============+================+===================+ | | | | | | CLASS XIII. | CLASS XIV. | CLASS XV. | CLASS XVI. | | | | | | | Nut-plants. | Plum-plants. | Berry-plants. | Apple-plants. | | _Nucariæ._ | _Drupaceæ._ | _Baccariæ._ | _Pomaceæ._ | | | | | | | Oleraceæ. | Astragalaceæ. | Umbellatæ. | Sempervivæ. | | | | | | | Sclerantheæ.| Hedysareæ. | Saniculidæ. | Galacineæ. | | Atriplices. | Astragaleæ. | Carrots. | Crassulaceæ. | | Amaranths. | Glycineæ. | Caraways. | Mesembryanthemeæ.| | | | | | | Polygonaceæ. | Trifoliaceæ. | Caprifoliaceæ. | Tamariscineæ. | | | | | | | Plantains. | Trifolieæ. | Mistletoes. | Tamarisks. | | Phytolacceæ.| Genisteæ. | Sambuceæ. | Buniaceæ. | | Illecebreæ. | Galegeæ. | Vines. | Hamamelideæ. | | | | | | | Thymelaceæ. | Phaseolaceæ. | Lythraceæ. | Saxifrageæ. | | | | | | | Nyctagineæ. | Vetches. | Trapeæ. | Saxifrages. | | Daphnoideæ. | Dalbergieæ. | Epilobeæ. | Cunoniaceæ. | | Santalaceæ. | Sophoreæ. | Salicariæ. | Lilacs. | | | | | | | Lauraceæ. | Mimosaceæ. | Melastomaceæ. | Rosaceæ. | | | | | | | Proteaceæ. | Detarieæ. | Rhexieæ. | Potentillæ. | | Aquilarineæ.| Mimoseæ. | Melastomeæ. | Neuradeæ. | | Laurels. | Cassieæ. | Grossularieæ. | Sanguisorbeæ. | | | | | | | Diclinisten. | Terebinthaceæ.| Myrtaceæ. | Fruit-trees. | | | | | | | Amentaceæ. | Empetreæ. | Lecythideæ. | Pomegranates. | | Nettles. | Celastrineæ. | Barringtonieæ.| Plums. | | Artocarpeæ. | Rhamneæ. | Leptospermeæ. | Medlars. | | Euphorbicæ. | Terebintheæ. | Myrteæ. | Apples. | +--------------+---------------+----------------+-------------------+
Third order, _P. axonales_. Herbs, shrubs, and trees, with similar blossoms, but many seeds.--_Saxifrageæ_, _Cunoniaceæ_, _Hortensiæ_ and _Lilacs_.
Fourth order, _P. florales_. Herbs and shrubs, with many stamina and follicles.--_Rosaceæ_.
Fifth order, _P. fructuariæ_. Trees with multistaminal-corollæ and fleshy fruits.--_Monimieæ_, _Pomegranates_, _Plums_, _Medlars_, _Apples_. (For their sixteen families vid. Tab. B.)
1754. In order to prove that each class of vegetables commences from below, and that all ascend in a parallel series beside each other, it is requisite only to place them in a tabular form. Further details upon this subject are to be found in my earlier works upon Natural History. The adjoining Table may here suffice. I place the families according to my "Lehrbuch der Naturgeschichte," although I am very well aware that all do not stand in the right place. That indeed no one will expect.
THIRD KINGDOM.
ANIMAL KINGDOM.
1755. The animal kingdom is the individual development of all four elements.
ZOOSOPHY,
1756. Is the development of the animal kingdom in consciousness. The repetition of the animal creation is spiritually divisible into Anatomy (Zoogeny), Physiology (Zoonomy), and Zoology.
I.--_ZOOGENY_.
1757. Zoogeny represents the idea of the animal or the developmental history of the individual animal.
IRRITABILITY OF THE BLOSSOM.
1758. The Highest to which the vegetable kingdom could attain was the blossom; and of this the sexual parts are the completion. At the instant when the sex originated, the vegetable functions became of a nobler character; for the sexual organs are only the inferior organs refined or purified by light. The electrical and chemical process of the vegetable body are again represented, but in a spiritual manner, in the blossom. The functions of the fruit were none other than those of the elevated chemism; they were only the nobler processes of digestion and nutrition. As their purest expression of life, and that which has been produced simply by their co-operation, is the motion which resides in cellular tissue, so also was it this only, which obtained a preponderance in the fruit, and that indeed at the cost of the material processes. The germen obtained a kind of motion; yet this appears to be still imparted by material processes. In the corolla, however, this expression of life attained to completion. It is no longer simple nutrition or accumulation of sap that moves the stamina upon the female stigma, but a purely polar act; the Immaterial, the Spiritual produces the phenomena of life. These movements of copulation are by no means a coalescence, are not a nutritive act, nor the result of mechanical decissation, as is the case in many capsules; but true elevated vital actions. The parts reassume after the motion their first or original position, a feat which is performed by no capsule upon its dehiscence. Of these movements, those of the leaves in the sensitive plants, e. g. Hedysarum gyrans, are the antetypes or prefigurations. There consequently originates with the highest development of the light-organs in the plant, a motion independent of the material processes, and of the terrestrial elements.
1759. A motion, that is liberated from the terrestrial elements, is free from their mechanism; it simply obeys the nature of the æther, which is of a spiritual, or voluntary, kind.
1760. The essence of volition or free-will does not reside, in a physical sense, in the consciousness of the action, but in the autocrasy; or in the ability to perform an action, without external or terrestrial influence. The æther-actions have originated from special polarity. Independent movements must therefore be such as have been produced simply by polarity, apart from material intrusion.
1761. The ability or power upon the part of organic bodies to apprehend polar excitation, to move themselves simply by its means, and again to restore themselves to their former state, without regard being paid to a material process, I call _Irritability_. That organ is irritable which can move itself without any other object than that of automacy or self-motion.
1762. Irritability belongs to the plant, but is only of that kind where the perception cannot express itself otherwise than by direct motion. In the sexual parts and probably in the highest leaf-formation, the plant is elevated to irritability; unto motion through simple perception, unto motion without design, to motion from mere lust or desire. The highest spiritual operation, of which the plant is capable, is irritability. But as everything which has attained its Highest, stands at the end of its development, so also has the plant terminated, when it has once exercised its power of irritability through copulation.
SEXUAL MOTION.
1763. All the irritable motion of the vegetable may be confined or reduced to the movements of the stamen-filaments, the other movements being merely precursors of these. What, therefore, the stamina would achieve by their motion, that the irritability does in a general point of view. The motion of the stamina is directed merely upon the stigma, in order to impart the male pollen to the female body; and thus, simply to evoke the spiritual tension, which resides originally in the male semen, as in light-æther from the dead mass, but that originally dwells in the female seed as in the dark mass of earth.
1764. Now, since the stigma bears simply relation to the contents of the ovary, and conveys everything to this, and thus, to a female utricle, which is the middle of the plant, or its body proper; so in the motion of the male organs, the conatus or effort upon their part to introduce an elemental matter, or rather its spirit into this utricle, or this body, is rendered manifest or revealed. The highest Spiritual of the plant is accordingly not a mere motion in the general sense; but one that is definite and wholly special--_a motion of Ingestion_. The direction or design of the first _independent_ motion is therefore _Ingestion_; this again not being of a general, but a wholly definite import, namely, an ingestion of the _male_ organ into the _female_.
1765. The Male is, however, characterized by its self-substantial polarity, and by its self-intrinsic life; the Female by the want of polarity, by a heterodependent life. The act of ingestion thus depends upon polarization, upon evocation and maintenance of an independent life. The fruit is vivified by impregnation, ay, for the first time then obtains life; the female becomes self-active through the vital spirit received from the male; the body is kept alive by, and only by, ingestion. The act of ingestion is the act that tends unto the self-substantial, independent life.
1766. The blossom dies, so soon as, by ingestion, it has attained this independent life. If we assume that it could not die, but retain the life that was momentarily won for several instants; then this would happen only by repetition of the first act, whereby it had obtained in an instant a self-substantial life; and thus by repetition of the ingestion. By ceaseless ingestion only can the blossom gain a lasting, self-substantial life of motion.
1767. But such a blossom, when self-dependently subsisting, could not continue in further union with the vegetable stock or trunk, for this is no longer requisite for its life's support; through the first act of vivification, through the once sprinkled pollen, it virtually becomes detached and falls as fruit to the ground; as a fruit certainly, or as a female body, unto which is wanting the _continued_ excitation produced by the male coition. A fruit thus detached or fallen off, and retaining the male filaments, which ceaselessly exercise the function of ingestion, will be of necessity engaged in constant motion; will be a blossom, that incessantly practises copulation.
1768. As in this blossom the motion of ingestion is that alone which sustains it, and nothing more can flow to it from a stem; so also will this blossom be occupied in constant motion; and it consequently comes to pass that the action, which broke forth at the last and instantaneously in the plant, being thus the highest or most individualized, is here the first, inferior, and most general action, or one that lies at the foundation of all other processes. The free blossom is naught but movement of ingestion.
1769. The blossom, however, concentrates in itself all the lower vegetable processes, is itself naught but the aggregate of such processes repeated in the body of the light; thus the blossom, having been set free, is a vesicle of ingestion endowed with all terrestrial functions.
ANIMAL FORMATION.
1770. The vegetable blossom loses its definition as plant, so soon as it has acquired self-substantial life; it loses its definition, because as blossom, it simply lives in light, while the plant must half dwell in darkness; it loses it, because the copulating motion can be frequently self-repeated.
1771. The self-moveable or automatic blossom has consequently passed over into a new kingdom, or into one whose very definition is the self-substantial motion.
1772. A blossom, which, when separated from the stem, maintains by its own motion the galvanic process or the life within itself, and gets its process of polarization, not from a body lying external to or coherent with it (as is the vegetable stem); but only from itself--such blossom is an _Animal_. An animal is blossom without stem or a flower, which of itself produces its stem, this being the reverse of what takes place in the plant. The essence of the animal consists in the maintenance through its own motion of the galvanic vital process. It has been already shown above, in speaking "quoad" the distinction of the organic essences, that the exclusive or only valid difference between plant and animal, was motion arising in the latter apart from any external stimulus. We have now been brought by quite another route to the same result.
1773. If the animal is the floral vesicle living from or by itself, it can no longer lie fettered like the plant between two elements; it must be nominally free from the chains of darkness, and thus from the earth. No animal is so joined by growth with the earth, that like as in a plant this should be a co-operating pole in its processes. No animal must co-exist in two elements like the plant, but it has all elements in itself, like the blossom includes all vegetable parts. It may be said that the plant has been immersed in the earth, water, and air; these three elements having, on the contrary, been immersed in the animal. The animal is in respect to them the continent, the planet; they, however, are the continent in respect to the plant. Thus the relations to the world are completely reversed in both.
1774. An animal is a floral vesicle freely separated from the earth, and living alone through its own motion in the water and air. And here it is not locomotion that is treated of, since this by no means belongs to the animal's essence. Yet on this account the poor oyster was formerly adduced as an argument against this animal character, but unjustly; for would a man frozen up in an ice-block lose his character as an animal? The oyster opens its shell and shuts it too, as well as the crocodile opens and shuts its jaws.
ANIMAL SIGNIFICATION.
We will now proceed to knit or associate with this genetic or physiological, the physio-philosophical mode of development.
1775. Every Organic originates from a mucus-point. If this mucus-point occur in the darkness, it thus becomes a terrestrial organism, a plant; if it enter into the light, which is only possible in the water and in air, it thus becomes a solar organism, independent of the planet, self-moving around itself like the sun, an animal.
1776. An animal is a light-mucus-vesicle, a plant, so far as the root is concerned, a darkness-mucus-vesicle; but it works its way unto air and light, and becomes a light-mucus-vesicle in the blossom.
1777. A free blossom is consequently to be philosophically regarded as equivalent to the primary mucus-vesicle, which has at once developed itself in the water. Now, such an aqueo-mucus-vesicle is directly that which the blossom can first become through a series of developments, and by divestures of the Dark.
1778. The plant is an animal retarded by the darkness; the animal is a plant blooming directly through the light, and devoid of root.
1779. The animal has been posited as a light-or æther-Total upon the planet; the vegetable as a planetary Total in light.
1780. The animal is a whole solar system, the plant only a planet. The animal is therefore a whole universe, the plant only its half; the former is microcosm, the latter microplanet.
ANIMAL PLACE.
1781. No animal can become developed beneath the earth, or where it is absolutely dark and dry. None simply or solely in air. Water is the origin or source of all animals.
1782. They have originated upon the sea-shore, but not in the midst of the sea, nor in that of the land. The deluge cast up the first men. They were littoral inhabitants, and without doubt carnivorous, as savages still are. For whence could they have obtained also fruits, cabbage, and turnips?
SENSATION.
1783. In so far as the animal vesicle is a whole solar system, to it, the characters that transcend the plant, such as motion, belong.
1784. But motion is not that alone which is displayed in the floral cyst when it has become solar; but with it there is yet something higher bestowed.
1785. Like light or sun the vesicle has the principle of its polarization or determination in itself; and it is certainly itself which moves its organs in conformity with this self-determination; but it is withal in antagonism towards the elements, like the sun is towards the planets. Through this antagonism the sun is destined unto the development of light. Although the light is its own product, still it perceives the object toward which the æther-polarity is directed. This perception of the direction, whither motion should act by means of the central polarity, is in the animal called _Sensation_.
1786. Sensation is the relation of the Central to the Peripheric, of the sun to the planet; motion is the relation of the periphery to the centre, of the planet to the sun. The Animal emerges into being from the alternating play of the supremest antagonism in the heavenly bodies, the Vegetable from that of the terrestrial antagonisms.
1787. Emission of light is discharging or unloading of the sun achieved by the influence of the planets; sensation is unloading of the animal by objects, by the world, and by its own organs.
1788. Sensation is therefore a positing of _want_ in the animal. Through sensation nothing comes into, but rather something passes out of us.
1789. The animal vesicle is a sentient or feeling blossom.
SEXUAL ANIMAL.
1790. As the essence of the blossom consists in the sex, so indeed is the blossom none other than the sexual system; thus it may be said that the animal vesicle is none other than a _sentient Sexual cyst_. This discovery is of the highest importance for the whole of zoosophy.
1791. Two fundamental properties are originally combined in the animal, at the very instant when an animal can exist. There is never one without the other, never simply sensation, but also motion; but the latter is not simply this, but is at once also a _movement of copulation_. The animal is a sentient genital.
1792. Both fundamental properties are however subordinated to each other. The basis of the animal organism is the sexual system; with this the animal commences; what is further developed in addition thereunto, is only higher completion. But what it, as sexual system, does, it does only through sensation.
1793. It is natural that the animal has not been concluded with the sentient sexual system; but that the terrestrial processes also are developed like as in the plant, and perfected indeed more individually than in the latter. The main distinction is, however, this; that these processes have preceded in the plant, and the sexual system grown out of them; in the animal, on the other hand, the sexual system is the foundation, the root, out of which these processes grow forth. The first and simplest animal vesicle is a sexual cyst, a _matrix_ or womb.
SENSITIVE ANIMAL.
1794. The animal is a twofold representation of the organization, being at one time the planetary, and at one time the solar. There is a planetary animal and a solar animal in the higher organism. The planetary animal is the plant in the animal, the galvanic animal; the solar animal is the sentient, the light-or æther-animal.
1795. The highest completion of the vegetable animal is in the blossom. There is thus a sexual or _Sex's animal_ and a sensitive or _Sensation's animal_.
1796. The completed animal consists of two animals, because it is at once planet and sun, plant and animal.
1797. The vegetable and the sensitive animal have been formed parallel to each other, yet in such a manner that the former being the lowest, contains only the dispositions unto the highest. There are consequently vegetative and animal organs, which take a parallel range. The animal grows upon a vegetable body. It may be aptly said, that the root of the plant becomes the mouth or head of the animal, the stem the trunk or visceral body, the blossom the sexual parts. The three parts therefore of the animal body, and the antagonism therefore of the head with the sexual parts, unite through the medium of the vegetable stem, or the visceral body.
ANIMAL ANATOMY.
1798. The parts of the animal body divide, as in the plant, into tissues, anatomical systems, and into organs proper or members. The tissues are the constituent parts of the systems, these of the organs, and all collectively, or associated, of the body.
I. _Animal Tissues._
1799. These are the mathematical primary forms, whereof the animal body consists, and divide into the animal and vegetative fundamental forms.
A. ANIMAL FUNDAMENTAL FORMS.
1800. As animality is the representation of the three conditions of æther, and thus of the gravity, with light and heat or motion, so are three tissues to be met with, which correspond to these three forms. The light emerges from the centre; the gravity occupies the whole mass; the motion oscillates between the two. The organic light-mass will therefore, as sun, occupy the centre, the gravity-mass, like the planets, the periphery, the motion-mass, like the heat, the radii between the two. The primary form is, however, the primary vesicle. If, therefore, new tissues appear in the animal, they can thus be only metamorphoses of the vesicle. The vesicle, can only resolve itself into three forms; either its contents become self-substantial--_Point_; or in like manner the envelope--_Line_; or, finally, both become an uniform mass--_Globe_.
1. _Point-tissue._
1801. We take up or commence the study of the animal substance, with the condition under which it has originated, that of a vesicle or sensitive blossom. The vegetable texture attained a form which was prescribed by the light. Now, as the animal vesicle is, in the first place, nothing else but a Sentient, so must the texture of the original animal vesicle be commensurate with this property.
1802. The highest perfection of the blossom was, however, resolution of the texture into the original form of vesicles or granules, a retroconduction of the organic mass to the primary condition, yet under the signification of light. The Highest of the blossom was an organized and designedly-prepared granular texture--pollen.
1803. Now a substance which absolves itself from the terrestrial forms, and would assume the form of the æther, and thus of that which is most discrete, can represent no other form than that of the point. The whole æther is an infinity of non-coherent atoms. This atomic formation, when metatyped or copied in a terrestrial mass, can be none other than a granular substance.
1804. The fundamental substance of the animal is point-substance; but, since the essence of the animal consists in its being a sensitive substance, so must it belong to the latter's essence, that it be atomic or punctiform. The point-texture is equivalent to the sensitive mass.
1805. It might be believed, that as the animal is a floral vesicle, the cystic or cell-form must also lie at its foundation; only, there is another relation beyond that which occurs in the plant. This animal cyst is an already organized cyst, an _organ_, no longer a component part of an anatomical system; this cystic form cannot, therefore, enter into the texture of the animal mass; yet meanwhile, as is natural, the sensitive mass admits of being reduced, but only as organic in a general sense, to the vesicular form. The lowest animals, such as the Infusoria, Polyps, Acalephæ, or sea-nettles, in short, all Protozoa or mucous animals consist of this point-substance, and are wholly sensitive mass.
_Nervous Mass._
1806. The sensitive mass is called in higher animals, _Nervous mass_. The nervous texture is a conjoined series of mucus-granules, which have become albuminous in character. The nervous mass is the least organized; it has selected the primary forms, which have been preconstructed in æther, or the densely fluid solar matter. The Dominant of the terrestrial organs can have also no other form, than such as agrees with the Dominant or governing primary mass of the planetary system; or it can have none other, because at the instant when it exists, it is sentient. At the first instant of the origin of organic matter, it can, however, originate only as infinitely numerous points; at the termination of the plant, this mass would be displayed as a light-organ; being thus engendered as such, it must forsake the vegetable forms, and assume the universal form-susceptive primary form, which is the form of the point. The floral mass, the delicate petals of the corolla, the stamina and the pollen are to be deemed the first onset or advance that is made to nervous mass. The cellular tissue becomes delicate and gradually resolves itself into granules.
1807. Granular or point-mass is, however, an accumulation of centres. The nervous mass is therefore, in accordance with the conception of the Organic, a repeated, multiplied centre. The nervous mass has therefore a light-function, i. e. the gentlest polarization in the organism. Nervous mass is light-mass.
1808. The animal substance has commenced with the nervous mass; thus with that which is the highest, and which physiologists have deemed to be the ultimate mass. The origin of the animal is from the nerves, and all anatomical systems are only free evolutions or separations from the nervous mass. The animal is naught but nerve. What it is further or in addition, is obtained elsewhere, or is a metamorphosis of nerves. The mucus of the Infusoria, Polyps, and Acalephæ is nervous substance upon the lowest stage or degree, where the other substances that are therein involved and merged, have not as yet been perfected in an isolated manner.
1809. The nervous mass indicates the absolute Indifferent in an animal, and consequently that which is polarizable by the gentlest aura or breath.
_Division of the Nervous Mass._
1810. The nervous mass behaves itself also in its production like the solar mass. As from this the planets have through antagonism on the periphery, emancipated themselves, so from the nervous mass have the anatomical systems, which are subservient unto lower purposes.
1811. The development of the animal organs is a constant division of the nervous mass, whereby it becomes more and more divested of their coarse coverings, and traverses the same like radiating æther, illuminant, heating, and moving æther. It is a positing of the centre in the periphery.
1812. When also the other systems have been formed out of the identical nervous mass, still the whole animal body is naught but nervous mass, only, in a crude or inert condition. There is, consequently, no point upon the body, on which the nervous phenomena are absolutely wanting, or where they may not appear under certain relations.
1813. What remains behind of the nervous mass, has now the form of filaments or rays, which are projected from one centre, the brain, to all parts of the periphery.
1814. The nerves being individualized, and withdrawn from the coarser mass, stand in need of no _actio in distans_, or no nervous atmosphere (although for other reasons such a one may exist), in order that every part of the body should have sensibility or feel; for every substance is verily but an aberrant nervous substance, in which the original spirit is still inherent or abides.
1815. Each part of the body has consequently irritability, and each one has the capacity for sensation; and that, indeed, through and in itself, or not borrowed from, what have been called, nerves; as it is indeed only the coarse nervous envelope of the more delicate nerves.
1816. Yet, meanwhile it is certain, that nothing feels but the nervous mass, because every thing, which feels, does, and hath the power to do so, only in so far as it has been _nervous mass_. Precisely as all metals are only magnetic in so far as they are metamorphosed iron, which is the primary metal.
1817. Certain formations or textures must, on that account, have different sensations, because they have deviated more or less remotely from the primary texture; _without_ in this any regard being paid to the number of nerves running to such systems.
1818. The transformation of the nervous mass upon the periphery is chiefly imparted by oxydation, because here the oxygen of the water exerts a direct influence. The transformation does not, however, simply occur upon the periphery, but also internally, and that indeed in a beamy or radial direction. The external parts will be harder, the internal or radial continue soft, but be more solid than the nervous mass itself.
2. _Globe-tissue._
1819. The nervous tissue alone cannot constitute the animal substance concerned in all functions, but it must with further development pass over into another. As the æther-mass cannot concentrate itself into the sun without, from the antagonism with the refraction of light, condensing into planets, so also a central mass cannot subsist in an animal, without converting itself on the periphery into one that is planetary or terrestrial.
1820. The antagonistic mass, originating in a peripheral relation in the nervous mass, will surround the residue of the latter like a cyst or cyst, just like the planetary masses, or the colours, have primordially surrounded the sun as great hollow globes. The aggregate and purer nervous mass becomes thus directly the central mass of the animal--the _brain_.
1821. The limitary mass originates through oxygenation. Thus the colours originate; they are an oxygenated light. Thus has every terrestrial mass originated through combustion. The planets are suns that have undergone combustion; the limitary mass is nervous mass similarly treated and deoxydized.
1822. As having been already subjected to combustion, it therefore becomes polarizable, and consequently susceptible of sensation in the least degree. The limitary mass must be rigid or fixed; for it has indeed originated through fixation of the poles, or through the strongest oxydation. The limitary mass is the most rigid in the whole animal; for it is the primary antagonism with the nervous mass, the ultimate planetary matter, which is characterized by immoveability of the atoms.
1823. The limitary mass must be typical of the earth-element, this being the most rigid, and the end of the oxydation. The limitary mass is the animal earth-mass, just as the median mass is the animal æther- or fire-mass.
1824. The texture of the animal earth-mass must be that of a crystal, but of a round globular crystal; for it is organic mass, and can consequently have been only deposited as vesicle; it is, however, earthy mass, so that the whole vesicle must be, with all its contents, rigidified. Now, the rigidified substance of a vesicle is a globe--the texture of the mass which is opposed to that of the nerves is consequently the globular form.
_Osseous Mass._
1825. The rigidified limitary mass, which exhibits histologically dense globes, consists of earthy substance, and surrounds the nervous mass, is _osseous mass_. The osseous or bony texture is a solid globe or rigidified vesicle, being thus ambitus or boundary, as well as complexus or contents.
1826. Bone can only originate through oxydation of the animal mucous or nervous mass, whereby it is converted into a vesicular form. These vesicles are, however, by virtue of the highest oxydation, which must necessarily enter into antagonism with the highest central organ, converted wholly and thoroughly into rigid substance or earth, which is the maximum of the oxydation or fixation of æther.
1827. The osseous mass, as the organic earth-mass, corresponds to the gravity. It is the materiality in a general point of view in the Organic, and consequently the Inert.
1828. Osseous or limitary organs will become more rigid in the air than in water. The air-breathing animals must have more perfect bones or harder limitary organs.
1829. Bones are therefore either wanting completely in the aquatic animals, or they are mucus scarcely oxydized, in other words, cartilage; or finally, almost entirely rough carbonated earth, in the corals and shells.
1830. This theory is most beautifully proved in the corals. Internally they consist of granular substance, like the polyps, or of sentient nervous mass; externally they are simply earth or globular form, which is the rudest antagonism presented to the likewise rude central mass.
1831. Bone essentially surrounds the nervous mass. The skull environs and incloses the brain, the vertebræ the spinal cord, the ribs, the visceral nerves the snail's shell all the soft parts of that animal, the coral stem its polyp-tube, the horny coat the insect.
1832. The purest and highest antagonisms in an animal are nerve and bone, and as such they are demonstrated on every occasion. The nerve is that which is soft, powerless, changeable, sentient, governing, and motion-imparting; the bone, what is hard, strong, unchangeable, non-sentient, governed, and becoming moved; the one properly speaking the spiritually vitalizing, the other the spiritually dead, or self-subsistent simply in a mineral point of view. The bone is the obedient planet of the nerve.
1833. Point-and globe-form are consequently, as regards the tissue of the substance, the first two forms of the animal body.
1834. What develops itself apart from nerve and bone in the animal, must either range between or below both; it must participate of both forms, or be only their incompletion.
3. _Fibrous Tissue._
1835. The nervous and osseous substance could not range opposite to each other, without a transition or a something interposing; as little as could æther and Terrestrial, or sun and planet, between which the moved æther or the heat oscillates, and conditionates the planetary motion.
1836. Between the soft point-form of the nerve and the hard globe-form of the bone, a semi-oxydation stands midway, just as the air stands between the æther and the earth. As this is the medium element, wherein the light is refracted into colours, and thereby warms and moves the planet, so must this median animal formation be the element, through which the nerve imparts its motion to the bones.
1837. This organ stands, like the air, upon the middle rate of oxydation; the oxygen becomes alternately united with it and set free; which is neither possible in the point-form, as being that which incessantly liberates oxygen, nor in the globe-form, as being that which always holds or retains the oxygen in union with it.
1838. This tissue must consist of firm or solid nervous granules, which have been serially co-arranged in lines or radii. Such organic lines are called _Fibres_.
1839. The fibrous is the third original tissue, which appears in the animal organization.
1840. The nerve acts upon the fibres as upon the bone, or as a Central upon a Peripheric, as the light upon the air.
1841. Thereby the soft fibre is polarized; the poles are mutually attracted and repelled, and motion of the fibres originates, their extremities approximating or withdrawing by virtue of the polarity. Contractile fibres are called _sarcose or fleshy fibres_.
_Flesh._
1842. The flesh is the median formation between nerve and bone. It is half nervous mass, therefore sentient, half bone, therefore moving.
1843. The essence of the motion resides in the muscle, not in the nerve. Such is the cause of motion, the muscle being the self-moving, the bone that which is moved.
1844. The flesh must surround the bone, as the air or water surrounds the earth.
1845. The flesh is a terrestrial substance, just as the bone is; the nerve is a cosmical substance, and on that account the mediator of everything.
B. VEGETABLE TISSUES.
4. _Cellular Tissue._
1846. There are three, and only three constituent forms essential to the animal substance, the point, the globe, and the line; equivalent to centre, periphery, and radius.
1847. Out of these three all other forms, of whatever kind, are developed, through degradation unto vegetable structure. This form can be none other than the _Cell-form_. In the animal there are therefore four fundamental forms, while in the vegetable only one occurs.
1848. The cellular form may be also called the water in the animal, the globe-form the earth, the fibrous the air, the point-form the fire. Thus is the animal even in its tissues a whole universe, for it cannot otherwise be thought of.
1849. The cellular substance is the last division of the point-substance, because the nervous granule becomes hollow. A true cellular tissue first makes its appearance therefore in the higher animals.
1850. Bone, flesh and nerve are the highest organs of the animal; the viscera, which mostly consist of cellular tissue, will indicate the Vegetative in the animal. Proper animal organs can only present the above-mentioned triplicity. What is not bone, flesh or nerve, is not animal, but vegetable.
1851. Nerve, flesh and bone are mutually excited, and are independent of the cellular tissue. They are moreover the animal in the animal, the thoroughly Free and Voluntary.
1852. These three substances have therefore nothing to do with the three terrestrial processes; they do not digest, respire, nourish nor transport about the galvanic sap, but live for themselves or to their own satisfaction.
1853. The origination of the three inferior substances out of the nervous mass, is perfectly similar to the original creative process of the three terrestrial elements from the æther. The animal organism is a second world-creation, for in the organic æther an organic air, earth and water, have been produced, or it has itself become these through fixation of the poles. This capacity for resemblance between the organic and inorganic elements is marvellous; but yet there would be more to wonder at, ay, it would be thoroughly incomprehensible, if the organic elements had been created according to some other type.
1854. When once the nervous mass has become separated from the three other masses, each then commences to be self-substantially perfected, and become a particular organ, though it is still under the supremacy of the primary mass.
1855. The principal mass which constitutes the animal body, after complete separation of the chaotic nervous mass, is without doubt the fundamental mass of everything organic; being the mucus or cellular mass, in which the other elements have only been included like veins of ore. It is the cellular mass therefore which we will first consider in its process of formative evolution.
1856. As cellular mass it must be the seat of the galvanic process, and thus of the life proper. In the cellular mass consequently the three vegetative processes, or the three terrestrial elementary processes, must be firmly established; as there digestion, respiration and nutrition are.
1857. With these three processes, the three superior elementary forms, which are peculiar to the animal, as nerves, bones, and muscles, will have nothing to do, excepting in so far as they govern them. As in the plant, so also in an animal, the terrestrial processes are only the appurtenance of the cellular tissue.
1858. In cellular tissue is therefore the seat of life. But the vegetative mass simply lives that it may live, while the animal lives, in order to combine the universe with the life. The animal elements live only in order to feel and move, in order to act freely like the world; the vegetative only, that they may subsist as planet. The latter are an image of the planet, the former of the world; the one deal with matter, the other with spirit.
_Integument._
1859. The cellular tissue does not continue a mere parenchyma in the animal as in the plant, but it obtains a definite anatomical form.
1860. The animal cellular tissue has issued forth from its highest formation in the plant, or out of the blossom, which is a great bladder or cyst composed of primary vesicles. It is the secondary cystic form, wherein the animal cellular tissue appears, when it becomes an anatomical system.
1861. The animal cellular tissue forms therefore everywhere _large bladders_ or _cysts_, whose walls consist of primary vesicles, or of the vegetable cellular tissue. Cyst-walls are teguments.
1862. The cellular system in the animal is _Tegumentary system_.
1863. The idea of the integument is the wall of a cyst. There is no tegument apart from the meaning of circumscription, inclusion, and limitation. There is no flat integument, or one that could be designed after the idea of the plane. Every tegument is periphery, just as there is nowhere a surface in the universe, which could have been produced according to the level or plane.
1864. All terrestrial processes, as digestion, respiration, and nutrition, are consequently tegumentary processes. All these organs must be tegumentary organs. Intestine, vessel, lung, in a word all viscera are naught but tegument.
1865. The vegetable tissue becomes in the animal, tegumentary tissue. The tegumentary formation is the vegetable in an animal--the parenchyma, the _Visceral_.
1866. The lowest distinction between animal and plant resides accordingly in this, that the vegetable tissue consists of actual vesicles, which form everywhere closely compressed masses; the animal cellular tissue on the contrary of granules, which inclose a hollow space. The animal body is a hollow globe of vesicles, the vegetable body one full of vesicles. (Ed. 1st, 1810, § 1870.)
1867. Every animal cyst is necessarily composed of the element of the vesicles, and is then for the first time an organ. The vegetable bladders are, however, single vesicles, and as such are already an organ. In the plant therefore the cellular tissue is upon the lowest stage, being only an aggregate devoid of secondary form; in the animal along with its aggregation a secondary form has been imparted. In this the higher character of the animal is at once demonstrated.
1868. The above is certainly a distinction between the two organisms, but it is not the _essential_ one; for with it, what is animal has been by no means expressed; this being first imparted in the three cosmic elemental forms, which manifest themselves through sensation and motion, and then admit of being recognized as an animal. The corolla is also a cyst, but without being an animal; because, to this animal-like tegumentary formation, the proper animal elements are still wanting.
1869. Now as the tegument is none other than the form, under which the cellular tissue exists in the animal, we must regard it as an elemental form, which has stepped into the place of the cellular. The tegumentary form constitutes the fourth form, and is none other than the primo-vesicular form elevated to a higher rank, by being composed of cell-granules, which have been formerly nerve-granules.
1870. The animal body must consist of nerve, muscle, bone, and tegument, and of no other fundamental form; in other words, of point, line, globe, and cyst.
II. _Anatomical Systems._
1871. All anatomical systems are developments and separations of the four tissues, which are prolonged as sheaths through the whole body, like as in the plant are the bark, liber and wood.
1872. They are divisible next of all into two great parties, into the terrestrial and cosmical, or vegetative and animal.
A. VEGETATIVE SYSTEMS.
1873. The vegetable systems can only be different developments of the tegument. They accord with the galvanic factors. Any further derivation of them is unnecessary. The tegumental development must be represented as the systems of digestion, respiration, and nutrition, since from it these have been sufficiently borrowed or derived.
1874. Except these three systems there can be no other tegumental system; and, if such appear to be present, they must be subordinate to these. For there cannot be subsequently any more than there was fundamentally or at bottom.
1875. In the animal, however, the galvanic processes do not remain entangled in _one_ mass, as in the plant. But they are even characterized as animal by their individual liberation from the whole mass. In the plant digestion or absorption, and nutrition or the course of the sap, were in one kind of mass or one kind of cellular tissue, all three processes (together with respiration) being in a tolerably confused condition.
1876. The animal appears in its dignity by separation of these processes, and by the perfection of each individually, or "per se."
1877. As all life consists only in the constant conversion of what is inorganic into the Organic, so is the process of digestion or absorption necessarily the first in animals.
1. INTESTINAL SYSTEM.
1878. The chemical process of the galvanism is conversion of the Inorganic into mucus, and thus an assumption of this matter into the organic body. Now as every limit of the body is tegument or cellular tissue, so can this assumption take place everywhere. Adoption of what is external into an organic body is _absorption_.
1879. Absorption originates from the antagonism of the body with the earth, which is organizable, and thus with the mucus.
1880. We call this slime or mucus, nutritive matter. Wherever such matter can operate upon the body, a corresponding organ of absorption, and thus a cell or integument, will be formed.
1881. The whole body is surrounded by integument; it was originally nothing but integument.
1882. The essence of the integument consists in absorption, or in the intervention of the chemical process.
1883. The integument is the root of the animal.
1884. The animal cellular mass is, however, in conformity with its origin, a bladder or cyst that has been opened by light and air. The integument is a large cyst not closed all round, but open at _one_ end. It is the open floral cyst, which has just become an animal. The original integument is thus _Intestine_. The intestine is the water-organ.
1885. The integument presents therefore to the external world, or to the nutritive matter, two parietes or walls, an external and an internal.
1886. Both walls are opposed to each other like light and darkness, like air and water. The external is the light-and air-wall, the internal the darkness-and water-wall.
1887. It is consequently only the internal wall that stands in the same relation as the root. The internal is "par excellence" root, and is thus a main organ of absorption.
1888. The external wall comes under the idea of the stem-bark, and in so far only as this has a root-nature in itself, is it likewise absorbent.
1889. As upon the external animal wall the light and air constantly operate--for without light indeed no animal originates--so is this wall more and more removed from the idea of the root, and becomes, by virtue of the influence of light and air, instead of an absorbent organ, an organ rather of decomposition--an _evaporation's organ_.
1890. As a cyst the internal wall incloses the nutritive matter, which originates from the mucus, and thus from the organic water. The internal wall is therefore constantly immersed in the water, and is consequently in every respect a root.
1891. As the animal is only developed in light, so must the function of the root languish in the external wall, and decay, because it is devoid of the earth, which protects it from or against the light. This deficiency is compensated for in another way, or by the formation of a cavity, into which the media of nutrition enter, and which is dark like the earth.
1892. Internal and external wall stand also opposite to each other, like water and air. The one is the water-, the other the air-wall.
1893. The nutritive matters are not decomposed upon the internal wall by extraneous influences, but they remain identical; ay, they become indifferent, because they enter into darker and warmer water.
1894. On the other hand such nutritive matters are decomposed upon the external wall; and there here therefore gradually originates, instead of the chemicalizing root-process, the polarizing process of air.
1895. In a perfect light-animal it is only the internal wall that is chemicalizing; the external has become oxydizing. The internal is a mucus-wall, but the external, on account of the decomposition of mucus, an oxygen-wall.
_Division._
1896. The more an animal has been exposed to the air and light, by so much the greater is the antagonism between its internal and external wall. In aquatic animals the antagonism is at its minimum, because externally and internally there is water; both parietes are therefore mucus-walls. The external wall of the fishes secretes an abundance of mucus, as does that also of the worms, snails, and molluscs.
1897. But an internal wall is still the more mucous of the two, because it is darker and warmer.
1898. At first the animal is content with the antagonism of the walls; especially so long as it remains occluded in dark and deep water, or within other animals. Many intestinal worms, polyps, and even Acalephous animals, are but simple sacs.
1899. When the animal organization, however, ranks upon a higher stage, light, or even air, operates more upon its external wall, but upon its internal, water; thus the antagonism of the two walls is carried out to the utmost degree.
1900. Through the different, ay, opposed processes, the two walls finally adopt another structure. The external becomes denser and harder, on account of the decomposition by light and dessication by air; the internal, however, retains its original structure and consistence. Soft, aqueous, indifferent, and constantly absorbent, it is only a viscous mucus.
1901. In place of an integument of similar tissue throughout, one will originate, whose external tissue is dense and oxydized, but whose internal is loose or spongy, and indifferent. The previously uniform integument will now separate or fall into two distinct layers; into a soft muco-cellular layer, and into a tough coriaceo-cellular layer.
1902. With the last attainable antagonism the layers finally separate; two bladders or cysts, disunited from each other, originate; of these the internal is the _mucous_, the external the _coriaceous cyst_.
1903. Now the internal cyst alone is the _intestine_, the external the _cutis_ or _skin_.
1904. Intestine and cutis belong to _one_ formation, or to the integument. They pass directly into each other at the mouth and anus. Their structure also is wholly similar.
1905. They are merely distinguished by darkness and light, but more closely by water and air. The intestine is the _water-tegument_, the skin the _air-tegument_.
1906. The functions of both are therefore co-related like dissolution is to combustion, along with which the evaporation has been bestowed.
1907. Intestine and skin stand in antagonism with each other.
1908. The first animal, as being sentient integument, is a sac; the first skin is also a sac; an animal around the animal.
2. DERMAL SYSTEM.
a. _Branchiæ or Gills._
1909. The external wall, being constantly exposed to the air that is in the water, can adopt no other than the aerial character, and is thus like the leaf of the plant. The skin is the organ of evaporation, and with this of oxydation also.
1910. A self-oxydizing integument is called a _Branchia_ or gill.
1911. The skin is essentially nothing else than a gill; and, if it subsequently appears as anything else, this happens only through a higher state of perfection being attained by its branchial function.
1912. The lowest animals, such as most of the worms, molluscs, and snails, breathe through the external integument; even the gills of fishes are none other than a piece of skin.
1913. Thus gills and intestine would be the first two organs, which are developed out of the tegumentary system by the antagonism of air and water. Through the gills, air, and through the intestine, water enters the body. The gill is the atmosphere of the animal, the intestine is its sea.
b. _Tracheæ, or Air-tubes._
1914. As the intestine, and in general every water-tegument is prolonged into mucous tubes or absorbent vessels; so also, with a more vigorous formation, the branchial membrane is drawn out into tubes, in order to conduct the air or oxygen towards the intestinal vessels, just as the intestine conveys through its absorbent ducts the water to the vessels of the skin.
1915. This saccular inversion of the skin constitutes the tegumentary lymphatic vessels, whose original function has been to transport the oxygen, combined with the water, to the intestine. They are the original respiratory vessels, which in the higher animals become, through the pure influence of air, true _Air-tubes_, like the spiral vessels.
1916. The air-vessels traverse the thickness of the body toward the intestinal membrane, like the mucous vessels do toward the branchial membrane.
1917. Thus an infinite number of air-vessels will and must originate.
1918. The air-tubes are consequently the formation which is properly opposed to the lymphatic vessels. They are for the air or for the skin, what these are for the water or the intestine. Air-vessels are first displayed in insects, then in fishes, reptiles, birds, and Thricozoa or Mammalia.
1919. If the infinitely numerous air-vessels concur to form one stem, they are then called _lungs_, as in the higher animals.
1920. The pulmonary vesicles are nothing but ramified air-tubes, such as the insect has.
1921. The formation of air-tubes is one of a higher character than that of the gills. For in it, indeed, the function has been separated from all other functions. It is simply destined to convey the air without water.
1922. As they pass into the dark, light does not operate upon them; and they will therefore less promote evaporation than mediate unto combustion.
1923. The anatomical idea of the air-vessels, or of the lung, is a saccular inversion of the skin. The skin is prolonged into, and ramifies towards, the body. The intestine is prolonged through the absorbent vessels, as being also small inversions, toward the lung, and becomes a stem--thoracic duct. Then the thoracic duct finally unites with the lungs through the medium of the heart, which is a new formation.
1924. Everything becomes stem which attains a higher grade, and which approximates the air and light. The stem strives to be a centre, but the ramification devolves upon the periphery; the former upon the Solar or Animal; the latter upon the Planetary or Vegetative.
1925. The nobler therefore a formation, by so much the more single and stemmy is it: such is the case with the trachea of the lungs, such with the lymphatic duct.
3. VASCULAR SYSTEM.
1926. The earth or the nutritive mass acts also upon its formation, and destines the tegument to a peculiar structure. The result of the electric and chemical process, or of the oxydation of the mucus, is precipitation, mass-and earth-formation; it is thus a process of nutrition, since through it the Solid of the body, with even the branchial and intestinal membranes, originates.
1927. The earth-system can only be developed where the two last systems coincide, or where the intestinal and branchial processes enter into mutual contact; in short, where the mucus is oxydized, and thereby separated into what is aerial and rigid.
1928. This spot is only in the middle between the two. The process of precipitation and the formation of elemental matter occurs, consequently, between the two layers of integument, or at present between the two teguments that have become self-substantial, or between intestine and skin.
1929. A new formation must be evolved, whereby the two shall be held together; a formation, whereby the antagonism shall be conducted from one to the other; and thus whereby the mucus shall be conveyed to the skin, but the air to the intestine.
1930. Were intestine and skin entirely separated from each other, each would perish; the former would no longer be oxydized, the latter no longer nourished.
1931. With their separation they must continue to be attached to each other at certain points, and thus undergo eversion at certain points. Thereby tubes originate in the intestine, which pursue their course toward the skin; in this again tubes, which pass to the intestine.
1932. A tube, which receives mucus from the intestine, air from the skin, and includes both within itself, is a long cyst. A cyst, which conducts mucus to the skin and air to the intestine, is a _Vessel_, a _vein_.
a. _Unclosed Vascular System._
1933. The vessel has, in virtue of its essence, two extremities, an air-extremity, which is polar, and a water-extremity, which is indifferent. Every vessel has been rooted in two systems, in the intestine and skin, and is subservient to both.
1934. The vessel is no longer a something single, like the last-mentioned or preceding cysts, but double. Every vessel hath two poles.
1935. The organization necessarily produces two kinds of vessels. A vessel, which conducts the mucus to the skin, cannot also convey the air to the intestine. There is consequently a mucus-vessel and an air-vessel, or a water-and an air-vessel, an indifferent and a different.
1936. The mucus-vessel is called _absorbent_, the air-vessel _respiratory duct_ or trachea.
1937. Air-and absorbent vessel stand in antagonism like skin and intestine, like water and air. The air-vessel is the skin or the branchia, which passes to the intestine, the absorbent vessel is the intestine, which passes to the air; the one the intestinal branchia, the other the branchial intestine.
1938. So long as intestine and skin were one in kind, this vascular process was in every situation. With their separation therefore the vascular structure has of necessity originated between two opposed situations. There is no point in the skin and none in the intestine, where there might not be an air-and a water-vessel, a respiratory and absorbent duct.
1939. There are therefore _numerous_ vessels, and consequently a _Vascular system_.
1940. Air-and water-vessel must abut against each other; because they are polar, because the one leads to this place and the other to that.
1941. The system of water-and air-ducts can form no closed vascular system; for they only grow towards each other, as did formerly intestine and skin.
1942. They would not have originated if both cysts had not separated from each other. These vessels are not therefore to be met with in animals that have no intestine. The transition of the water-into the air-ducts takes place in the higher animals through the union of the thoracic duct with the subclavian vein, which conveys the blood directly to the lungs.
1943. The vascular system is properly the primo-cellular tissue, which occupies the middle, and at whose extremities the two cysts remain approximated, in order that they may continue to live.
1944. The vascular is the original system, since its two extremities already carry in themselves the air-and mucus-process, so that branchia and intestine are only to be viewed as peculiarly perfected conditions of these extremities. I would even have developed the vascular system first of all, had not another course been required to render the subject intelligible.
1945. The first animal cyst is a vessel with two kinds of extremities, whereof, like the plant, one draws its supply from the water, the other from the air; but in the animal this is effected by its own motion.
1946. The branchia is nothing but a vascular tissue in the air, the intestine none other than a vascular tissue in the water. Thus, do I wish that these organs should be understood, and not as mere cystic walls.
1947. Every point therefore in the intestine and skin absorbs, and thus every part is perforated with infinitely numerous holes or pores. Here they absorb air, there water.
1948. Every absorbent point of integument is drawn out as a tube towards the respiratory system, in order to let what it has absorbed become oxydized. These tegumentary prolongations into tubes are the _lymphatic vessels_.
1949. In all teguments there are necessarily lymphatic vessels, but more of them in the water-than the air-teguments. On that account the lymphatic vessels are much more numerous in the intestine than the skin. They are there called chyliferous, or lacteal vessels.
1950. The lymphatic vessels are the first of all vessels. Many animals, as perhaps the Acalephæ and Distomata, appear to have only this kind of vessel.
1951. Lymphatic vessels are present in the skin, only in so far as it has resigned the respiratory function to special organs.
_Meaning of the Unclosed Vascular System._
1952. The action of this unclosed vascular system is wholly similar to the motion of sap and air in the plant, there being only an ascent of the first, and a fall or descent of the last. In the absorbents the sap ascends out of the root (intestine) into the leaves (branchiæ); in the respiratory vessels the air descends from the foliage (skin) to the intestine and the whole body of cells.
1953. This vascular system is therefore the pure remnant of the plant, and has as yet assumed no properly animal character, except that both its sets of vessels or tubes are self-substantial, and ramify, while in the plant they are only intercellular passages or non-ramified spiral vessels.
1954. In insects this system has been most perfectly evolved; there the air-tubes, being well parcelled out, are in great number, and run directly to the intestine and dorsal vessel, which is, as it were, only the trunk of the lymphatic vessels, or the chyliferous duct.
1955. Now, such a vascular system, merely oscillating, as it were, between intestine and skin, can be but persistent in animals, which express only the vegetable type of organization. If other significations be introduced, so also will this vascular system be otherwise evolved.
1956. The unclosed vascular system will be present along with an energetic antagonism between skin and intestine; with an antagonism that is almost suppressed, there will be none. There are then only cells, or there is only point-substance, as in the Infusoria, polyps, and Acalephæ or sea-nettles.
1957. The dermal vessels exist only in the air-insects, because in them nothing but air and water are engaged in conflict. Externally there is desiccated horn, internally mucous water.
1958. A perfectly unclosed vascular system appears to be developed only in animals which respire air. At least there are only genuine air-tubes and lymphatic vessels in such as breathe air; e. g., in the Mammalia, birds, reptiles and fishes, the latter set of vessels being probably not present in insects.
1959. Through the predominance of the air-process, as in insects, the mucus that is conveyed to it becomes so rapidly decomposed, that no more remains behind, for which a new vessel would be necessary.
1960. The galvanic process is at every instant annulled, and only renovated by a new afflux or supply. Here the galvanism does not subsist in itself as a peculiar and independent system.
b. _Closed Vascular System._
1961. The unclosed vascular system is not yet self-substantial, because it is a cæcal eversion of the intestine or inversion of the skin, being itself only a ramified intestine and skin. Every system, however, attains its perfection, by being rendered independent of its origin. Thus the leaf is the spiral vessel that has become free, the root the cellular tissue in a like condition, the blossom the liberated vegetable trunk. The vascular system will therefore aspire also to the achievement of its blossom.
1962. If the decomposition that is effected by an inferior amount of polarization, does not happen rapidly enough, for what has flowed thither, to disappear, during its afflux, by evaporation or precipitation; the rest of the mucus which has obtained the air-polarity, is now repelled by the respiratory vessels, because both have become synonymous.
1963. The oxydized superfluous mucus is at once, however, attracted by the intestine, because they are not synonymous. There consequently originates a vessel in the mass of mucus or parenchyma, that has been secreted between the intestine and skin, which begins in the respiratory membrane and terminates in the intestine.
1964. This vessel will commence at the end of the absorbents, or at a point, where it devolves upon the respiratory vessels, to take up their contents, namely the oxygen, together with the nutritive substance, and convey them to the intestine. The mucus, which previously stagnated and moved but slowly from one spot to another, is now again carried back by another vessel without interruption to the intestine.
1965. The vessel which conveys oxydized mucus from the respiratory to the intestinal system, is called _artery_.
1966. On the intestine, however, this polar mucus is again reduced to ordinary mucus. It has now become synonymous with the intestine, is repelled from it and attracted by the branchial membrane.
1967. One and the same fluid or sap is consequently brought back from the branchia to the intestine, and from thence again to the branchia. This last vessel is called _vein_.
a. _Arteries._
1968. The artery is, according to its signification, an air-vessel, which is prolonged or elongated as far as the intestine. Essentially, the artery conveys nothing but air, though this is effected by means of a medium or vehicle, which is the undecomposed mucus (blood). It is an air-tube, that has been self-substantially dismembered from the skin, in order to become a special independent air-tube, or such as is commensurate with animal nature.
1969. In the artery the external integument has been repeated in the nutritive system, and hence the galvanism has become of a continuous character.
1970. The artery must therefore be the highest vessel, the most total of all terrestrial processes. For it is the air-duct united with the intestinal vessel. It contains mucus, which carries the properties of both poles of the body in itself; there is oxydized mucus.
1971. The artery contains the whole body, lung and intestine, in short, the whole animal, (whereas it was previously dispersed in two vessels,) in a fluid state within itself. From the artery therefore nutrition will directly take place; from it the animal will be formed.
1972. The air or respiratory vessels may be viewed as arteries carried to the very extreme. In the lung the arterial system has attained its highest purity, the oxygen only, without the indifferent substance, being contained therein. The trachea is the roughest artery--Arteria aspera.
1973. The arterial system, in accordance with its signification, makes its first appearance in water, because the aquatic mode of respiration is less energetic, and thus the mucus is more feebly decomposed. The Mollusca, snails, and many worms have a perfect arterial system. The branchiæ do not, like respiratory tubes, pass into the body, as in insects; but there are vessels which take up the oxygen and convey it into the body.
1974. As lung and absorbent belong to the pure air or the pure water, so do artery and vein to the water combined with the air. The two former are therefore present only where aerial respiration occurs, and the two latter where water merely is respired.
1975. The last system is only present in animals, in so far as they are aquatic in their habits.
1976. Insects, as being purely aerial animals, have therefore arteries and veins only so long as they are in the larva or worm-like condition, and as flies or perfect insects may continue to live without them. On the contrary, the purely aquatic animals appear enabled to live without true respiratory and absorbent vessels. It in fact appears, that lymphatic as well as respiratory vessels are wanting in the molluscs, snails, and worms, since the water directly bathes or washes the arteries.
1977. Animals with both systems of vessels, the unclosed and closed, must be more perfect in structure, and must at once combine worm and insect in themselves. They are insects from having absorbent and respiratory vessels, but worms as having arteries and veins.
b. _Veins._
1978. The veins are developed as mucus-vessels at the intestinal extremities of the arteries, which absorb the arterial mucus (blood), after it has deposited its air on the tegumentary substance, just as the lymphatic vessels absorb their fluid from the intestine or any other part of the body.
1979. As the artery is a respiratory vessel that has become self-substantial, so is the vein a similarly conditioned, and dismembered lymphatic vessel. In the one it is the lung, in the other the intestine, that has become the free vascular system. But in the proper vascular system both lung and intestine are repeated, the former as artery, the latter as vein.
1980. These arterio-lymphatic vessels (veins) necessarily convey their arterial mucus or blood into the stem of the original lymphatic system, or the thoracic duct. For every Indifferent must be brought toward the respiratory organ.
1981. The tegumentary lymphatic vessels (absorbents) consequently unite with the arterio-lymphatic vessels (veins), before arriving at the respiratory organ, and pursue their course thither in common, where they pass over into the air-vessels. The usual notion or idea is, that the lymphatic vessels, from conveying their fluid into the veins, should be subordinated to the latter. But the true philosophical view, is of reverse import, although the veins are larger than the thoracic duct. In the investigation of such relations, recourse can by no means be had to quantity or size, but to the importance of the quality or contents.
1982. The veins are, properly speaking, subordinated to the lymphatic vessels, just as the arteries are to the tracheæ, or air-tubes, and the former therefore pass over into the lymphatic vessels.
c. _Circulation._
1983. Through the veins, as arterio-lymphatic ducts, the vascular system has become a closed system in itself, because, on account of the polarity, the vein unites at both extremities directly with the artery. It is a continuation of the artery, like the air-duct is of the skin, and the lymphatic vessel of the intestine. Thereby a _Circulation_ of the arteriose mucus or blood originates.
1984. In its essence the circulation is a combination of the intestinal with the branchial system into _one_ anatomical system.
1985. The circulation is therefore a higher formation, since through it the vascular system repeats in itself the totality of the vegetable organism.
1986. On that account the circulation is the vital process proper.
1987. But for that reason also, the circulation is impossible in the plant, since it is devoid of arteries and veins.
1988. It makes its appearance in the aquatic animals, for they have, generally speaking, vessels. Molluscs, snails, worms, and crabs already possess a circulation; it is wanting, on the contrary, in those animals that are without intestine, and ceases in insects, when, or in whom, the air-vessels obtain the preponderance.
1989. In circulation the galvanism is restricted. In the skin, intestine, air-and lymphatic vessel there is also galvanism, but distributed upon organs that are remote and subservient to different purposes.
d. _Blood._
1990. The blood signifies the earth in the animal, combined with water and air.
1991. The blood is the proper nutritive matter for the animal.
1992. The blood is earth, which carries all terrestrial elements in itself, such as the air through the medium of the gills, the water through that of the intestine, and is consequently a complete planet.
1993. The blood is a fluid planet.
1994. The blood is the fluid body.
1995. The body is the fixed or rigid blood. Blood and body are wholly equivalent, have the same elements in themselves; only here the latter are stationary, there they course along. Both consist of mucus or gelatine, albumen and fibrine.
1996. The blood is half combusted mucus, the body mucus, that has been wholly subjected to combustion.
1997. After the vascular system has attained its own circulation, or to the closed galvanism, no higher development of the tegumentary formation is any longer conceivable; as little as after the blossom anything more could originate.
1998. After all three elements are united into _one_ point, into _one_ system, as is the case in the circulation, where the venous blood represents the water, the arterial blood the earth and air, no new system can further originate in the vegetative body.
4. SEXUAL SYSTEM.
1999. In so far as the animal adopts into itself, or is rather based upon, the whole plant, the blossom or _Sex_ is also developed in it. The vegetable sex consists of seed, capsule, and corolla.
2000. The seed is the first part of the blossom which is put forth in the plant. The most inferior or asexual plants have only seeds devoid of capsule and corolla, and produced without the concurrence of female and male parts. The blossom in the animal is therefore in the beginning also nothing but seeds or ova. Thus these animals are asexual. Such as the Infusoria, whose body directly divides into new animals, like the fungi.
2001. The second floral organ is the capsule, which contains on its borders or dissepiments the seeds, and on its apex the stigma, or the opening of the cyst.
2002. So also in the succeeding forms of animal life the animal capsule or uterus originates. The orifice corresponding to the stigma is the mouth of the womb; the seeds upon the septum become _ovary_.
2003. The lowest animals, as the Polyps, are fundamentally none other than such an uterine system. The polyp's mouth is the os uteri; the sac formed by the polyp's body is the uterus, in which ova-cysts or ovaries develop, that open into the margins of the month.
2004. In such animals the uterus and intestine, as likewise the mouth and uterine orifice are fundamentally _one_ organ; nutritive matter and ova are also one in kind. External tegument as branchial organ is at the same time also a tegument of the uterus.
2005. The higher animals are distinguished from the lower by separation of all these intricate and, as it were, coalesced organs.
2006. Digestion, respiration and nutrition, growth and propagation, are originally of one kind. But with further development come the male parts also, which belong to the category of the organs.
_Parallelism of the Animal and Vegetable Body._
All vegetative systems of the animal body being now developed, the attempt to co-ordinate them with those of the plant admits of being made.
2007. What the _sexual parts_ correspond to in plants, needs no exposition.
2008. And just as little that the _lung_ is the parallel organ to the leaves or _foliage_. The relation of the other organs is, on the contrary, difficult; the striking resemblance, however, of the sexual parts and the lungs to the same systems in plants, is ground sufficient for assuming also the parallelism of the other organs.
2009. If the _root_ be compared with the intestine, then the _stalk_, as being the medium of the _vascular system_, must be regarded as _heart_.
2010. The _bark_ will correspond to the _skin_, the liber to the _veins_, the _wood_ as a tracheal body to the _arteries_.
2011. The vegetable tissue will be transformed into the lowest organs of the animal; the _cells_ into the _mucous tissue_, the _intercellular passages_ into the _lacteal vessels_, the _tracheæ_, or spiral vessels, into the lowest kind of respiratory tubes or the _cutaneo-lymphatic vessels_. We have accordingly the following parallel series:
A. _Tissues._
1. Cells Mucous tissue. 2. Ducts Absorbents. 3. Tracheæ Cutaneo-absorbents.
B. _Systems._
4. Bark Skin. 5. Liber Veins. 6. Wood Arteries.
C. _Members._
7. Root Intestine. 8. Stalk Heart. 9. Foliage Lung.
D. _Sexual parts._
10. Seed Ovum. 11. Ovary Uterus. 12. Corolla Testes.
B. ANIMAL SYSTEMS.
2012. As in the blossom the light allows the whole vegetable trunk to be once more developed, though with coloured signs or marks of distinction, so also is the animal body taken up into the sense of light, and the vegetative systems are elevated unto light-or rather æther-systems.
2013. A new animal originates upon the old animal and is equivalent to it. Every perfect animal is twofold in its nature, being a planetary and solar animal, a vegetative and animal being.
2014. Three animal systems must be developed out of the three vegetative, an earth-, water-, and air-system, each being refined or purified through the agency of light, and forming the bones, muscles, and nerves, which correspond to the gravity or materiality, the heat or motion, and the light or tension.
2015. Everything that is higher in its nature can only be developed out of that which is its direct antecedent, like the blossom from the leaf. Now the artery is the ultimate Vegetative. In this therefore, must the elements of the osseous, muscular and nervous systems reside. Now, the artery consists, of four parts, of the cellular, fibrous, dense serous coat, and of the blood. The cellular coat or tunic is the remnant of the whole tegumentary formation. The fibrous tunic is the embryo of the muscular system. The serous tunic is the embryo of the osseous system; for in old age it attains to ossification, bony lamellæ being deposited around it. The blood is the embryo of the nervous system. It is only requisite for it to coagulate, and it is then nervous mass. Nervous globules are blood-globules quiescent, or in a state of rest. Accordingly, in the artery, the whole body has been actually prefigurated or typified.
2016. Every animal system necessarily exists under a duplex character, being once in the service of the vegetative systems and once for itself; or it exists as trunk and as blossom. There is, therefore, a vegetative and animal nervous system, and, under such binary conditions, osseous and muscular systems likewise.
2017. All three systems are offsets from the arteries, and therefore their constant companions. The animal systems are called _flesh_.
1. NERVOUS SYSTEM.
2018. The nervous mass is coagulated blood. The nervous system is thus a higher arterial system. The highest arteries, however, are the air-tubes. The nerves therefore run parallel with the tracheal systems.
2019. What the tracheæ are for the vegetative body, namely, the vitalizing and motor principle, such are the nerves for both the animal and vegetative body.
2020. The nerves are distributed like the tracheæ of plants, in individual threads or filaments, which run near to each other arranged in a fascicular manner, and then mutually separate.
2021. The nervous filaments everywhere accompany the arteries, and, as far as their ultimate ramifications, like the tracheæ of insects pass to all parts of the body.
2022. The nervous mass has been separated into an arteriose and venous mass, the former being the cineritious or gray substance, the latter the medullary or white. The nervous system is therefore an entire blood-system, with two poles; it is therefore alive and active for itself, or independent of other systems.
2023. Both nervous masses are in a state of constant tension against each other, and consequently in constant tension with the whole body.
2024. The nerves are filaments that, upon the separation of the parts of the body, have remained behind, just like the arteries are tubes individualized in the general parenchyma, throughout which the sap was previously dispersed as in the vegetable trunk. The first individualization necessarily takes place at the oral extremity. The first nerve is a ring surrounding the pharynx or gullet. Thus, if the body consists of several consecutive rings or cysts, as in the Worms, each ring will or can have its nervous ring, which inferiorly gives off nerves to the vegetative organs, and superiorly to the animal, if these be present. Such a point for giving off nerves is called a ganglion; since also every ganglion sends off nerves to join other ganglia; so both below and above a nervous cord or string will originate in the longitudinal direction. The nerves, which have been left behind in the vegetative parts, form the vegetative nervous system; those that remain in the animal parts, the animal system.
a. _Vegetative Nervous System._
2025. The vegetative nervous system is the nervous mass that has remained behind, after the greatest part of it has become converted into tegumentary formations. Now, as these tegumentary formations, being surrounded in the higher animals by flesh, were thus viscera, so may the vegetative nerves be called also _Visceral_ or _Splanchnic nerves_.
2026. These visceral nerves govern the vessels, the intestine, and lung; with the sexual parts also, yet in their case in combination with the animal nerves, because the sexual parts are at one and the same time organs of vegetative and animal life.
2027. These visceral nerves everywhere accompany the vessels, and are therefore like these distributed in a cystic manner between intestine and skin. They form a large cyst which concentrically surrounds the intestine.
2028. They do not, however, like the intestine, form any closed cyst, but only a cystimorphous net, like the vessels.
2029. The two nervous masses are in them separated from each other, like as the branchiæ have been distributed along the whole body and separated from the intestine. The gray or branchoid substance has separated itself from the white medullary substance into individual _ganglia_, or as it were into individual nervous branchiæ. The medullary substance has also retained its connexion though only in a ramular manner, and not uninterruptedly like an integument. It is called _plexus_.
2030. The ganglia and the plexuses stand in mutual opposition, like the branchiæ and intestine, like artery and vein, like blood and lymphatic vessels. The ganglia oxydize, polarize; they are the active. The plexuses suffer, digest, and are the recipient.
2031. The visceral nerves, like the viscera, act for themselves, being unconcerned for the animal systems.
2032. The visceral nerves have a vegetable sensation for themselves, a sense of touch, such as the blossom might have in the instant of the pollenization.
2033. As all formations have become symmetrical through and in accordance with the vascular system, so does the vegetative nervous system separate into two parallel stems which accompany the arterial trunk. They are called intercostal nerves. These ramify and form plexuses that accord with the visceral organs, which they govern.
b. _Animal Nervous System._
2034. The animal nervous system is the repetition of the vegetative, and is combined therewith to constitute an unity. The nervous cyst that was previously dissevered in a reticular manner, becomes a closed tube, which is placed upon the light-exposed side of the other animal systems, and thus upon the vertebral column. This closed nervous tube is called the _myelon_, or _spinal cord_.
2035. The spinal cord is worth as much as all the visceral nerves taken together; it is the felted system of intercostal nerves; and is, properly speaking, none other than the posterior double cord of ganglia. It consists therefore also of ganglia and plexuses, but both have coalesced, on account of the increase of the mass and its endeavour towards the attainment of union.
2036. The ganglia form a tube, which is inclosed by the tubes of the plexuses. The ganglion-tube is the gray, the plexus-tube the medullary substance.
2037. If the gray substance appear to reside within the medulla, this is the result only of involution or a folding in. The parietes of the spinal marrow stand therefore in everlasting tension with each other, like ganglia and plexuses, and like arteries and veins.
2038. The spinal marrow is the content of the bones and muscles, like the intercostal nerves or the visceral cord are the contents of the abdominal cavity, and as the blood is that of the internal and fibrous coat of the arteries. Bones and muscles are the animal parietes of the nerves, as the two vascular membranes are the vegetative walls of the blood. The skin, as branchia, incloses all, like the cellular membrane does the vessels.
2039. As the intercostal ganglia give off plexus-forming ramules; so does the spinal marrow; there are the spinal nerves. The spinal marrow is, first of all therefore, the coalescence of the two intercostal nerves.
2040. These spinal nerves are, however, animal plexuses, which partly encroach upon the visceral nerves, and partly pass to the animal systems.
2041. There are therefore two kinds of spinal nerves, vegetative and animal, and as many of them as there are divisions in the viscera and in the animal systems.
2042. The nerves pass off symmetrically from the spinal cord, because the nervous mass belongs to the symmetrical osseous system. They form therefore rings both anteriorly and posteriorly.
2043. The nervous system does not consist of individual cysts, like its two animal teguments, or bone and flesh. It is at one time the type or image of the vascular trunk and its ramifications; at another the indifferent æther-mass, which does not crystallize; it is lastly the organic primary mass that has remained persistent, and must thus be coherent in texture. It is the blood continually streaming from the animal divisions of the heart.
2044. The whole animal nervous system is a tegumentary cyst with tubes passing off from it symmetrically in the form of rings.
2045. The spinal cord cannot be the highest. It has only the lowest signification, in so far as it stands in the service of the viscera and the sense of touch, and thus follows the position and arrangement of the bones. Thus the spinal cord is first of all an _osseo-nervous mass_.
2046. The nerves, as running for the most part _forwards_, are musculo-nervous mass; those running backwards or outwards are tegumentary or sensitive nerves. This signification furnishes us, also with the physiological function of these two divisions of nerves. The nerves are homologous with the flexors, the spinal cord with the extensors; the nerves with the air, the medulla with the earth; the former with the arteries, the latter with the veins; the nerves are thus what is more active, the medulla that which is more inert.
2047. On that account the nerves only are in intercourse with the world, while the medulla broods within itself. Consequently, both these nerve-formations are not as yet the pure self-substantial nervous blossom, which no longer imitates flesh and bone, but only itself.
_Brain._
2048. The highest point attained by the lower systems are the orifices of the viscera, the mouth and the nose. The mouth is the first animal sign, which the plant gives off in the blossom from itself. The Noblest lies therefore at the anterior extremity of the animal, or in man in the direction _upwards_.
2049. It consequently occupies the middle point, or one that is between the anterior flesh and the posterior bones, and at the same time the spot, from which all vital processes emerge, or the mouth.
2050. The oral nervous mass is the _Brain_. It forms originally the posterior ganglia of the pharyngeal ring.
2051. The situation of the brain is essentially in front of or above the body, in opposition to the sexual parts, which are the lower totality.
2052. It is, however, above and behind; for it is originally situated posteriorly. The brain can therefore originate only, when the posterior medulla inclines from above forwards, making a curve in the latter direction; the brain is a spinal cord that has been bent from above forwards.
2053. The more the spinal cord is curved forwards, so much the nobler is it. This is self-evident.
2054. The brain is a spinal cord which makes the transit from the signification of bone to that of flesh.
2055. In the brain therefore the tendency must principally reside, to give off nerves, and perfect them into self-substantial nervous organs.
2056. In man the brain with its nerves curves round like a crosier, and that more perfectly than in any other animal. The spinal cord therefore in the highest formation of the brain returns again parallel to the direction, in which it has ascended.
2057. In the brain there is of necessity the greatest quantity of nervous mass. The brain is the nervous trunk, as the _liver_ is probably the vascular trunk.
2058. In the brain the cystic formation has been most purely represented; as e. g. in the cerebral cavities or ventricles. The brain is the stomach of the nervous system or its lungs.
2059. The brain consists essentially of two substances, of one accommodated to the flesh, and one to the bones, or of one arteriose, and the other venous. The former is the gray or cortical, the latter the white or medullary substance. The cortex is the lung of the brain, the medulla the liver or the intestine.
2060. The bark or cortex is the polarizing, active, oxydizing; the medulla the patient or suffering.
2061. This nervous pulmonary substance is continued along the spinal cord and even along the nerves, there as veritable gray substance, here as the vascular membrane of the nervous mass.
_Head._
2062. The brain, as being a system that has been separated superiorly from the other systems, determines the _Head_. The head is only there in so far as a brain is there.
2063. Head and trunk are antagonistically disposed, as Animal and Vegetable, or still more exactly, like nerve and bony flesh are to the viscera.
2064. The head is naught but a nervous organ.
2065. The concomitants of the nervous mass follow the brain, but, instead of the medulla having been previously subordinated to, or at least co-ordinated with, these, it is they that are thus related to the brain. The bones of the brain are the brain-case or cranium, the flesh of the brain, the face or countenance. On the head, bones and flesh have been disposed in the strictest manner according to their dignity or worth. Posteriorly there is almost pure bone, in front almost pure muscle.
2066. The cranium can be none other than the vertebral column continued around the brain. It consists of three vertebræ, the face of one. This will become clear in what follows.
2067. If the bones of the head are the repetition of those of the trunk, so also must the flesh of the head be a repetition of that of the trunk. Pectoral and abdominal muscles are ennobled in the muscles of the face.
2068. The face must have been principally formed by the orifice of the intestine--the mouth, and by the opening of the lungs--the nose, and by the apex of the vascular system--the members which are repeated as jaws. The month is the stomach in the head, the nose the lung, the jaws the arms and feet.
2069. The salivary glands are the liver in the head, as the mouth is its stomach. The liver that was originally also symmetric in form has become wholly symmetric in the higher organized head and forms two glands. The salivary ducts are the hepatic or biliary ducts.
2070. The tongue is the pharynx elongated upon the anterior side, because in front there is more flesh. The tongue is the extremity of the intestine converted into muscle.
2071. The nose includes pectoral muscles, the mouth arthric muscles or those of the limbs.
2072. If pectoral and abdominal muscles are repeated in the face, so also must the anterior bones, ribs, and limbs be repeated. It will be shown, in treating of the organs, that the nose is a vertebra, the jaws members, and their muscles those of the limbs. The head is the whole trunk with all its systems. The brain is the spinal marrow, the skull the vertebral column, the mouth intestine and abdomen, the nose lung and thorax, the jaws are members.
_Senses._
2073. The perfect animal again consists of two animals, the spiritual or solar, and the terrestrial or planetary. The animal nervous system does not continue to remain simply in the service of the other systems, but seeks also to gradually render itself self-substantial or independent. Now, the operation of the nervous system for itself is sensation. The parts of the nervous system having become self-substantial, will be therefore pure organs of sensation. Yet as the nervous system cannot emancipate itself from the other systems, so will its highest development be attained only in combination with the highest development of the other systems. There are therefore as many stages of the self-substantial nervous development, as there are special anatomical systems.
2074. Sensation must be modified according to the processes performed by those systems, with which the nervous system combines. These systems are, however specifically distinct from each other. Sensations that are specifically distinct are sensorial sensations. Organs of sense are accordingly the combination of the highest part of an anatomical system with the nervous system. Sensorial sensations are different processes of the anatomical systems perceived in the nervous system.
2075. The first combination of the nerves with the vascular system that has become free, or with the integument, is the sense of feeling--vascular sense. The intestinal system emancipated and combined with the nerves, is the tongue--gustatory sense--intestinal sense. The lung upon its highest evolution with the nervous system is the nose--olfactory sense--pulmonic sense. These are thus the sensorial organs of the vegetative systems--senses of vegetative life.
2076. There are indeed three animal senses; but as the osseous and muscular system form in their conjoined operation but one system or the motor system--there can be therefore only 2 animal senses. The osseo-muscular or motor sense is the ear. If the nervous system becomes wholly self-substantial, the nervous sense thus originates, or the eye, in which the brain itself has been planted outwardly, and acts independently of all other systems.
2077. The vessels form the general system, and therefore the tegumentary sense surrounds the whole body. Its brain is the spinal cord.
2078. The four remaining senses are perfections of individual systems at their perfect extremity, and thus in the proximity of the mouth and the brain. They together form the head. The jaws and the tongue obtain their nerves from the medulla oblongata, and this is therefore the brain of the gustatory sense. The brain for the nose is the gray cerebral substance, because the olfactory nerves are its elongations. The ears obtain their nerves from the cerebellum, which is consequently the auditory brain. The eyes are developments of the great brain or cerebrum--optic brain. Such is the rationale and signification of the divisions of the brain.
2. OSSEOUS SYSTEM.
2079. The nervous mass consists of indifferent, deoxydized blood-globules. If these be peroxydized, then the highest oxyd of the planet is deposited in them, namely, the earth, and that indeed which was the last remnant in the order of their production, or the calcareous earth.
2080. Vesicles or cells replete with calcareous earth are globes. The osseous texture consists therefore of globes; is only a dense cellular tissue, and thus ranks nearest to the vegetable structure. The basis of the bones is at first a cellular gelatine, which, with increased oxydation, is converted into cartilage. Finally, calcareous earth is deposited in this cartilage.
2081. In the lower organized animals, who breathe for the most part by means of branchiæ, the acid combined with the calcareous earth is an inorganic, or the carbonic acid, i. e. oxygen combined with carbon, or the earthy Inflammable; in higher animals it is an organic acid or phosphoric acid, i. e. oxygen combined with phosphorus, or the aerial Inflammable. Phosphoric acid may be regarded as peroxydized gelatine, as acid of gelatine. The bone is therefore earth, salt and Inflammable.
2082. The first appearance of the osseous mass is in the oxydizing organs. It is formed from the dense or internal coat of the arteries, since in old age bony lamellæ are deposited upon this. In the hearts also of many animals bones are formed.
2083. The first regular formation of bones is exhibited in the trachea or air-tube, which has been directly exposed to the oxydizing process of the air. These first forms of the bones are rings.
2084. The antetype of the bone is, however, the intestine, as the air-vessels are the antetype of the nerves. The bone is a tube, an ossified intestine.
2085. There are two osseous systems, a vegetative and an animal; the one surrounds the tegumentary systems, as, e. g. the scales of Fishes and Reptiles, horny rings of Insects; the other the nervous systems.
a. _Vegetative Osseous System._
2086. The vegetative osseous system is divisible into dermal, tracheal, intestinal, and vascular bones.
2087. The dermal bones are tegumentary rings, which surround the whole body, and are tracheal rings in so far as the skin is originally a respiratory organ. Such are the rings of the body in Insects, the shells of the Gasteropods and Molluscs, with scales and scutes in general.
2088. The tracheal bones are the branchial arches and tracheal rings.
2089. The intestinal or splanchnic bones are tubes environing the intestine, as in the corals, or imperfect annular segments, which at one time are found in the stomach, as in the Mollusca, at another in the pharynx, as in the Worms, Snails, Sea-urchins, and Holothuriæ; constituting what are called pharyngeal maxillæ. The branchial organs are fundamentally also none other than pharyngeal rings. The lingual and palatal bones, with the intermaxillary bones, belong also to the same category.
2090. The vascular bones are displayed in the hearts of many animals. The three last divisions may be called visceral or splanchnic bones; and then we have tegumentary, splanchnic, and nerve-bones.
b. _Animal Osseous System._
2091. The animal or nervo-osseous system must separate from the vegetative system of bones, and be placed upon the side exposed to the light. The side of the inferior animal that is exposed to the light, or averted from the earth, is the upper surface, dorsal region or _back_.
2092. The back holds the same relation to the ventral side as light does to the darkness, as sun to the earth; therefore the dorsal side is of a dark, the ventral of a faint or pale colour.
2093. Back and belly are related polarwise to each other.
2094. Through the medium of the bones the distinction between back and belly has been definitely established in the animal, and, as a consequence thereof, the distinction also of right from left. Before a formation of bone exists, the animal is for the most part a round cylinder.
2095. The osseous system can in itself be only symmetrically constructed.
2096. The osseous is the only symmetrical system in an animal. The other organs are so only in so far as they follow the arrangement of the osseous system.
2097. The animal osseous system is, from its being a repetition of the intestinal canal, a tube. This tube is surrounded, like the trachea, by rings, between which the tegumentary tube suffers constrictions.
2098. The back is a series of numerous bony rings.
2099. These bony rings are the _bodies_ of the vertebræ.
2100. The vertebræ have originated through polar repetition, through the muscular cysts.
2101. In addition to the series of vertebræ on the back, a vertebral column is formed moreover along the ventral surface, and without doubt there only where the air-organ, the branchia or lung, is situated. This inferior vertebral column is the breast-bone or _sternum_.
2102. All the systems, even those that are of a subordinate character in the animal, follow the direction of the main vertebral column. The intestine, as well as the vessels, are placed in accordance with it. Thus the principal trunks of the vessels take their rise along the vertebral column, the other vessels being given off from them in this situation, like the lymphatics from the intestine.
2103. The vascular ramules which surround the intestine and the skin, run out from a main stem, and are directed in a symmetric manner downwards and upwards (in the horizontal body of animals), or towards the belly and back.
2104. If new bony rings originate, they must also take these directions. They accompany the vessels that run in a circle, as the vertebral column accompanies the vascular trunks. These annularly-disposed bony twigs or apophyses constitute in the direction downwards the ribs, in that upwards the vertebral arches. Anterior and posterior to the vertebral column there consequently originates a long canal formed by the bony rings. In the anterior canal lie the galvanic or vegetative organs, in the posterior or upper, the organs of light must be situated "par excellence." The former of these canals is called the thoracic and abdominal cavity, the latter the vertebral canal. The vertebral canal is not the bony or medullary cavity itself, but it has been formed by several bony cysts in the same manner as the thoracic cavity. It consists of the body and the two arches. These are thus posterior (superior) ribs. The vertebral canal has the same signification as the thoracic cavity has; it is only a posterior thoracic cavity. It therefore contains, like the anterior canal, viscera dissimilar in kind to bone; the one including the spinal cord, the other vessels, intestine, and lung.
2105. The bony cysts that have originated through constriction do not all harden into calcareous matter, but they remain as alternating membranous cysts. Between the rings there are permanent cysts. The membranous cysts form the _joint_, or articular capsule. An articular capsule is a bone which has remained soft.
2106. This change in the ossific process takes place through the attachment of muscles, concerning which we shall treat in the sequel.
2107. The whole osseous system is consequently a symmetrical arrangement of several polar cysts and rings.
2108. The vertebra is not a single ring, but is at once a tolerably compound osseous system. The whole osseous system is nothing but a vertebra repeated.
2109. The number of the vertebræ necessarily conforms to that of the pairs of nerves or ganglia of the spinal cord; for they are indeed only the periphery or envelope of the latter. The number of nerves is, however, adapted to that of the organs, which they have to take care of.
2110. Now, the nervous organs are the senses. There are consequently as many divisions of vertebræ as there are senses. Thus there are vertebræ appertaining respectively to the senses of touch, taste, smell, hearing, and sight. Now as the four latter senses make up the head, but that of touch or feeling is distributed over the whole body, and superintended by the spinal nerves, the vertebræ thus divide into two principal sections, into vertebræ of the head and trunk. The number of cephalic vertebræ is 4; namely, nasal, ocular, lingual, and auditory vertebræ.
2111. To a perfect vertebra belong at least five pieces, namely, the body, in front the two ribs, behind the two arches or spinous processes; every vertebra of the head consists therefore of five pieces also. In those vertebræ which are removed from the respiratory organ, the ribs are smaller, as in the ventral ribs, and anchylosed with the body, as in the cervical vertebræ, where they are represented by the perforated transverse processes, and in the lumbar vertebræ they disappear entirely.
2112. The formation of the cervical vertebræ, where the ribs have been impacted or interposed between the body and spinous processes, is continued into the cranial vertebræ. These are only expanded cervical vertebræ. At the base of the skull four vertebral bodies lie in a series one behind the other; the body of the occipital bone, the two bodies of the sphenoid, and the vomer. Upon the sides of each of these bodies are situated alar processes, which correspond to the transverse processes of the cervical vertebræ, or to the ribs; e. g. the articular heads or condyles of the occipital bone, the alæ majores and minores of the sphenoids, and the two sides or lateral surfaces of the vomer. Behind these are placed the two broad cranial bones, which correspond to the spinous processes; as the occipital ridge or crest, the parietal, frontal and nasal bones. The occipital vertebra consists of the body, the two condyles and the occipital crest. The parietal vertebra consists of the body of the posterior sphenoid, the alæ majores, and the parietal bones. The frontal vertebra consists of the body of the anterior sphenoid, the orbitar wings or alæ, and the two frontal bones. The nasal vertebra consists of the vomer, the ethmoid and the two nasal bones. The occipital vertebra is the auditory vertebra; it incloses the auditory bones and the cerebellum, which gives off the nerves of hearing. The parietal vertebra is the lingual vertebra; the maxillary and lingual nerves passing through its alæ majores. The frontal vertebra is that belonging to the eye; through the orbitar plates or wings the optic nerves pursue their course, and it environs the cerebrum, from which these nerves originate. The nasal vertebra contains the olfactory nerves.
2113. Each cranial sense has thus only one vertebra, and the skull will consequently be formed of four vertebræ, whereof three appertain to the cranium, one unto the face. (See Oken's Ueber die Bedeutung der Schädelknochen, 1807.--Isis, 1817, S. 1204.)
2114. Several vertebræ are, however, found for the sense of feeling or touch, because it includes all the organs of the trunk. There must be therefore as many vertebræ in the trunk as there are particular organs placed therein. Of these there are three, the respiratory, digestive, and sexual system, or thorax, abdomen, and pelvis. To the thorax belong the neck, the arms, and the entire set of ribs. 5 vertebræ must appertain to the arms, because they have 5 digits and 5 nerves. But the ribs, and consequently also the digits, are determined by branchial vessels, are only repeated branchial arches, whose number in almost the entire class of fishes is 5. There are therefore also 5 thoracic or pulmonic vertebræ. Since the larynx consists of the 5 original branchial arches, and lies in front of the neck; so must the 5 superior cervical vertebræ stand in the signification of branchial vertebræ. The odontoid process of the second vertebra must be regarded, from its being separated from it in the foetus, as a particular cervical vertebra. Accordingly, in the Thricozoa there are eight cervical vertebræ. The 3 inferior cervical, and the 2 upper costal or rib-vertebræ, give off through their interspaces the nerves of the arms, and are consequently the brachial vertebræ. The 3-7th rib are thus appended to the 5 proper thoracic vertebræ, which stand in the signification of the pulmonary vertebræ. To these succeed the 5 short ribs which belong to the abdomen; their vertebræ are thus intestinal vertebræ. The succeeding vertebræ belong to the sexual system, and are indeed the 5 lumbar or pedal vertebræ, because they furnish the pedal or foot-nerves, the 5 sacral vertebræ being the proper sexual vertebræ. The coccygeal or caudal vertebræ correspond to the cervical vertebræ, and are present for the sake of the sexual branchiæ; usually one and the other is arrested. Thus there are,--
3×5 Respiratory vertebræ. 3×5 Sexual vertebræ. 1×5 Digestive vertebræ.
The number of the sensitive vertebræ is consequently 7×5=35, which are distributed into three groups, in accordance with the principal cavities of the body, whereof the 2 terminal groups consist of 15, but the abdominal group, which combines or unites them, only of 5. The body is accordingly not merely laterally, but also in length, a perfectly symmetrical structure, which has been parcelled out in the following manner into its five stockworks, stories or floors:
I. _Dermal vertebra._ A. _Sexual vertebræ._ a. Coccygeal vertebræ 5, b. Sexual vertebræ 5, c. Pedal vertebræ 5.
B. _Abdominal vertebræ_ 5.
C. _Thoracic vertebræ_. a. Pulmonary vertebræ 5, b. Brachial vertebræ 5, c. Cervical vertebræ 5.
II. _Auditory vertebra_ 1. III. _Lingual vertebra_ 1. IV. _Optic vertebra_ 1. V. _Nasal vertebra_ 1.
This regularity is found too only in the human skeleton. The animals are irregular men. (Vide Oken's Zahlengesetz in den Wirbeln. Isis, 1829, S. 306.)
_Cavities of the Trunk._
2115. The osseous system forms the trunk or body, because it follows the vascular system; the two other galvanic systems, the dermal and intestinal systems, form the large portions of the trunk or its cavities; to them is added the sexual cavity or pelvis.
2116. There are thus three truncal cavities, a pulmonary, an intestinal, and a sexual cavity, or thoracic, abdominal, and pelvic cavity.
2117. The osseous system will develop itself most feebly around the abdominal, because it is the indifferent cavity. Therefore there are either no ribs at all, or they are so short that they do not reach to the anterior vertebral column or the sternum. The short or false ribs are, according to their physiological sense, _abdominal_ or _splanchnic_ ribs. The thoracic ribs must be perfectly developed, i. e. abut against both vertebral columns, be entire ribs; the entire or perfect ribs are thoracic or _pulmonary_ ribs. The sexual ribs are arrested on the pedal and coccygeal vertebræ, but on the proper sexual vertebræ, namely, the sacral bone, they are still present as rudiments.
3. MUSCULAR SYSTEM.
2118. As the intestinal system reappears in what is animal under the condition of a vertebral column, so also does the aggregate of the vascular system ascend, and the vessels become animal.
2119. The animal vessels are the muscles, or filled-up vessels. The polar process enters the body through the vessels; thereby the tubes obtain two strong poles, and are drawn out lengthways. They are _fibres_, and consist of a series of strongly oxydized blood-globules.
2120. The fibre is chiefly apparent in those vessels, such as the arteries, in which the influence of air operates more forcibly. Now an artery has, in addition to the external cellular coat, two coats, like the first animal body, or one wall turned towards the mucus, and one towards the air. The internal arterial wall is enteroid, the external dermoid in character; the one being simply granular, the other fibrous membrane. The two membranes separate into two cysts or tubes, which likewise adhere within each other like intestine and skin. The external will become fibre, the internal bone.
2121. There is a vegetative and an animal fibre-or muscle-system. The one is associated with the tegumentary formations, the other with the bones and nerves.
a. _Vegetative Muscles._
2122. The muscles of vegetative life are simply fibrous tunics, as in the arteries, and are found in the skin, in the intestine, and in the vessels.
2123. The tegumentary muscles lie under the skin, and are inserted into it, or into the dermo-osteous system, when such an one is present. If the fibrous membrane be strongly developed under the skin, then it receives the name of a _Panniculus carnosus_.
2124. The intestine has also its fibrous coat, which, upon the stomach, anus, and pharynx, is frequently developed as a panniculus carnosus.
2125. The same holds good of the vessels, especially of the arteries and trachea.
2126. The fibres are either elongated or else annular fibres. The latter obtain the preponderance at the extremities of the tubular formations, as on the pharynx, anus, lips, and eyelids.
2127. We have consequently a dermal, splanchnic, and osseo-muscular system.
b. _Animal Muscles._
2128. In the oxydizing part of the vascular system the formation of fibres must be preponderant over that of cells, and thus at the root of the lungs.
2129. The vessel becomes in the lungs a fleshy cavity. The fleshy vessel is the _heart_.
2130. The heart is a vascular fragment, with a preponderating development of fibrous membrane.
2131. This fibrous membrane is developed at a spot where all vascular systems enter into mutual proximity, such as the respiratory ducts and the intestinal lymphatic vessel, the artery and the vein.
2132. In the union of all the highest galvanism is attained, and then the formation may launch out into what is animal.
2133. The heart is the animal in the plant.
2134. The first heart is an arterial heart. There is originally no venous heart. In the embryo, particularly in that of the bird, this is extremely distinct; the arterial heart also emerges for the first time, and per se, in the lowest animals, in the Molluscs, Snails, and even in the Fishes, although in the latter it is regarded as a venous heart.
2135. The arterial heart is the central, the venous heart the peripheric.
2136. The heart is the prototype of the muscular system. All muscles must be a metatype of the heart.
2137. The muscle is hollow. It is a cyst.
2138. The muscular system is a manifold and serial juxtaposition of fibrous cysts or of hearts. In this respect the muscle has been formed in a corresponding manner to bone. Both are rows of cysts.
2139. But the muscle, as being the external fibrous tunic, is the enveloping or external cyst.
2140. In idea, the muscle can only directly envelop the bones, and not the other parts, for it ranks upon the same grade of development with the bone; it is the arterio-fibrous wall, while the bone is the internal arterial wall.
2141. Bone and flesh stand in antagonism like air and earth. The muscle is that which is polarizing--moving, the bone what is polarized, moved. The muscle is heart, the bone the moved blood. Bone and muscle are related as that which is contained and what is containing. The muscle is the wall of the cyst, the bone, the fluid which has been secreted from it and rigidified.
2142. As therefore the muscle is an individual cyst, a heart, which cannot invest the whole body like a single large cyst, so also must the muscular contents be only a discrete cyst. The uninterrupted character of the bones depends therefore upon that of the muscles, and the latter upon the meaning of the heart.
2143. _A physiological rationale of the joints is accordingly afforded in the heart._
2144. A bone is a rigidified, ossified heart; the osseous system is a series of mutually dependent, alternately ossified and non-ossified (arteriose and venous) hearts.
2145. The muscular cyst includes the soft bone, or the joint.
2146. At its two extremities there is preponderance of oxydation, whereby the soft osseous cysts are combusted into hard calcareous earth.
2147. The muscle is a cause of the alternating ossification.
RELATION TO THE OSSEOUS SYSTEM.
2148. As the formation of bone predominates upon the side exposed to the light or the nervous, so does that of muscle upon the shady or vascular side. The abdominal side of the animal is the muscular, just as the dorsal side is the osseous. Upon the thorax, abdomen, on the members which belong to the anterior region of the body, and on the face, the muscular layer is by far the most predominant. Posteriorly, however, or upon the back, it is slightly wanting, and the bones there project.
2149. The back is related to the anterior surface (when regarded in man) as bone is to muscle. What is _in front_ is muscle, what is _behind_ is bone. The anterior side is therefore more active, nobler, more powerful and more spiritual than the posterior. Posteriorly stands the earth inert and rigidified, while in front is the air in ceaseless capacity for, and actually in, motion. The anterior muscular layer is more active and powerful than the posterior.
2150. In every muscular cyst there are two kinds of layers, an anterior and a posterior, or stronger and weaker layer.
2151. The stronger layer is the flexing, the weaker the extending. For the limbs have been necessarily bent forwards. The direction alone of the joints at once resides in the structure, which is even determined by these relations. Such a muscular layer, which mostly consists of several bundles, is termed a muscle.
2152. A muscular cyst consists of flexor and extensor muscles. The individual muscle is therefore only a piece of a cyst, and therefore not in itself hollow. It is only an entire muscular layer of flexors and extensors which is the pattern of the heart. The flexors are the strongest and are placed in front, the extensors lie behind.
2153. In the heart the flexor layers have not yet separated from the extensor layers, because the vegetable flesh has as yet no symmetry in itself.
2154. Flexors and extensors occur in pairs; because the osseous system is in pairs.
2155. There resides in the osseous and in the muscular system no cause for a diversity in the two halves of the body. Thus, if there be a difference, it must only reside in the asymmetrical galvanic systems.
3. _Organs._
2156. Organs are parts of an anatomical system, which separating, combine with a part of another system, and thereby obtain a peculiar or special function.
2157. For each system therefore there are as many organs, as there are possible combinations of systems. There are vascular, intestinal, pulmonary, sexual and tegumentary organs, with furthermore, osseous, muscular and nervous organs.
A. VEGETATIVE ORGANS.
1. INTESTINAL ORGANS.
2158. The intestinal or splanchnic system falls first of all into three great divisions, into that of the viscera, the sex and the head; the visceral intestine separates moreover into pulmonic, venous and tegumentary intestine, and this in constant accordance with its combinations and functions.
a. _Visceral Intestine._
2159. In the digestive system it is the chemical process that takes precedence. This, however, divides into three moments, into that of _solution_, _separation_ and _formation_, or crystallization, which is here _absorption_. Thus the intestine separates also into a solvent, secernent and absorbent intestine, and that indeed through its combination with the pulmonic, vascular and tegumentary system. The solvent viscus is the stomach, the second or secernent, the duodenum, the third or absorbent, the small intestine (jejunum and ileum).
_Pulmonic Intestine._
2160. All solution is accompanied by oxydation. The gastric juice is in its action an acid.
2161. The gastric juice obtains its oxygen from the spleen. The spleen is the stomachic lung. This is attested by its position and close approximation to the stomach; by its black, venous, and deoxydized blood, which in certain diseases has been secreted even in the stomach; by its want of any excretory duct; by its tissue, which resembles that of the oxydized placenta; furthermore by the natural character of this function when contrasted with the unnatural quality of other useless functions that have been assigned to it; and lastly too, by the fact that, without this opinion be admitted, it must remain a superfluous organ, and as regards use, unknown. After a series of years, during which this doctrine has been combated upon all sides, without, however, a single reason, save and except that it was not believed, being advanced by its opponents, I must still persist in the correctness of the above view.
_Vascular Intestine._
2162. In the duodenum the _analysis_ of the food takes place through the medium of the bile. It is consequently the _biliary_ intestine or stomach.
2163. The biliary intestine does not stand upon a par with the other intestines, but has an equal rank with the stomach. It is therefore not confined within the mesentery, but can expand itself like the stomach; it has its vessels and nerves. In it the separation of the chyme into chyle and excrementitious matter takes place.
2164. What the spleen is for the stomach, that is the liver for the duodenum; it is the hepatic stomach, and consequently the vascular stomach.
2165. The liver is the ramification of the intestinal canal along with the whole vascular system.
2166. Now, as the analysis is the principal function in the whole process of digestion; so is the liver the chief organ of all digestive organs.
2167. The liver is the centre, the brain of the digestive system, because it is the blossom, the synthesis of the vascular system. From it everything emanates, and upon it everything which concerns digestion, ay, the whole body, retrogressively operates. If the liver suffers or undergoes functional derangement, the whole vascular and tegumentary formation then becomes a liver, as is exemplified in the disease called jaundice.
2168. The bile effects the analysis or separation through its basic or alkaline character, since it combines with the acids of the chyme, and thereby forms the excrementitious matter.
2169. That which mediates between the acid and what is alkaline is the fluid secreted by the pancreatic gland. The pancreatic gland is the ramification of the intestine along with the arterial system.
_Tegumental Intestine._
2170. That in the jejunum and ileum or the small intestine proper, and in that alone, absorption and thus the tegumentary function, but nothing else, occurs, is well known. By this absorption the chyle becomes removed from the intestine, so that the excrement alone is left.
b. _Sexual Intestine._
2171. The sexual is pre-eminently that which is exsecernent or excretive, since one sex strives to redintegrate itself upon the other, becomes ingestive for the other, but egestive for or in relation to itself. It is therefore an essential property of the sexual parts, that they secrete and excrete.
2172. Every galvanic system, which has been adjoined or appended to the sexual parts, is excretive. They are, taken in a rigid sense, the only system of excretion. The co-operative processes of the sex are those of the vegetative systems, but they take place in the reverse direction. The latter convey inwards, the former outwards. The kidneys are a reversed or evacuating lung, an excrement-forming liver; the urinary cyst is an exspirant, a trachea containing refuse matter; the urethra is a glottis reversed; their diseases are therefore similar. They thus expel products of the individual respiratory system.
2173. And in this is the sexual apparatus distinguished from them, namely, that it expels the products of _all_ systems, the products of the whole organism, or the organism itself. In the semen the whole male body with all its parts transudes, or passes over in a fluid state, into the female parts; in a child, the female together with the male body, passes over--formed or fashioned into the world.
2174. The sexual intestine must therefore be expulsive also. It is that which conducts the intestinal juice and the refuse of the aliment out of the body.
2175. The evacuating or efferent intestine is the large intestine. It is consequently the _Sexual intestine_.
2176. The large intestine is related to the abdominal or small intestine, exactly like the urinary cyst is to the kidney, and this to the vascular system. The small intestine passes therefore by perforation into the large, not the latter into the former. The small enters by perforating the large intestine, and therein evacuates its alimentary residue, as into a special-cyst, that has nothing to do with the intestinal system. The large intestine is the _excrement_-cyst, just as that which has been called bladder is the _urinary_ cyst.
2177. The large intestine no longer digests, but only receives what is left from the process of digestion and expels or casts it out.
2178. The large intestine begins with a cæcal extremity, or with an obtuse cyst, and opens into the anus, exactly like the original animal cyst, the polyp. The blind extremity is called cæcum. With this the small intestine communicates by perforating it at an acute angle, and that indeed in a direction which runs towards the obtuse cystiform extremity; so that both intestines lie in mutual juxtaposition like a fork, whereof the pharynx and the rectum are the two prongs, while the cæcum is the handle.
2179. The two intestines are consequently by no means correlated. In the perfect animal there are two intestinal systems thoroughly distinct from each other; two intestines, which belong to two different animals, the sexual and encephalic animal, or the plant and the animal. The genesis of the large intestine and all its relations, which are principally seen in the cæcum and rectum, plead for this philosophical derivation of the two intestines.
2180. The rectum appertains wholly and entirely to the sexual system, and especially to the uterus. It is devoid of mesentery, and supplied by special vessels; it stands in most obvious sympathy with the uterus, with its diseases, and with menstruation. Even hemorrhoids are a sexual disease, a disease of the _sexual_ intestine.
2181. The anus is thus the intestinal mouth of the sexual animal. In the lowest organized animals, the oviducts seminal and urinary ducts together run into it, as into a proper mouth. The anus is a true oral cavity in the fishes, reptiles, and even in the whale.
2182. The pharynx opens into the mouth, and so does the rectum into the anus; the trachea opens into the month, and so does the urethra into the anus; the salivary ducts open into the mouth, and so into the anus do the oviducts, with the spermatic canals in the lower animals. The rectum moreover lies behind the urinary cyst, like the gullet behind the trachea.
2183. The sphincter or occlusor muscles of the anus have been formed like the sphincter muscles of the pharynx. The anus is a mouth without a head, and therefore a mouth without lips, or an pharynx.
_Cephalic Intestine._
2184. The union or combination of the intestine with the animal systems, such as the nervous, muscular, and osseous, occurs in the head.
2185. In so far as it combines with bones and muscles, it becomes an organ of motion, and, when with the nerves, an organ of sensation. The animal or sarcal intestine is pharynx and mouth.
2186. The motor organ is an organ of prehension. The organs of prehension are endowed with independent motion, and therefore move towards the food. The first general organs of motion are the members of the body. In the higher animals the thoracic members are at once the prehensible organs. Instead of the aliment being obliged to flow, as unto the plant, in a state of suspension in water, the animal moves itself toward its food.
2187. The members are the first organs of prehension. But these are repeated in the head as jaws and teeth. The teeth are the second organs of prehension, but the first which belong to the cephalic intestine; they are called _organs of seizure_ or prehension.
2188. The organ of digestion is, however, one of a chemical character. It must be therefore repeated as such in the head. This is shown in the _salivary glands_. The saliva is the animal gastric juice, and is therefore directly solvent in its properties. It is _poison_.
2189. After and during the operation of the saliva, the aliments in the mouth are subjected to the action of the molar teeth and ground or _chewed_. These comminuting or crushing organs are only a repetition of the act of prehension, and consequently belong to the prehensile or biting organs.
2190. The mouth is the stomach repeated in the head.
2191. The union of the intestine with the nervous system is the _tongue_.
2192. The intestine, when repeated in the head's muscular system, is the organ of deglutition, as is seen in the pharynx and oesophagus.
2193. The prehensile and manducatory, venomous, gustatory, and deglutitory organs, are the forms into which the intestinal system divides, when it is repeated in the after-animal, or in the sphere of animal life. The gustatory organ is the nerve-intestine; the prehensile organ the bone-intestine; the organ of deglutition the muscular intestine; while the poison-organ is the proper cephalic intestine, or the stomach.
2. VASCULAR ORGANS.
2194. The vascular system regarded "en masse," or in a general sense, has to participate in the nutrition of the body; so far it takes the place of the cellular tissue, and cannot therefore be developed for itself into any special _organs_. If, however, certain vessels separate from the general system, and combine with other systems unto the performance of special offices or functions, _organs_ then originate, which, nevertheless, do not, in a rigid point of view, belong to these systems.
2195. There are therefore as many vascular organs as there are possible combinations; thus with the tegument, the lung, the intestine, the sexual parts, and with the systems of animal life.
a. CUTANEO-VASCULAR ORGANS--BRANCHIÆ.
2196. The development of the vessels unto a special organ in the integument are _Respiratory organs_; or, more properly speaking, the development of the integument to constitute a special organ in combination with the vessels, is a respiratory organ.
2197. In the commencement the branchiæ are only a vascular network upon, and therefore subordinate to, the integument. They pass, however, through all possible stages of development, until they have subjected and converted also the integument into a vascular system; a point which is attained in the formation of the lungs.
2198. The branchial membrane already commences in the earthworm to concentrate itself and dilate, so as to form what have been called the sacculi or pouches; in the leech it is folded inwards, so as to form lateral vesicles or cysts, that prognosticate the air-passages or stigmata of insects; in the Nereids the vessels project upon the back from above the integument as free branchial ramules, a formation, which is again found in the nudibranchiate Gasteropods.
2199. These branchial ramules form at first two rows extending along the whole dorsal region; by degrees, however, the posterior set disappear, and the cervical branchiæ only are left, as antetypes of the gills of fishes.
2200. In the Mollusca the branchial vessels unite to form laminæ upon the ventral sides, and are already surrounded by a kind of thoracic cavity, the mantle, as is exemplified in the tectibranchiate Gasteropods. Here again the branchiæ are either a simple vascular net enveloped in a fold of the mantle, or elongated into filaments disposed like the teeth of a comb, or into laminæ, &c.
2201. The lateral branchiæ of the Nereids usually emit filaments resembling feet, upon the root of which the true branchiæ are then placed. In the Crustacea these filaments harden into actual feet provided with joints. The feet are therefore nothing else than branchial filaments, which have lost their vegetative function.
2202. In many worms the same branchial filaments are converted into hairs or bristles, which are therefore none other than desiccated branchial filaments.
2203. Even the hairs of Mammalia and the feathers of Birds have been left as remnants of the original branchial formation.
2204. Where the branchiæ have assumed the laminated form, they are surrounded by a similarly formed covering or operculum. The shells of the Mollusca are, as regards their signification, none other than branchial opercula, or gill-covers; the same holds good also of the gasteropodal and crustacean shells, and at bottom of every calcareous and horny covering of the body.
2205. To the same category belong also the opercula of Fishes, and even their scales. Fundamentally, the whole epidermis is only a product of respiration or oxydation.
2206. The first saccular or cæcal inversion of the integument, as in the Leeches, Molluscs, and Snails, is at once a predominance of the tegumentary, obtained through the medium of the branchial formation, whereby the skin commences to become a self-substantial organ of respiration.
2207. In the Scorpions the branchiæ are introverted sacs or cysts, into which, however, instead of water, air already enters.
2208. This insaccation is converted in the Spiders into more distinct air-cysts, which finally ramify in the higher insects and become true tracheæ.
2209. Lastly, the respiratory system obtains the upper hand to such an extent, that, together with internal air-tubes, external branchial laminæ are also developed, as in the Molluscs; but in them the tracheæ obtain the preponderance over the blood-vessels, so that these laminæ dry up and become wings or fins.
2210. The wings of Insects are branchial laminæ, converted into air-organs.
2211. The wing-coverings or elytra are branchial opercula, and correspond to the shells of bivalve Mollusca.
2212. Every insect therefore must properly possess four wings and two wing-coverings, of which last, however, rudiments only appear to be left in the nocturnal Lepidoptera.
2213. In the higher organized animals those gills only which are nearer the head are persistent, the posterior or lateral branchiæ being gradually arrested.
2214. These lateral branchiæ remain in Fishes as lateral mucous openings, which constitute the lateral line.
2215. The cervical branchiæ are limited to the number five, which has already begun to be established in the Crustacea; namely, at the origins or roots of the five anterior pairs of feet.
2216. The number five probably derives its origin from the vegetable kingdom, and that indeed from the genesis of the pinnate leaves, so that one kind of numerical law appears to prevail with respect to this organ in both organic kingdoms. It is probably also an imitation of the five organs of sense.
2217. The vessels of the branchiæ of Fishes are accompanied by bony rings, which correspond to the feet of Crustacea.
2218. All Fishes have, with few exceptions, five branchial arches.
2219. When in Fishes the sarcous system begins to produce viscera, then the five branchial foramina pass inwards, and only a single respiratory aperture is left for them in the fleshy body; namely, the external branchial foramen.
2220. In the lower animals water or air passes in and out through the same respiratory aperture, but in Fishes these two courses are preserved distinct; for, except in the Lampreys, the water enters through the mouth, and passes out through the branchial aperture.
2221. Here the attempt is manifested in a still greater degree, to bring the process of respiration wholly under the control of what is animal, this being in the next place, and for the first time attained, when respiratory apertures only are left upon the head.
2222. The respiratory apertures of the head are the nostrils, which are suddenly manifested in Fishes, but are in them simply subservient to the sense of smell, and not as yet to the respiratory act.
2223. All the higher animals have, like fishes, branchial apertures in the neck, only they coalesce and become obliterated at an early period, or so soon as that respiratory process appears, when air passes through the nostrils. In the Salamanders and Frogs these branchial foramina are persistent for a longer time, frequently through the whole period of life; but in Birds and Mammalia they disappear, while they are in the embryo state.
2224. When the branchial apertures close, the vessels separate from the arches, and betake themselves to a glandular body lying in front of them. The thyroid gland is the remnant of the former branchial formation, and is therefore found only in Reptiles, Birds, and Mammalia.
_Lungs._
2225. In Fishes the internal organ of respiration is already indicated by a cæcal eversion of the pharynx, which is surrounded by the branchial arches. This membranous cæcal eversion is called the swimming-cyst, which in the higher animals becomes double on account of their symmetry, and is then called lung.
2226. In the Fish the aquatic and aerial process of respiration are present together, the former being external, the latter internal.
2227. The branchial arches having coalesced, are converted in the higher animals into tracheal rings, into the larynx and the posterior cornua of the hyoid bone, if these are present, as in the Reptiles. The larynx is therefore no special organ, but only a remnant of the branchial respiratory apparatus.
2228. The laryngeal vessels are, like the thyroid glands, branchial vessels, and in Fishes, therefore, the branchial vessels do not correspond to the pulmonary vessels, but to those of the trachea. The pulmonic vessels of Fishes are the blood-vessels of the swimming-cyst, which convey blood directly into the heart, whereby this organ obtains the signification of the left or arterial heart.
2229. When the branchial apertures have coalesced, the nose then opens into the mouth or into the trachea, and in this way the nostrils assume the complete character of aerial foramina.
2230. The nose is therefore originally an organ of smell, and then a part of the respiratory system. It is the animal lung.
2231. As the secretion of bone is a product of the more powerful process of oxydation, so do the bony rings multiply beneath the branchial arches or the larynx, and are called tracheal rings. In the feebly respiring Reptiles the trachea is therefore still in a membranous condition, but in Birds and Mammalia, it is surrounded by many rings, or is a repetition of the larynx.
2232. In Birds even a kind of inferior larynx originates, which is provided with muscles and can produce tones.
2233. The ramification of the trachea into two bronchi or branches is constantly progressing, and at length these tubes divide into a great number of vesicles, which together form the lungs. The lung, which was in the commencement a simple saccular inversion of the integument, has now become a self-substantial organ, to which the respiratory vessels have been subordinated. The lung also divides upon each side into five lobes.
b. VASCULAR ORGANS OF THE INTESTINE.
_Liver._
2234. The self-substantial development of the vascular, and its separation from the general, system, is most perfectly attained in the _Liver_.
2235. In the liver, as being the vascular system, which combines with the intestinal canal, the venous system has become independent. The portal vein arises from the intestinal canal, collects into one trunk, and again ramifies in order to unite with the biliary ducts, which are only a ramified saccular eversion of the intestine. This conjunction is represented by the liver.
2236. The liver as being a venous organ stands therefore in opposition to the lung, and produces, instead of oxydes, a basic body, or the _bile_. The liver, as being the venous system which has become free, is to be regarded as the highest development or blossom of the vascular system.
2237. It is for the vegetative body, what the brain is for the animal; and hence, therefore, the similarity of structure and the sympathy between both organs.
_Spleen._
2238. As opposed to the liver, the arterial system also develops itself upon the intestine as a respiratory or branchial organ. These intestinal branchiæ are found in several of the lower organized animals, especially in the Holothuriæ and Aphroditæ.
2239. In the higher animals they are aggregated into a special organ, through which the gastric juice obtains oxygen; this is the spleen. The spleen is the branchia of the stomach; it has therefore no excretory duct and requires none.
2240. Lastly, the salivary glands also in the mouth as well as in the duodenum, the abdomino-salivary glands, and even the odoriferous glands on the rectum, such as the castor-and civet-pouches, are complications of vessels with intestinal ramifications.
C. VASCULAR ORGANS OF THE SEX.
_Kidneys._
2241. The vascular organ of the sexual system is the kidneys.
2242. As the urine is chiefly characterized by the urea and thus by a basic principle; it thus corresponds to the bile, and consequently the kidneys directly to the liver or the lung reversed.
2243. But there is also a sexual branchia in those inferior animals which breathe through the anus, such as many aquatic larvæ.
2244. The remnant of this in the higher animals appears to be the allantois, and in the body itself probably what have been called the primordial kidneys.
2245. The combination of the vascular system with the animal systems is the sense of touch.
3. RESPIRATORY ORGANS.
2246. The respiratory organ is a development of the integument.
2247. The perfect organ of respiration is an air-organ or _lung_; combined with the tegumentary system, it is a water-organ or _branchia_.
2248. There are _dermo-branchiæ_ or the _branchiæ_ proper, as in the Worms, Molluscs, Snails, and Crabs.
2249. Intestinal branchiæ in the Holothuriæ, constituting in the higher animals a _spleen_.
2250. The sexual branchiæ are probably the primordial kidneys.
2251. The branchiæ when united with the osseous system are the branchial arches of Fishes, which subsequently separate into larynx and thyroid gland.
2252. The self-substantial development of the integument into the respiratory organ is a lung.
2253. When united with the vascular system or the vegetative systems generally, the integument forms the tracheal system of Insects.
2254. The tracheæ are spiral vessels as in plants.
2255. The respiratory organ, united with the motor system, is the lung proper, which is situated within the thorax, and covered by ribs.
2256. The proper lung divides also, like the intestine, into two parts, into the cystiform, pharyngeal expansion of the larynx, and into the pulmonic substance, which is equivalent to the stomach, and wherein the analysis of the air takes place.
2257. The trachea, and especially the larynx, is moreover an entire thorax, a ribbed skeleton upon a small and membranous scale. In the larynx, the animal thoracic structure, consisting of ribs and muscles, resides prognosticated. The larynx has originated from the coalescence of branchial arches. The ribs are a repetition of the branchial arches.
2258. The diaphragm is a formation, which admits, by no reasoning from its anatomy, but only genetically, of being explained. Originally, the whole body was only an abdomen, upon the external surface of which the branchiæ were appended. A remarkable instance of this is at once afforded by the Snails, and also the Fishes. As the gills were converted into lungs, a special body or the thorax originated for them, which encroached upon the abdomen. Now, the abdominal wall left between these two cavities, was just the diaphragm.
2259. The diaphragm is not a transverse wall or septum. The idea of such a cross-rail gainsays all sound physiology. It has been the abdominal wall.
2260. The union of the lungs with the nervous system is the nose.
2261. The nose is the cephalic thorax; but it has also the thoracic contents, or the very lung repeated in itself.
2262. The multicavernous ethmoid bone is the lung in the nose, the two nostrils being the stigmata or foremost openings of the trachea. The nasal muscles are homologous to the cartilages of the trachea, and especially to those of the larynx.
2263. The velum palati is the diaphragm between nose and mouth, or between the cephalic thorax and the cephalic abdomen.
_Coverings._
2264. The animal coverings are desiccated respiratory organs appertaining to the integument.
_Capillary Vessels._
2265. The principal function of the vessel is that of secretion, whereby nutrition is imparted. This secretion must take place in the whole body, in so far as it is opposed to the lung. The vessels then pass over into the finest canals, and are called _capillary vessels_.
2266. The capillary system of vessels is an organ in antagonism to the lung; what enters the latter organ, passes out through the former.
2267. The capillary system of vessels is the property of the tegumentary system. It may almost be said that wherever there is integument, there capillary vessels are present. The most perfect evolution of capillaries is the integument. It is the proper exsecernent, in opposition to the intestine, which is the absorbent organ.
2268. Evaporation is the essential tegumentary process.
2269. The product of the evaporation is mucus.
2270. In the evaporation, however, the mucus becomes analyzed by the influence of the air and light.
_Epidermis._
2271. The external mucus exuding from the skin becomes oxydized, the inferior on the contrary reduced; the oxydized becomes vitreous and transparent. It is the _Epidermis_.
2272. With the maximum of oxydation the epidermis passes over into a vitreous transparent horn, e. g. _scales_.
2273. The scales, which invest the toes and digits are called claws, and finally become nails. The finger-nail is nothing but a scale, which in this situation has become particularly large and strong.
2274. That which is reduced beneath the vitrified epidermis determines the colour of the skin. With a demi-oxydation it is uncoloured, appears white. Where the integument is thin, the red colour of the blood appears through it; such integument is therefore white upon the whole, but red in individual places.
2275. With the most complete reduction effected by the highest operation of light, its substratum or under-layer becomes black. The mucus passes over into reduced carbon. Beneath the vitreous epidermis there is thus a metallic _pigmentary membrane_.
_Hairs._
2276. Capillary vessels, which simply convey mucus, but are elongated singly above and from out the cutis, are _Hairs_. The idea of the hair is a _capillary vessel_, the contents of which are no longer blood, but reduced mucus. It is an indifferent capillary vessel. The hair is hollow and contains an oil, which determines the colour.
2277. In the hairs the nutritive system is prolonged above the body.
2278. The hairs and scales are the general terrestrial system of the body determined by the air.
2279. Thus the earth has sprung up into a plant. Scales and hairs are like true vegetable leaves, which still continue their process in the animal; they can no longer it is true make their respiratory process hold good for the animal process itself, but are at present content with only oxydizing the material of evaporation. It is properly the integument only, not the entire body, which respires through hairs and scales.
2280. The hairs are dried branchial filaments, and therefore continue to occupy in Man, those situations only where in the lower animals branchiæ or tentacula are placed. Around e. g. the mouth, upon the head, under the arms or in the axillæ, and around the sexual apertures.
2281. The feathers are dried ramified branchiæ, pinnate leaves.
2282. The hairs participate in the electric process of the whole body.
2283. What determines the colour in the plant, determines it also in the animal; only in the plant the colour is more coarsely precipitated, and nothing therefore glimmers from the interior of the body, but all is green. In an animal, however, the colouring matter is transparent, and the interior is thus rendered visible.
2284. With the external coverings, such as scales, and lastly with the hairs, all organs of the trunk, in so far as they are derived from what is vegetable, are exhausted. The vegetable animal, as trunk, is completed, and we must now therefore turn to the sexual organs.
4. SEXUAL ORGANS.
2285. The sexual organs are developments of the integument upon a higher grade, and combinations of the same with the animal systems, like as the blossom is a repetition of all vegetative systems.
2286. They range in the middle between the vegetative and the encephalic animal, and are therefore a totality for themselves, or a sexual animal.
2287. There are vegetative and animal sexual parts.
I. _Vegetative Sexual Organs._
2288. Are special developments of the intestine, vessels and branchiæ.
a. SEXUAL ORGANS PROPER.
2289. The proper sexual parts are a repetition of the digestive system on its transition to what is animal, or to the organs of sense.
1. _Female Organs._
2290. The female parts are a floral capsule, consisting of an ovary or sac, stigma and ova. All higher development takes place, however, through separation of the complicated organs and processes.
2291. The three parts therefore of the gestative sac or matrix separate, each part becoming self-substantially perfected. The orifice elongates into a cervix or neck, which is gradually rendered more distinct from the matrix, and in its highest and self-substantial completion is called oviduct.
2292. The matrix elongates also at its cæcal extremity or fundus, as well as at its open end. The germinal bodies also become self-substantial, separate gradually from the fundus of the matrix, and are then independent ovaria.
2293. As in the highest animal formation they also assume the animal symmetry, while at first they were only single, or manifold, as in the plicated capsules of Star-fishes, and therefore remained stationary at the number two; so is the ovisac prolonged into two long cornua, or trumpet-like tubes, which at the commencement, indeed, still include the ova, as in Insects and Fishes, but are subsequently left entirely free.
2. _Male Parts._
2294. The leaf-formation is elevated to the corolla or the male parts in the plant. They are, however, only the repetition of the plant upon a higher stage. Hence in the animal also the male parts will be upon a higher grade of position to the female.
2295. As the stamina surround the ovarian capsule in the plant, so do animal stamina stand around the orifice of the oviduct; as _penes_.
2296. In the lowest ovisacs the penes are circularly disposed around the orifice; as is exemplified by the tentacles of a Polyp; by degrees, however, they are resolved, for the sake of symmetry, into two, and then project upon the sides of the orifice, as in the Serpents and Lizards. In the higher animals the two penes coalesce into _one_.
2297. This penis situated at the orifice of the oviduct is the clitoris. The vagina is separated from the clitoris; but in the male organs they both combine or blend with each other, and the vagina becomes the seminal canal or that of the penis.
2298. As the orifice begins to assume a male character, and the external parts to be more forcibly developed in the direction outwards, so, on the other hand, do the internal parts recede the more, and remain simply as ovaria or ovisacs, in which the ova, instead of being formed as such, dissolve into pollen, mucus, or male semen. These ovaria, which now secrete semen instead of vitelline vesicles, are called testes.
2299. The testes originate, while the mucous ova are being reduced to primary mucus, or to infusoria.
2300. What is male originates through an organic process of decomposition or putrefaction of the ova. The semen is an organic product of decomposition.
2301. The semen must contain infusoria. A semen which does not contain infusoria, is ovum-like, or feminine. Except at the time of heat or rut, and thus when the animals have a female character, the semen possesses no infusoria, and is in that case simply albumen.
2302. Semen, which is devoid of infusoria, is incapable of impregnation. How can one wretched female ovum impregnate or fructify another?
2303. As the trumpet-like tubes or oviducts belong to the ovaria, so are they developed with the testes, and now convey semen instead of ova. The female tubes become the vasa deferentia or _spermatic ducts_, the uterine cornua, the vesiculæ seminales.
2304. Between the vesiculæ seminales and the vagina or the penis the uterus shrivels up into the prostate gland, into which the vasa deferentia open, like the oviducts do into the uterus.
2305. As the matrix is the proper female organ, so will the spermatic ducts seek to combine with the penis, or at least open self-substantially upon the os uteri. The testes open either through the spermatic ducts into the vagina, as in Fishes, Reptiles, and Birds, or into the penis, as in the Gasteropods, Insects, and Mammalia.
2306. Male and female parts are therefore perfectly homologous, the former having a greater development of the external division, the latter of the internal.
2307. The female parts have undertaken the business of vegetation, or that of the viscera, the male that of the animal excitation.
2308. Since the male parts are no new formation, but the female parts themselves, only characterized by internal arrest, and external increase of development, so do male and female parts appear incapable of occurring together in any animal. Perfect hermaphrodites would, accordingly, be impossible; for where testes are, no ovaria could be, because the testes are themselves the ovaria, or the latter only changed.
2309. Hence androgynism would be possible only by one ovarium remaining as such, and the other being converted into a testis.
2310. This development appears only possible when the two halves of the body are unequal. Asymmetrical animals only can be androgynous. In the snails one of the two molluscan shell-valves has been more largely developed than the other, and therefore one half also of the body is greater than the other. On this account many hermaphrodites are found among these animals.
2311. Thus, there ought to be no androgynous animals with at the same time two ovaria and two testes. Nevertheless, this very peculiarity occurs in many of the lower organized animals, e. g. the worms.
2312. Accordingly, the principle of androgynism is in general to be found in the want of symmetry. Symmetrical animals are as a rule dioecious. No hermaphrodite is found among the Insects, Fishes, Reptiles, Birds, and Mammalia.
2313. If such occur, they are formations that have remained persistent at the lower stage of development, or upon the transition of the embryo through the Snail-type of organization; they are thus monstrosities, or malformations.
2314. These malformations also, when occurring in the higher animals, can never possess more than one testicle and one ovarium. The uterus then ranks midway between its condition as such, and that of a prostate gland; the spermatic tube or urethra opens after a female fashion below the root of the penis.
_Impregnation._
2315. Since the male sex is related to the female, as corolla to capsule, as leaf to stalk, as air to water, and as light to matter; so is it related also as integument to intestine, as lung to lymphatic vessel, as artery to vein, as nerve to flesh or muscle, as Animal to Vegetative.
2316. Copulation is therefore an _irradiation_.
2317. Already, in the course of the heavenly bodies, has the highest act of the animal, that of copulation, been preindicated or portrayed. The creation of the universe or world is itself nothing but an act of impregnation. The sex is prognosticated from the beginning, and pursues its course like a holy and conservative bond throughout the whole of nature. He therefore who so much as questions the sex in the organic world, comprehends not the riddle or problem of the universe.
2318. If the female parts have effected a complete transition into the male, so are the sexes necessarily separate and distinct.
2319. Since the male parts are the female that have been more highly developed, so there resides in the latter the constant conatus or effort to convert themselves into the male.
2320. This metamorphosis, or conversion, is, however, no longer possible in the female parts, when already finished or fully formed, but is only attainable in a new attempt being made upon their part, to transform the fluid mass into ova.
2321. Gestation or pregnancy is none other than the propensity of the Female to convert itself into the Male.
2322. The foetus is the male in a female, or the foetus is the male sexual parts in the female.
2323. In idea, every foetus should be male. But, if mascularity be attained with the first production, then the second necessarily subsides into the female. In this manner there of necessity originates an equiponderance in the number of both sexes.
2324. Moreover, if the sexual parts be regarded according to their proper signification in the animal, they are the upper intestinal system, as it has been developed in the mouth, and that indeed in such a manner, that the female parts are the vegetative form or the oral cavity, the male, the animal form, or the tongue with the salivary glands; the former represent the process of deglutition, the latter that of taste.
2325. Into the uterus or into the prostate gland the excretory ducts of the sexual glands, or the spermatic canals and oviducts, pursue a convergent course, like the salivary ducts do into the oral cavity.
2326. The testes, as also the ovaria, are forebodent of the _salivary glands_. The semen and oviducts are salivary ducts; they open by twos and symmetrically. Semen and ova are secreted like saliva. Semen and ova have also a salivary function, though of this the former is endowed with more than the latter.
2327. The ova correspond, as being an object of the semen, to the object of the saliva, which is the food or aliment. The saliva imparts to the bole or mouthful of food its first animal signification; it renders it for the first time capable of passing over into animal organs; it impregnates the morsel. The semen renders the ovum capable of effecting the transition into an animal; it spits upon the ovum.
2328. Impregnation is a process of smearing with saliva, conception a process of deglutition.
2329. Pregnancy is a process of digestion and formation of blood.
2330. If the internal sexual parts denote the internal visceral parts of the mouth, so must the external parts of the one correspond to the external of the other. The labia pudendi correspond to the lips, and the clitoris to the tongue, which is more perfectly represented in the penis. Both tongue and penis consist of two halves; in cases where the former is fissured or divided, the latter is so also, as in the Serpents and Lizards. In many animals, as the dogs and other Mammalia, there is likewise a bone in the penis, which corresponds to the os hyoides or linguale. The salivary ducts have combined with the penis; or, taken in a more strict sense, it may be said, that in the penis the tongue has coalesced with the oral cavity, so that both form a canal,--that of the penis, into which the salivary ducts (vesiculæ seminales) open.
2331. The sexual passion or venereal desire is a gustatory process of the sexual animal, the copulation being at one and the same time a matter both of taste and deglutition.
3. _Germ or Embryo._
2332. In the embryo the whole animal already resides in miniature, as does the plant in its seed.
2333. The embryonic intestine is the vitellus or yelk.
2334. The embryonic integument is the amnion.
2335. The embryonic vascular system is the chorion.
2336. The embryonic sexual system is the allantois. The above propositions can only be perfectly developed in the physiology.
b. VASCULAR ORGANS OF THE SEX.
2337. The hæmatopoietic or blood-preparing vascular system is the lung; the blood-destroying, exsecernent vascular system is the kidneys. The kidneys are the lung reversed. The liver decomposes the venous, the kidneys the arteriose blood.
2338. The _kidneys_ are the individualized vascular system of the sex, as the liver is that for digestion. They accord with the liver in their glandular structure, in the renal pelvis, which is like the gall-cyst, in the ureters that resemble the biliary ducts, and lastly, in the general signification of the urine as a product, wherein the whole organism, the whole blood-system has been excreted, like the bile, in which the venous blood undergoes the same process.
2339. Every disturbance of the digestive function acts in a striking and very direct manner upon the urine. The jaundice is apparent in urine; and what else is diabetes than a malady analogous to diseases of the liver? In urine we recognize what the bile has done with the food; the urine is the fluid nutritive system, and consequently the fluid organism "in toto;" the sexual blood, or sexual bile.
2340. The urine is the purest mirror or reflex of the bodily condition, and hence ourology or the doctrine of urine is of the most universal importance in semeiotics or symptomatic pathology: the ureters correspond to the tracheal branches or bronchi; thus, the urinary cyst to the trachea; the urethra to the larynx.
2341. In many animals the ureters open directly into the cloaca, as in many Fishes, Reptiles.
2342. By degrees the cloaca is retracted towards the ureters, and then originates a cloaca, which is both urethra and urinary cyst, as in Birds.
2343. In the higher animals, where a perfect ourocyst or urinary cyst has been evolved, the urethra opens into the anterior wall of the vagina, in front of which the urinary cyst then lies, in a similar manner to the trachea in front of the pharynx.
2344. In many Fishes the urinary cyst is absent, their pulmonic sac being also but feebly developed and only remaining as an asymmetrical swimming-cyst--the ureters too open directly into the cloaca, just as the swimming-cyst opens into the oesophagus. The pharynx of Fishes, as being surrounded by the branchial arches, is at once pharynx and larynx, like as in many animals cloaca and urinary cyst are of one and the same kind.
2345. In many Reptiles (such as the Tortoises and Frogs) the urinary cyst gives off two cæca or blind sacs, as is the case in the larynx of many apes.
2346. In the Bird the two cul-de-sacs of the urinary cyst are much more developed and have assumed the form of two cæcal intestines, so that they have been actually viewed as such, and their number two been assigned as characteristic of the bird, the other animals meanwhile having only one. The cæca intestinalia of Birds are the lateral and upper extremities of the urinary cyst. The true cæcum of the bird is the vitelline canal or duct, just as in the fishes and all higher organized animals; this being distinctly retained in the aquatic birds.
2347. In the Bird the rectum properly opens into the urinary cyst between the two blind sacs or cæca, and that indeed with a regular vulva, which is a sphincter muscle.
2348. The cloaca of the Bird is an urinary cyst, into which the anus opens.
2349. The orifice of the cloaca is properly that of the urethra; ova and fæces are moistened with urine. In a bird both these egesta are combined.
2350. Like the urethra, so is the trachea membranous in Fishes, and in most Reptiles also.
2351. Urinary cyst and urethra stand in sympathetic relation with the trachea and larynx, and have also similar diseases, such as catarrh, inflammation, &c.
2352. The proper proof, however, of the urinary cyst belonging to the respiratory system resides in its genesis. It proceeds from the allantois, which in Birds is a decided organ of respiration, a branchia.
2353. Out of this urinary cyst issue in the embryo what have been called primordial kidneys, which shrink or dwindle down at a later period, but have entirely the structure of branchiæ.
2354. Here then is a sexuo-respiratory process, which corresponds to the anal respiration, upon a higher stage, of many worms and aquatic larvæ, e. g. Holothuriæ and Libellulæ. This anal respiration is in its signification a sexual respiration.
2355. Even in the acephalous bivalve Mollusca and Snails the respiratory openings are almost always situated in the proximity of the anus; their respiration is still a sexual respiration.
2356. For the first time in Insects it becomes a respiration of the trunk; and for the first time in the higher animals a truly animal, namely, a cephalic respiration.
2357. The urinary system is a double system; it conjoins in itself the two highest galvanic processes, that of secretion and excretion.
2358. Secretion is an hepatic character, excretion a pulmonic character. Secretion belongs to nutrition, excretion to respiration. Excretion is an exspiration, secretion an influx or infusion. Secretion is related to excretion as water is to air, as liver to lung, as basis to oxygen.
2359. Secretion takes place, e. g. of bile and saliva, in so far as the processes of the body, especially those of the digestion, are promoted. Excretion is only effected in so far as that the organs, into which what is secreted enters, may obtain a tracheal signification, i. e. evaporation. All excretory orifices are in a certain sense larynges or tracheal orifices. Thus, this relation between secretion and excretion would have been discovered without recourse to conjecture being had upon our part.
2360. The urine is "par excellence" a double product of this kind. It is secreted in the kidneys for simply one object, like the bile. It is excreted, because it belongs to the sexual system, which is essentially exsecernent.
2361. The object or purpose of the urine has not been subverted in all animals. In Birds, where the urinary cyst and intestine are confluent, the urine enters like the bile into the intestine, at least at a spot where there is intestinal matter, which it renders fluid.
2362. In Insects and Snails it appears to invest as a mucus the ova, and serve for their attachment. The same appears to hold good of the spider's web. What has been called purple juice and ink (in the Cephalopods) ranks probably in the signification of urine.
c. SEXUAL INTESTINE.
2363. The sexual intestine is the colon or large intestine, which, in this respect, belongs to the sexual system, as has been indicated at § 2171.
II. _Animal Sexual Organs._
2364. The bones of the sex are the feet with their appurtenances, such as the pelvis with the lumbar, sacral and coccygeal vertebræ. The muscles, as well as the nerves, are self-understood. But of these we will speak in the sequel.
B. ANIMAL ORGANS.
2365. All organs, which are purely animal, are penetrated or traversed by the nervous system, just as the lower systems are by the tegumentary formation. No higher organ is the wholly pure evolution of _one_ system, but the systems ever combine with each other; and this combination, when individually represented, constitutes the organ.
2366. Organ is distinguished from system in not pursuing its course through the whole body, nor consisting simply of one and the same mass, but by its occupying a definite part of the body and being composed of several systems.
2367. Every organ has therefore a special and specific function also.
2368. The systems of animal life divide only into two kinds of organs, into those of sensation and those of motion, into the solar and planetary, or central and peripheric.
a. MOTOR ORGANS.
2369. Bone and muscle are not societies, but only poles of one system. There is therefore no mere bony organ, and no mere muscular organ. Meanwhile we will here regard them in particular or "per se."
1. _Osseous Organs._
2370. The first bones were branchial arches or tracheal rings. When the lungs were developed from the branchiæ, the branchial arches were repeated as ribs or pulmonary arches. Lastly, should bones be formed, which are to be wholly in the service of the animal or the nervous system, so also must they be wholly liberated from the vegetative organs, and become self-substantial, i. e. have nothing else to do, but move. Free motor organs can be none other than ribs that have become free.
2371. These free ribs must inclose the respiratory organ or the integument, which has become an organ of animal life; they are the members or _limbs_. If we think of ribs, whose office is no longer to inclose lungs, which must no longer be subservient to the uninterrupted vital motion of respiration, and which are no longer united by pleura into a closed cyst or sac--will not such simply retain the self-substantial voluntary motion in themselves? will they not abandon the inferior cystic form, and represent the same, though but ideally and voluntarily? will not such a thorax open in front, like the intestine has opened at its nobler extremity?--will not such ribs be members, arms, digits? The members or limbs are the _members of the trunk_, or ribs that have opened in front; they are the thorax that has opened in front; and hence are nothing new, but only something emancipated or set free. Such ribs can be none other than motor organs; for they were previously nothing else. Then, however, they performed motion in the service of the viscera; at present, where they are absolved from this service, they execute it only in accordance to the will of the head, simply according to its will, for they are _verily nothing more than motor ribs_. Where, however, or in what region of the body will the ribs attain unto such freedom? Without doubt, in the neighbourhood of the head, and thus at the very spot where the lungs derive one of their extremities. The limbs are therefore _cervical ribs_.
2372. The arms are a thorax, consisting quite purely of bone and muscle, and represented as isolated or detached from the viscus, or the lung; on this depends their nobility, or, in other words, upon what is vegetative having been wholly left behind.
2373. The arms, when clasped together by the fingers, are a thorax without viscera, without heart and lungs; they are destined to inclose a whole body in the embrace.
2374. By an embrace that which is embraced has been made our viscus; it has been adopted as our animal heart, and as our animal vital organ--or lung. The embrace has an exalted physiological signification, and precisely that which it unconsciously possesses in the state of pure love. Nature always thinks more nobly than we do. We follow blindfold her beautiful regulations, and she rejoices in the sport.
2375. As the fundamental number of the branchiæ is five, so also must the limbs represent five ribs; they split into five digits or fingers. The feet of the Crustacea and Insects generally do not correspond to our feet; but to our fingers. The lower organized animals have only toes, no feet. The five thoracic feet of the crab correspond to our fingers; its five abdominal feet to our toes.
2376. There are three limbs in accordance with the three totalities of the body, truncal, sexual, and cephalic members, or arms, feet, and jaws.
2377. The members of the body or trunk belong to the thorax, because it is the respiratory system. The abdomen has no members; what have been so-called, are in their signification sexual limbs.
2378. Had the animal no sex, it would have no posterior limbs.
2379. As the three lower cervical and the two upper dorsal vertebræ belong to the arms, so also they appear to commence with five ribs, but then to become arrested, and again emerge to perfection in the digits.
2380. The shoulder appears to consist of five ribs, but this does not as yet admit of being clearly pointed out. Meanwhile, it is certain that the scapula, acromion, and coracoid are particular bones, to which may be added the clavicle.
2381. The middle finger is the elongated radius, and is therefore the longest or radial digit. It is that which is persistent, if only one finger has been left, as in the horse. The ring-finger is the ulnar finger. It is that which, along with the former, appears in the bi-ungulate animals; the spurious or dew-claws are the auricular and index-finger; the thumb is the last ramification, is therefore always arrested, and frequently present only as a wart-like excrescence or papilla.
2382. All animals, which have true digits, are furnished with five of them, more or less completely developed. If what has been called the metacarpal bone of the thumb be numbered, which it must, as a digital articulation or phalanx, every finger has thus one carpal bone, and each bone of the fore arm also one.
2383. The sexual members or feet correspond in all their pieces to the arms: the pelvis is the shoulder repeated; and certainly, the iliac bone is equivalent to the scapula, the ischium to the coracoid process, the os pubis to the acromion, the marsupial bone to the clavicle.
_Cephalic Members._
2384. Both pairs of limbs are repeated in the head, because in it the whole trunk is repeated; the upper jaw corresponds to the arms, the lower jaw to the feet. Each jaw consists of two members, which are ankylosed in the higher animals at their point of meeting or in front, but in Fishes are partly, and in Insects completely, separated.
2385. Each jaw consists of the same bony divisions as the limbs of the trunk, of scapula, humerus, and fore-arm; or of pelvis, femur, and tibia. This is easily to be demonstrated in Birds, Reptiles, and Fishes.
2386. The digits are repeated in the teeth. The teeth are claws.
2387. There are therefore five kinds of teeth, which correspond to the five digits. The thumb becomes the canine tooth; the index-finger the false molars; the middle finger the laniary molar, the ring-finger the second, and the little finger the third true molar tooth.
2388. The intermaxillary with its incisor teeth belongs, as well as the palatal bones, to the pharynx; and is a visceral or intestinal maxilla.
2389. The lower animals therefore, as the Fishes, have almost nothing but intermaxillary and palatal teeth. They act principally upon the lingual teeth. The reptiles have still palatal teeth, which higher up in the vertebrate series disappear.
_Symmetry._
2390. As the cervical ribs have nothing more to inclose, have no longer to respire, but only to move; their symmetrical development is thus undelayed. The symmetry is at first wholly attained by the act of opening or apertion.
2391. The limbs are the most symmetrical organs. They are symmetrical in each smallest part, and these parts again assume a symmetrical arrangement in regard to each other. They are the ideal of symmetry.
2392. They are, however, the free living symmetry. They can create by their motions symmetrical forms. The symmetry consists chiefly in, and has been produced only by, the motion.
2393. The symmetry of the motion is the most exalted, for it is that which is endowed with life. The symmetry of the form is that which is dead.
2394. The symmetry of the form appertains to inorganic nature, the symmetry of the motion is the property of animals.
2395. Dancing and acting are the highest organic symmetrical movements, and also the highest symmetries. They are the symmetry of the motor members, wrought by motion.
2396. Music is a much higher symmetry of motion.
2397. Speech is the highest spiritual symmetry; it is the dance and histrionism of the spirit or mind.
2. _Muscular Organs._
2398. The muscles are everywhere attached to the bones, and help to form with them the same set of organs.
2399. The muscles of the larynx are therefore the antetypes of the costal muscles, these of the member-muscles, and the dorsal muscles of the scapular and pelvic muscles.
2400. The muscles of the limbs are found in a state of treble repetition. That the muscles of the arm and leg are of one kind, admits of being demonstrated with tolerable facility. But it is necessary, that in doing this regard be paid to the ligaments.
2401. The ligaments are only stunted muscles. Without bringing them into account, the muscular system does not admit of being explained and understood.
2402. The crural muscles are again found in the lower jaw.
2403. The brachial muscles, on the upper jaw or face.
2404. The movements of the facial muscles correspond to those of the limb-muscles. Upon this depends the interpretation of dumb-show, or the art of physiognomy.
3. _Nervous Organs._
2405. Nervous organs are liberations of individual parts of the nervous system, and their endowment with a peculiar function or sensation.
2406. The liberations of the nerves are combinations with the other anatomical systems at the spot where they have attained their highest evolution.
2407. Each system, however, has its peculiar process. Through the reception or adoption of the systems into the nervous system, a peculiar sensation must therefore originate.
2408. Peculiar sensations are sensations of sense.
2409. The combinations of the anatomical systems with the nervous system, whereby the former have been subordinated to the latter, are consequently _sensorial organs_, or organs of sense.
2410. In the sensorial organs the processes of the several systems attain unto sensation. They are brains of the anatomical systems.
2411. There are as many senses as there are different anatomical systems, and consequently senses belonging to vegetative and animal life.
2412. The number of vegetative systems is 3; the vascular, intestinal, and pulmonary system.
2413. The most complete combination of the vascular with the nervous system is the integument--constituting the tegumentary sense, or _sense of feeling_.
2414. The most complete combination of the intestinal system with the nerves is the tongue--intestinal or _gustatory sense_.
2415. The most complete combination of the lungs with the nerves is the nose--pulmonary or _olfactory sense_.
2416. Among the three animal systems, bones and muscles produce, from their association, only one action--the motion. The most complete and perfect combination of the motor system with the nerves, is in the ear--osseo-muscular or _auditory sense_.
2417. The nervous system has become a self-substantial organ in the eye--the nervous, or _visual sense_.
2418. There are therefore only 5 senses; they are none other than repetitions of the anatomical systems in the sensation; they are the highest developments which are possible in the lower systems, being the blossoms or heads of such systems.
2419. These systems are, however, processes of the universe taken up into the organization. In its organs of sense, the processes of the universe are thus felt or perceived. The senses are world-organs, and are therefore placed in contact with the world, or occupy an outward position.
2420. The vascular system is the nutritive system. In it the blood coagulates into the solid parts of the body. The feeling-sense is therefore sensible of the nutrition or rigidifying process of the body. Now, the Solid of the planet is the earth. The sense of feeling perceives therefore opposition--it is an _earth-sense_.
2421. The function of the intestinal canal is digestion. In taste, the process of digestion is felt. But digestion is a solvent, a hydrapoietic process; in taste the water is therefore felt--it is the _water-sense_.
2422. Respiration is a process of oxydation. In smell the respiratory process is felt. Oxydation is, however, an air-process--it is _air-sense_. Thus do the three vegetative senses feel the elements of the planet--are _planetary senses_.
2423. The animal systems are symbols of the æther, of the gravity with the heat or motion, and of the light.
2424. The motion is only moved matter, and thus a combination of the muscular and osseous system. The ear therefore perceives the motion of the primary matter, or the atomic motion--it is _gravity-sense_, _æther-sense_.
2425. The light is the tension-process of the æther; to see, is therefore in an organism to emit light or shine--_light-sense_.
2426. The signification of the senses is twofold; they are anatomical _systems_ which have become nerves, and on that account also _elements_ that have attained unto sensation.
1. Sense of feeling--vascular, tegumentary, nutritive sense, _earth-sense_.
2. Gustatory sense--intestinal, digestive sense, _water-sense_.
3. Olfactory sense--pulmonic, respiratory sense, _air-sense_.
4. Auditory sense--osseo-muscular, motor sense, _æther-sense_.
_Matter-Sense._
5. Visual sense--nervous, tension-sense, _light-sense_.
2427. The sensorial organs are not simply combinations of the anatomical systems with the nerves, but also with the bones and muscles. These have been entirely taken up into the signification of the animal body.
2428. Each sense has its own nervous, osseous, and muscular system.
2429. The sense of feeling has its bones and muscles in the limbs.
2430. The sense of taste, its bones in the lingual bone, its muscles in and upon the tongue.
2431. The sense of smell, its bones in the nasal bones, its muscles being frequently very much developed in the snout or proboscis.
2432. The sense of hearing, its bones in the auditory ossicles, its muscles in the auditory conch.
2433. The sense of sight, its bones in a ring surrounding the sclerotic coat and in the eyelids, its muscles in the ocular muscles.
2434. Besides the proper sensorial nerves, each organ of sense is provided with nerves for the motor system and for the vegetative systems, especially those of secretion.
2435. The integument has, besides the nerves of the tactile papillæ, vascular and motor nerves.
2436. The tongue is supplied with nerves of motion and digestion, and has therefore three pairs of nerves.
2437. The nose receives motor and respiratory nerves from the fifth pair.
2438. The ear has likewise three kinds of nerves; the auditory nerves, facial nerves, and a branch from the fifth pair, not to mention those that are distributed to the auditory conch.
2439. The eye has, in addition to its wholly special motor nerves, a number of others, which superintend its vegetative systems, such as the iris and the secretions of the humours.
a. VEGETATIVE SENSES.
1. _Vascular Senses._
2440. All senses are only conditionated by the peripheric nervous mass because they are combinations of the nervous mass with the blossoms of the inferior systems.
2441. The most general system of the animal is the vascular system, which is represented externally as integument. The animal was in the commencement nothing but integument, and this again naught but vascular and nervous mass, so that the whole integument was thus an organ of sensation.
2442. Through the integument the animal becomes an individual, or a something distinct from the aggregate of nature. Now as the integument is principally the organ of sensation, so is the primary sensation that act, by which the animal is distinguished from nature. The tegumentary sense is the sense of _distinction_, of limitation.
2443. Through the act of discriminating, a something foreign or extraneous is granted us. The immediate perception of what is extraneous, is called feeling. Tegumentary sense is _sense of feeling_.
2444. The feeling-sense is the first in the animal.
2445. It is that which is general in the animal.
2446. The whole animal is naught but a sense of feeling.
2447. Out of the feeling-sense all other senses must be developed, just as all other systems are developed out of the tegumentary formation.
_Organs of Touch._
2448. Where, however, the skin has attained to a higher grade of formation, or where it has combined with higher systems, there also will the sense of feeling be supplied by the former.
2449. The combination of the integument with the osseous and muscular system, and with a nervous system of its own, takes place in the limbs. Since the motor members are only a liberated thorax, so no other sense can belong to them but that of feeling, which the thorax previously possessed.
2450. But these sensitive organs are moved, and therefore voluntary organs, digits, or former branchiæ. Moveable or voluntary organs of feeling are called _tactile organs_. The feeling of the motor members is touch.
2451. The highest feeling necessarily consists in touching, because it has in that become active, while before it was only passive.
2452. In the situation of the sensitive papillæ, the origin of the digits from respiratory organs admits of being still recognized. They are arranged in spiral lines upon the points of the fingers.
2453. The sexual organs belong as tegumentary developments to the sense of feeling. There is no special sexual sense.
2. _Intestinal Sense._
2454. Opposite to the general feeling or the integument, the function of the intestine is evolved. In the trunk it is simply busied with its own processes; when it first mounts or ascends into the head, it becomes subordinated to the nervous action.
2455. The combination of the intestine with bones, muscles and peculiar nerves, is presented in the tongue. This is the sensitive organ of the intestine.
2456. The tongue is a feeling-sense in water, just as the skin was that in the air. For it is the blossom of the digestive process. To the tongue therefore belongs the digestive or water-organ of the mouth, which is constituted by the salivary glands.
2457. The sensation of what is fluid in its chemical relations is called _taste_. Gustation is not a peculiar process, but obviously only the nervous commencement of the digestive process. On that account also the gustatory sense still lies concealed in a cavity. The whole buccal or oral cavity still belongs to the sense of taste.
2458. As in the sense of feeling the motor system still predominates, so does it also in the tongue, as being the second sense, which has been liberated from the plant. The nervous mass is in this sense not preponderant over the muscular and bony mass.
2459. The tongue is still to be regarded as an organ of touch, though one in which the flesh or muscle has gained a mastery over the bones, while in the true tactile organ the bones determine the principal forms and functions. The tongue is a nervous organ in the muscle, the hand is such in the bone.
2460. The hyoid or lingual bone is none other than the first branchial arch, and consists pretty nearly of the same pieces as the arm.
2461. Compound lingual bones, such as occur in many Reptiles, have originated from coalescence of several branchial arches.
2462. Like the limbs, so is the tongue originally a double organ. In most Reptiles it is longitudinally fissured. Such animals have usually also a double penis. In all animals the tongue is divided into two moieties, which are only connate by means of suture. The penis also consists of two connate penes.
2463. As in the tegumentary sense, the nerves could not be peculiar or special in kind, but took their origin from all parts, and particularly from the spinal cord, so also is this the case with the intestinal sense, which is still only an internal tegumentary sense. The lingual nerves proceed from several situations, and that too from the upper part of the spinal cord.
2464. The oral cavity also consists, properly speaking, of mere tactile organs, which have been repeated in the head. Thus, there are tactile organs which are subservient to the gustatory sense, in biting, chewing, and swallowing.
2465. The lips are tactile organs upon the threshold or brink, as it were, of the gustatory sense.
2466. In the oral cavity, however, the glands of the intestinal canal are repeated. The salivary glands secrete fluid, like the pancreatic glands.
2467. The sense of feeling is present in all animals. They are only animals by virtue of it; but the sense of taste appears to be first formed at a later period, after the intestine has separated from the integument; in the animals that have no intestine, its existence is problematical, and even in Fishes and Birds it is but poorly developed.
3. _Pulmonic or Lung-sense._
2468. When the respiratory organ is hoisted up into the head, and there becomes an organ of sensation, it passes over into a sense.
2469. That the nose is the thorax, together with its viscera, repeated in the head, has been already remarked. The many convolutions of the turbinated or olfactory bones correspond to the ramifications of the trachea; the nasal cartilages to the tracheal or laryngeal rings; the olfactory membrane to the pulmonic vesicles.
2470. The process of the lungs repeated in the head becomes smell, like that of the intestine became taste. The olfactory sense is the highest blossom of the arteriose vascular system or the branchial net. On this account the olfactory membrane is the most delicate, and the densest tissue of arteries and veins.
2471. The nose is related to the mouth, as the thoracic is to the abdominal cavity; the olfactory membrane to the tongue, as the lung is to the stomach. It is a cephalo-thorax. The nose is not therefore so completely closed as the mouth, but is opened through the two most anterior air-holes or spiracula. The nasal apertures or nostrils are the last persistent remnants of the spiracula, after all those upon the sides of the body have been closed up.
2472. It is the last organ of sense, which has been evolved from the trunk. It is therefore nobler than the two others, and has also a nobler object, the air.
2473. The nerves of the olfactory organ are peculiar to it, and are encephalic nerves. As the sense of smell is the pulmonic or arteriose sense; so also does the arteriose substance of the brain combine with this organ. The olfactory nerves consist of cineritious or gray substance, and are only prolongations thereof.
2474. This is the only phenomenon of the kind met with among all nerves, but it is commensurate with the character or signification of this organ. A sensible pulmonary organ can only have arteriose nerves. As the liver is throughout venous, so is the nose throughout arteriose in quality.
b. ANIMAL SENSES.
2475. It now only remains for us to consider the motor and the sensitive system proper upon their highest stage of development. The motor system, when represented in the nervous system, is a peculiar organ of sensation, as is likewise the nervous system itself upon attaining its highest state of development.
4. _Osseo-Muscular Sense._
2476. The lowest condition of the motor system is the limbs, which represent no peculiar sense, but only the sense of feeling refined or set in motion. This motor system ascends into the head, and there no longer exercises its motive powers in prehension, progression, &c., but solely unto sensation. Now, a system, which converts its function into that of sensation, is a sense.
2477. The sensorial organ, which simply through motion, or a resistance presented to that of the atoms, produces sensation, or wherein the motion as such is felt, is the _Ear_.
2478. The ear is none other than the ultimate development of the bone and muscle, when they are brought under the direct dominion of the nerves.
2479. The auditory ossicles are the limbs subtilized or refined in character. They have joints, are provided with muscles, and move exactly like the limbs. It may be said; that the stapes is the scapula, the incus the humerus, the malleus the fore arm, and the concha with its cartilages, the hand with its digits.
2480. The ear, like the bulk of the limbs, has originated from branchiæ. In Fishes, the auditory ossicles have degenerated into the branchial opercula.
2481. The ear-trumpet (or Eustachian tube), which opens into the mouth, is the internal branchial aperture.
2482. The motor system, however, belongs to the trunk, whose viscera are repeated also in the ear, or in a cavity which has been called its labyrinth. The three semicircular canals appear to correspond to the intestine, the cochlea to the trachea.
2483. The ear has not only a nerve, but likewise a brain, of its own.
2484. The cerebellum is the auditory brain, and from it the acoustic nerves take their rise. For, since the ear is the sense of the whole motor system, and consequently of half the animal, it inevitably follows, that an appropriate or special nervous mass must be developed for it, just as the myelon or spinal chord has been for the trunk. An organ, which is so constantly active, must necessarily have a large nervous mass. The cerebellum is consequently not a brain in the general sense, but one which is wholly individualized. It participates in the motion, which, as sound, is transmitted to the animal.
2485. By its signification as well as by its own brain, the ear gives us to recognize its elevated rank above the other senses.
2486. The ear must stand in relation with the body's limbs.
2487. The ears first make their appearance in animals, associated with a tolerable development of the limbs. With the exception of a few instances, the ears first become apparent in Fishes, being in them at least furnished for the first time with true ossicles and semicircular canals. The ear, like the limbs, with which it constantly maintains a parallel course, is very slowly perfected. In the Fishes which have only fins (as instruments of locomotion) it is wholly concealed within the cranial bones; in the Reptiles it emerges or is more exposed to view; but for the first time in Birds and Mammalia where, generally speaking, the limbs also are first perfected, it attains its completion; in them only is the cochlea fully developed, and an auditory meatus which opens externally.
5. _Nervous Sense._
2488. In all lower organs, and even in the senses which have hitherto been commented upon, the nervous system was not the chief, but only the co-ordinate agent. It has only by its conjunction with, assisted in elevating the character of, other systems, so that their material might be converted into spiritual processes. But the nervous is also a self-substantial system, and must therefore attain likewise a free development.
2489. With the highest organs of the nervous system, the relation which has been hitherto maintained, must be reversed. The inferior systems will now be the co-ordinate agents.
2490. The highest nervous organ can only possess that function, which is originally peculiar to the nervous system, i. e. the most delicate polarization, the light-function. There is the _Light-sense_.
2491. The eye is nothing but a nervous system represented in a state of purest organization, just as the ear was the purest system of motion.
2492. In the eye it is the brain itself, which expands, in order to turn unto or face the light.
2493. As the ear has its own brain, so also has the eye; the cerebrum is the optic brain. (Ed. 1st, 1811. §. 2317.) This is the rationale of our having two brains.
2494. Now the cortical or gray substance has been already diverted from the cerebrum to supply the olfactory sense. Its medullary or white substance remains for the eye. The medullary brain is the optic brain. The medullary is consequently the nobler part of the nervous system.
2495. The brain's medulla is homologous to the light; its exterior or cortical portion is related to the material light, the air.
2496. The eye is only a medullary brain, posited peripherically and in a nervous manner. The brain itself has elongated and become integument.
2497. The visual membrane (or retina) is the cerebral substance cystically expanded. It must be regarded as being originally a closed bladder or cyst. (Ed. 1st, 1811. §. 2321.)
2498. The optic nerve is itself hollow, and unites the cerebral with the orbitar cavity.
2499. The vitreous body, which fills out the cyst of the retina, is the cerebral medulla which has become transparent, or a semi-fluid albuminous mass.
2500. The sclerotic coat of the eye is the continuation of the brain's dura mater.
2501. The vascular or choroid coat of the eye is the continuation of the encephalic pia mater. All parts of the brain have been consequently continued into the eye.
2502. Now, what the brain is for the earthly body, that also must it be for the eye. The eye is not simply brain, but a representation also of the whole body. The brain can, forsooth, be nowhere without its body; if therefore it is elevated into the eye, so also must it take up and elevate the body along with itself.
2503. The eye is an entire body, a whole animal. Again, in the next place, the animal systems, such as limbs, thorax and abdomen, have been most distinctly represented in it. The light is seized, respired, digested, and hence felt by the eye.
2504. As the light represents chaotically the whole of nature, but this material nature enters completely into the animal through the processes of the trunk, so does the light enter it through the eye. The eye is the chaotic representation of all material processes of a body.
2505. The limbs or members of the eye are repeated in the ocular muscles and the sclerotic or bony ring; in many Fishes the eye stands upon a flexible pedicle, as in the Crabs. The ocular muscles move the eye in different directions like a hand.
2506. The sclerotic corresponds to the corium, the cornea to the digital unguis, or finger-nail.
2507. The choroid or vascular coat, is the respiratory system or lung in the eye. The iris corresponds to the larynx, the pupil to the glottis; its expansion and contraction is a respiratory movement.
2508. The choroid coat incloses also an osseous mass, the lens--a vertebral body. The morbid states of the lens are osseous diseases, such as gout.
2509. In the chambers of the eye, water, as being a product of digestion, is constantly secreted.
2510. The orbitar cavity is a mouth with salivary glands--giving vent to tears.
2511. The lachrymal canal is a branchial duct, which opens into the nose, like the Eustachian tubes did from the ear into the mouth.
2512. The eyelids consequently correspond to the lips, and are in like manner fringed with hairs.
2513. As the body has everywhere two halves, and laterally also presents two entire organisms, so also is the formation of the nervous system a double one. Each eye is an entire body.
2514. In the two eyes the halves of the body have completely separated as entire bodies, and each has attained self-substantiality. Each eye is a free animal in the animal body. Each eye is therefore circumscribed by its own _integument_--is a free animal. It is endowed, like the hand, with omnilateral motion; it has cavities, i. e. its bodily cavities and humours, or inclosed masses--_viscera_.
2515. An organ, which again repeats in its miniature the whole animal itself, of which it is only a part, must necessarily be the highest point unto which an organism can attain. With the eye the organization, and consequently nature, has been concluded.
2516. The eye is a parasitic animal, of the same kind with the animal upon which it exists.
2517. In a certain sense all sensorial organs are parasitic animals in the animal; only they are not all of the same kind with it. No one of the other senses has e. g. repeated _all_ lower systems in itself, and it is therefore to be regarded only as a subordinate or half-animal, which lives upon that which is more perfect.
_Senses of the Sexual Animal._
2518. In essaying to speak concerning the sensorial organs of the sexual animal, we shall only encounter in the latter the emotions of the vegetative senses, and these indeed disposed according to their rank.
2519. The sense of feeling is most perfectly developed in the legs, whereof the pelvis represents the scapula.
2520. The external sexual parts are the analogues of the gustatory sense; the female parts being indeed those of the mouth; the male, which are frequently furnished with bone, of the tongue. The jaws are not repeated in a sexual animal, except in Insects, namely, as the pharyngeal maxillæ.
2521. The analogue of the nose is wholly stunted, and is only left persistent as a trachea or air-tube in the urethra.
2522. In other respects the cavity of the sexual parts is a trunk-cavity "per se," like the thoracic and abdominal; the pelvic cavity contains the viscera of a whole animal.
II.-_PHYSIOLOGY._
2523. Physiology is the doctrine or science which treats of the functions of the animal. Like the organology was, so also must the science of the functions be developed. There will be functions of the whole animal, and of the tissues, systems and organs.
_A._ FUNCTIONS OF THE ANIMAL IN GENERAL.
2524. The first act of the animal is an assimilation to the universe, whereby it also receives or takes up into itself the primary function of the universe. This is the perception of the circumscription and totality in itself, its self-manifestation, the _feeling of self_. The first action of the animal mass is that of self-sensation. Now, through the self-sensation, self-substantiality has been granted.
2525. The animal is consequently a Whole in Singulars only through the feeling of self.
2526. As the universe is only an analysis of the self-consciousness of God; so also can the development of the animal and the formation of its organs, be none other than an analysis of the self-sensation. All other functions (just as all masses are only a metamorphosed nervous mass) are but differently polarized self-sensations, or these as it were dispersed.
2527. The self-sensation of an individual body is not, however, inclosed within itself, like the primary relation of the universe; because it is not the universe, but only a fraction of it, which has sprouted forth like a bud from the great planetary body. The self-sensation is not therefore a simple feeling of self, but a feeling also of something _foreign_ or extraneous, and thus the animal self-sensation becomes, in reference to Nature, an act of discrimination or discernment.
2528. Now, the animal is distinguished only from Nature by the act of being liberated from it. It is therefore comprehended in a constant liberation or severance. The life of the animal continues only by a constantly renewed and indefatigable severance, by a desertion or falling off from Nature.
2529. In so doing, however, it detaches itself from Nature, as being a part thereof; the severance is therefore a conversion of the nature into an animal. In this consists the reciprocal action of both, viz. that the animal is constantly seeking to assimilate the nature unto itself.
2530. The ability or power to assimilate the nature, is called _excitability_.
2531. Excitability is the most general phenomenon of the organic mass, and appertains both to plants and animals.
2532. But in the animal excitability, the free self-sensation, within which a free motion is necessarily inherent, is superadded or originates. This excitability unto motion I call _irritability_.
2533. Thus this irritability belongs only to animals.
2534. Irritability does not depend directly upon motion, but throughout upon sensation. Without sensation no irritability is possible. If the sensation ceases, so also does mobility, or the capacity for motion, cease.
2535. Since irritability originates from the antagonism of the animal with the world; so is it parallel to an antagonism of the heavenly bodies, or to that of sun and planets. The mutual operation of these two heavenly bodies is, however, an interchange of polarity, a polar excitation. The irritability is a polar process; but one which is pure and devoid of material excretions, just as the sun excites the earth without eliciting therein any material change or transition. The animal becomes polarized by the incentive agent or stimulus.
2536. Through the irritability there originates a double polarity in the animal. In the first place one between the world and the animal; in the second, one between the exterior of the animal and its interior. The world-polarity gives the feeling or sensation, the body's polarity the motion.
2537. In the sensation the animal always transcends itself; there is thus only excitability. In the motion the animal abides or remains within itself; but the self-sensation proceeds from both conditions. Accordingly, in self-sensation the world, and the animal within the animal, convene or come together. The animal is itself universe, and it at the same time comprehends the great universe. Now, in both conditions or in both functions has the animal been turned towards the world and also towards itself. In feeling it turns itself towards the world, in order to adopt this spiritually into, or repel this from, itself; in motion it turns itself towards the world, in order to materially adopt or to repel it. In both cases of assumption it turns itself towards itself.
2538. Both these properties belong to the animal body; if individual organs are deficient in them, it happens therefore, from their having superadded or adduced other properties, which are predominant, to the original ones. This is the essence of what is animal in a living body. Thus, if from an animal all the vegetable systems could be subtracted, it would do nothing but feel and move.
_B._ FUNCTIONS OF THE ANIMAL IN SINGULARS.
I. _Functions of the Tissues._
a. FUNCTIONS OF THE ANIMAL TISSUES.
1. _Functions of the Point-Tissue._
2539. The point-tissue is also spiritually represented, and this spiritual manifestation or expression is the stamp or impress of the arrangement of the matter into points.
2540. The point-matter is the matter of sensation; the sensation is, however, imparted by polarization. The act of feeling is a conduction of the polarity from point to point; this is the light-polarity. This light-like polarity transmitted from point to point is sensibility.
2541. The sensibility is not the result of a peculiar nervous fluid, or even of mechanical vibration, &c., of the nerves; but of an antagonism between animal and world, brain and tegument, sun and planet.
2542. The point-tissue is in every respect the æther which has become the mucus, and has in every respect also to act in accordance thereunto.
2543. The sensibility or the nervous power, viewed in relation to the smallness of the body, operates instantaneously upon every spot. It is not first conducted or tardily conveyed hither and thither; but everywhere, where an antagonism, a stimulus, a _planet_ is, there also is the "vis nervosa." Where a planet is, there also is the solar tension.
2544. But abstractedly or ideally considered, the nervous force certainly requires a time to elapse for the propagation of the stimulus, just as the light-tension darts through the æther only in a given time. As light traverses at the rate of 40,000 miles per second, so must the nervous power, if it obey similar laws to those of its prototype, and the length of a man be estimated at 1-5000th of a mile, glide through the human body in 1-200,000,000th of a second, which, "quoad" the phenomenon, cannot, of course, be observed.
2545. Accordingly, it is theoretically certain that the nervous power does not operate with absolute or unconditional velocity. To the validity of this statement, observations of many kinds, especially those made in diseases, and by astronomical computations of the strokes of the chronometer, afford additional evidence.
2546. Thus, for a stimulus impinging upon the brain, to act or be transmitted from thence into the toes, there elapses 1-200,000,000th of a second. In the event of disease, a retardation in the process is conceivable, and may admit even of being observed.
2. _Function of the Globe-tissue._
2547. The dense or solid form is the matter, when it has died away; the function also, or the crystallization, died therein, so soon as the crystal was represented. The office of the bones is only to preserve the counterpoise to the nerve, to supply a limit, and therefore a support to its action, by which means the achievement of the latter first becomes possible.
2548. The business of the bone, is to play an antithetic part to the nerve, and nothing else, or to serve the latter as a "terra firma," upon which it may execute its plans. The bone suffers, and in this consists its office. The other offices of the bone, such as its being the firm and solid framework of the body, its protecting the nervous mass, &c., are subordinate matters, which are at once clearly understood.
3. _Function of the Fibrous-tissue._
2549. There is the active motion. The primary motion is, however, a result of the polar tension induced in the æther by the light. Just as the heat has been produced in the æther by the light-polarity, so has been the animal motion in the fibres by the polarity of the nerves.
2550. Every fibre, when it is in motion, has necessarily two polar extremities. For the fibre stands between the artery which is oxygenous, and the nerve which is basic, or between a zinc and silver pole.
2551. In the conditions of rest the two poles must be neutralized, or, generally speaking, not in existence. This is only possible by disjunction of the galvanic circuit or chain.
2552. Since the artery is constantly operating upon the fibre, but the nerve only, when it has been stimulated; so must this change in the condition of the fibre reside in the change of the nervous influence.
2553. In unirritated conditions the nerve does not act upon the fibre; the latter is not _illuminated_, nor does it stand in a state of tension with the nerve, which is not in itself polar, but is only, when stimulated, a sun towards the fibre, or polar within itself. If the chain be then dissevered, the fibre is in a state of indifference.
2554. Thus, in the event of tension, the nervous extremity of the fibre becomes negative, the arterial extremity positive; both ends necessarily attract each other in order to discharge or unload themselves. This attraction is an _abbreviation_ or contraction.
2555. After the discharge the fibrous extremities are homonymous; they repel each other, and relaxation or extension ensues.
2556. The motion of the fibres is consequently a perfectly galvanic process, devoid of _material_ interference, between nerve, fibre, and blood. The nerve is silver, the artery zinc, and the fibre moist card. All theories of motion based upon swelling of blood, elementary change, and such like causes, are not animal, though organic in character; the former are injections, the latter chemical processes.
2557. Through the fibrous tissue being shortened and elongated, the entire animal is self-displaced.
2558. Through the fibrous tissue the animal _appears_ as animal, through the point-tissue it is (and that in essence) an animal. God is in himself, but he first appears in the world. This elucidates or explains the relation of "noumenon" and "phenomenon."
4. _Function of the Cellular Tissue._
2559. This tissue has no other function than that in the plant. Many saps or juices are effused into the cells. The latter are engaged in constant expansion and contraction, whereby they decompose and propel these juices. They are the proper seat of the nutritive process, since they coincide with the capillary vessels.
2560. But as every rigidification is combined with volatilization, so does a process of evaporation occur in them at the same time.
2561. This process is the property of the teguments or membranes, both of those lining the internal cavities, as also of such as are placed externally.
2562. But the process of evaporation is principally the function of the true skin or corium. As an organ of evaporation the corium is similar to the lung. It is consequently aeriform water, which of necessity constitutes the principal mass of evaporation.
2563. The water is, however, mucous in character, the mucus becomes oxydized in the air, and through this also carbonic acid is formed.
_Heat._
2564. The chief function of the cell-formation is the process of heat. The cell-process and heat-process are one in kind. Heat is the product of the process of condensation and rarefaction, and thus of the nutrition and evaporation, both of which happen in the cells, or, if we please, in the capillary vessels.
2565. In the skin the process of temperature is individualized.
2566. All temperature depends upon the process of condensation and rarefaction. At one time it is the nerves, at another the vessels, anon, external influences, which alter or modify it. Animal heat, like the cosmic heat, has been produced by change of the fixation. This change, however, takes place principally in the nutrition and evaporation.
2567. The fat is the "residuum" or the antagonism of the process of evaporation or water-formation. It is therefore deposited everywhere on the water-organs, beneath the integument, along the intestine and the vessels, around the kidneys, and so on.
2568. It is, like the adipocire, a product of the aqueous decomposition. On this account it is subservient to the process of temperature. It is the only isolator of heat, while all other animal bodies are conductors.
II. _Functions of the Systems._
2569. The functions do not concern the whole body, but only its principal parts.
a. VEGETABLE FUNCTIONS.
2570. The vegetative functions are matter-changing processes, which, consequently, correspond directly to the chemical processes, or are rather of one kind with these, but altered by organic elemental bodies.
1. _Functions of the Intestinal System._
2571. The general function of the integument consists in the secretion of juices, of which there are principally two kinds, the general and special.
2572. The general intestinal juice is mucus; it acts by rarefying.
2573. The gastric juice is of an aqueous, mucous, and acid nature. It acts as an acid and is endowed with chemically solvent properties.
2574. The bile is of a basic, inflammable, alkaline nature. It acts also chemically by analysing and precipitating.
2575. The saliva is the gastric juice of the head. It is a juice _secreted_ under the influence of the sensibility, and is on that account indifferencing, and nullifying in its effects; it is the highest poison.
2576. As the indifferent saliva precedes the gastric juice, so does the indifferent pancreatic fluid the bile.
2577. The proper function of the intestinal system is the digestion with all its divisions. There is an animal and a vegetative, or oral and abdominal digestion.
a. _Oral Digestion._
2578. Oral digestion is a mortifying or putting to death of the food.
2579. Since what is organic only serves as aliment for the animal, but nothing can be assimilated to the latter, without its having been previously reduced to the original condition of Infusoria, so also must the first act of the digestive process depend upon this, or converting the organic into primary organic bodies.
2580. This reduction to the primary condition is a putting to death of the organic individual. Organisms only which have been killed, can be converted into infusorial matter, and are then nutriment for the animal. The first act of digestion is consequently an act of putting to death.
2581. The act of killing consists of two moments, the mechanical and dynamical, or in lacerating and poisoning.
_Laceration._
2582. The mechanical act of putting to death commences with the search after nutriment, and thus with the movement of the feet; to this succeeds the prehension, or seizure with the claws or hands.
2583. This motion of the limbs is then repeated in the cephalic members, or the jaws.
2584. The seizure of the food with the cephalic arms or jaws, is the infliction of a wound commensurate with the position and form of the teeth. The teeth are digits of the cephalic limbs, or being devoid of any fleshy layer, claws. A gripe with such digits is in itself the infliction of a wound. For, in order that the food be grasped with only sufficient firmness to admit of its being drawn into the mouth, the sharp digital points of the mouth, i. e. the teeth, must make an incision therein.
2585. To seize, bite, wound, and kill, is in an animal one and the same act. For, so soon as the food or prey is seized, and consequently wounded, it is drawn deeper into the mouth, and therefore a new grasp and bite is essayed.
2586. Upon this the food is twirled about in the mouth by the muscles, or is chewed, whereby the parts are separated into mechanical atoms.
_Poisoning._
2587. Mechanical atoms are, however, not yet dead, because the vital polarity is not extinguished. It is requisite for a dynamic act, which suppresses all galvanism, to operate upon them.
2588. As this act of killing has directly for its object the death of the Organic and that alone, so is it a venenation or _process of poisoning_. To poison is not merely a chemical act, but one having for its immediate purpose, to separate the connexion of the organic atoms.
2589. The first act performed by the animal is that of poisoning.
2590. Poisoning, as being a direct aggression upon and destruction of life, is a destruction of the galvanic process. Now, the galvanic process is destroyed by suppression, destruction, and separation of its poles. The demolition of poles is a _polar equilibration_. Poisoning is equilibration of poles on the galvanic organism, or their neutralization.
2591. The primary change of poles is, however, in the blood. Poisoning is a balancing of the blood-poles.
2592. Indifferent blood is no longer blood, but chyle or infusorial primary mass.
2593. The nutritive fluid or chyle is poisoned organic substance. There is no aliment except what is wrought by poison.
2594. The poisoning of the aliments must take place through the medium of their blood, or their sap. If poison therefore does not get into the blood, the death does not ensue. The digestive juice is poison only for the blood, but not for the other systems.
2595. Chemical bodies which induce death, act by destruction of the bodies themselves, not merely by their neutralization or excessive polarization. They do not operate very differently from iron when in a state of red heat. Such is the case with sulphuric acid, nitric acid, alkalies and arsenic.
2596. Now, if these bodies be called poisons, there are thus three kinds of poisons, mineral or chemical, vegetable and animal poisons.
2597. The chemical poisons destroy the mass; they convert the Organic into minerals.
2598. Vegetable poisons reduce the animal to the plant; they do not effect the destruction of the mass in a general sense, but only that which is purely animal, or the nervous system--they are nervous poisons.
2599. The animal poisons destroy what is vegetable in the animal, or the galvanic process--are blood-poisons.
2600. _The saliva is the digestive poison._
2601. The saliva is not present, in order to macerate, and so prepare the dry aliments for digestion, but to _poison_ them. Everything else is only a subordinate operation. Now, the process of poisoning takes place only in the blood. The saliva does not act without inflicting a wound.
2602. Strictly regarded, all saliva is poison. There are examples of the saliva of birds, and even of men when inflamed with passion, having acted as poison.
2603. All other animal poisons, contagious principles or miasmata, are analogous to the saliva, being partial salivæ, partial poisons. Diseases of the skin produce tegumentary poisons or salivæ; glandular diseases, glandular poisons; nervous diseases, nervous poisons; diseases of the lungs, pulmonary poisons. The saliva is the blood-poison; the hydrocyanic acid, the nervous poison.
2604. During mastication saliva imbues every atom of the food, whereby the mass is throughout neutralized, or annulled.
_Docimasy or Testing._
2605. The oral digestion or that of the mouth cannot, however, occur by itself, without its being a nervous process also. The intestinal function taken up into the nervous system is the _taste_.
2606. Taste is the chemism resident in the head, the digestion in the nerves. Taste therefore happens according to the theory of the digestion or the chemism.
2607. The lowest element of the chemism is the water, the highest repetition whereof in the earth is--the salt. The former is therefore the object of abdominal digestion, the latter of nervous digestion. To taste is to digest salt. In order to taste the salt, every part of the food must be brought in contact with the tongue; this is only possible through comminution of the aliment by means of the teeth. The mastication dissolves the aliments mechanically into atoms, just as digestion will resolve them chemically into "infusoria."
2608. That which should be digestible, must have a saline character, must be soluble. The tongue is accordingly the test-organ of the digestive process.
_Deglutition._
2609. At first the tongue tastes only with its apex or tip; but after the salts which operate upon the tip are neutralized and examined, the tongue will also taste with its root, the two extremities of the tongue and their gustatory sensations being thus mutually opposed. It therefore takes the morsel upon the root and presses it against the palate, whereby the pharyngeal muscles clasp and swallow it.
2610. In deglutition the object of the tongue is not to give over or surrender the morsel to the pharynx, but to enjoy it "per se." During this fruition, however, it is robbed of it. Deglutition is therefore a result of an undesigned mechanical contrivance. Each organ works for the other, while fancying that it works for itself, which again it actually does, while it reaps the enjoyment so derived. Has the tongue finished tasting the food, then the pharynx obtains it involuntarily.
2611. To the saliva, and thus to a morsel, the acidulous gastric juice is polarwise related; both therefore seek to unite, and with this their organs also, namely, pharynx and stomach. The stomach obtains the preponderance, because it is acidulous; the pharynx moves towards it and with it also the morsel.
_Rumination._
2612. If the food, when received into the stomach, continue acid by virtue of its nature, or from not having been properly chewed and imbued with saliva and so neutralized, it is then homonymous with the gastric juice. The stomach therefore seeks to neutralize it, by restoring it again to the action of the saliva.
2613. Acid aliments cause vomiting. Grass which has not been masticated, and therefore enters acid or non-killed, and susceptible of fermentation, into the stomach, is regularly brought back into the mouth, and to the saliva, i. e. is ruminated or rechewed.
2614. Rumination is a regular act of vomiting, which has originated from the antagonism of the saliva and gastric juice and from the acid nature of the morsel.
b. _Intestinal Digestion._
2615. The intestinal digestion is the perfect chemical process taken up along with all its moments into the animal.
_Gastric Digestion._
2616. The stomach exercises by means of the spleen the oxydizing process of the intestine, the solvent function, and thus the action of the water. The gastric juice is related to the food, like water is to earth. The gastric digestion is liquefaction, unto which oxydation makes the preliminary step.
2617. Through the process of liquefaction, the poles are only potentially augmented, but are not dissevered, nor new substances formed. The gastric digestion creates no new bodies, but only mixes the old in the most homogeneous and intimate manner with each other.
2618. The gastric digestion is an animal process of fermentation. In deranged states of the stomach there is therefore a propensity to acid eructation, development of carbonic acid, and even formation of sugar.
2619. This digestion consequently reduces the (animal) food to the signification of the plant. Gastric digestion is a process of vegetable germination. The salivary process is a reduction to the animal death, digestion a reduction of this dead something itself to a lower kingdom.
2620. The gastric juice alone, like an acid, effects the solution of the aliment; with this, the movement of the gastric walls, which only tends to produce an easier mixture of the alimentary particles, having nothing to do.
_Hunger._
2621. Through the digestive process the gastric juice is consumed by the food, and the stomach deoxydized. If there is a deficiency of dephlegmatizing and deoxydizing aliments, then the peroxydation of the stomach must produce a feeling--called appetite. If this be not appeased, the oxygenic tension in the stomach is elevated or increased, and then begins to become unpleasant; this is _Hunger_.
2622. Here the feeling of the stomach's peroxydation is an obstructed process of fermentation, dependent upon want of food and alkaline principles.
_Thirst._
2623. The feeling of the reverse condition to the above is _Thirst_. It originates through a too rapid deoxydation of the stomach, through deglutition of the gastric juice on account of an excess of food. But it may also originate from a deficiency of gastric juice, or from an alkaline tendency in the latter; just as hunger resulted from a superabundance of gastric juice, or a disposition to form acids.
2624. Thirst is the feeling of too powerful a digestion, or of too rapid fermentation, whereby the product or leaven of the latter becomes, as it were, bankrupt; upon this the blood flows in greater quantity, in order to secrete the gastric juice; the arterial nature becomes elevated, and finally, an inflammatory condition, associated with a sense of dryness, originates, and is propagated as far as the mouth.
2625. Thirst and heat rank, like hunger and cold, parallel to each other. The feeling of dryness appears as heat, that of moisture, as cold. Cold therefore at once extinguishes or quenches the thirst; but produces hunger, which again is mitigated by heat.
_Biliary Digestion._
2626. The stomach is the pulmonic intestine; the duodenum, through its combination with the liver, is the vascular intestine, having a predominant venous character; or the one is the arteriose, the other the venous stomach.
2627. The gastric digestion has reduced the food to the state of a vegetable; but this does not yet suffice for it to become a nutritive matter, namely, a new organism. As the primary organism originates, so also must the body originate in the process of nutrition, and thus from the element of the Organic. Now, this primitive matter or element is the mucus, the "infusorium." There is still therefore a process necessary to redissolve the vegetable and convert it into protoplasma or primary mucus.
2628. This, however, occurs only through the process of putrefaction being wrought in the food. This process is not simple solution, but decomposition, or separation of the constituent parts. The acid vegetable parts, which had been active through the process of fermentation, must be therefore decomposed, and in such wise, indeed, that what is mucous shall be separated from them.
2629. This separation or analysis takes place by means of the _Bile_, which combines with the acid of the chyme.
2630. The biliary digestion is an alkalizing or saponaceous process. Through it the chyme becomes separated into nutritive juice or chyle, and into excrement.
2631. The excrement is what is vegetable, or the product of fermentation combined with the bile. The chyle is what is infusorial, or the product of decomposition. Nutrition therefore begins anew. Its process is a "generatio originaria."
2632. In digestion the processes of both organic kingdoms, namely, the fermentative and putrefactive, are repeated.
_Absorption._
In the intestine, not merely lung and vascular system, have been represented, but also the integument or organ of absorption.
2633. After the animal food has been reduced to the vegetable, and this again to the infusorial state, it can be taken up by the body. It now becomes absorbed in the small intestine. The small intestine is the tegumentary system, or the root-bark.
2634. The chyliferous or lacteal vessels stand in antagonism with the lung, or the skin as being the original respiratory organ. It is only, therefore, the infusorial chyle that has been absorbed, not the excrement, because between the latter, as the product of oxydation, and the lacteal vessels, repulsion takes place. The chyle, having been absorbed, enters into the thoracic duct, and from thence into the lungs.
_Evacuation._
2635. Through the absorption of what is fluid, that which is excrementitious becomes more solid, and is thus given over or transferred into the vegetable, sexual, or large intestine.
2636. The excrement is now found in another, i. e. in a lower, or vegetable animal. It therefore obtains the direction of all sexual secretions; it is thrown out or ejected, and in a reverse direction, because the anus is the sexual mouth.
2637. Digestion is thus through all predicaments, from its incipient dealings with the highest life unto the plant, and from this to the mucous globule, a thorough process of putting to death.
2638. The nutritive will be through all predicaments, from the infusorium to the plant and to the animal, a thoroughly vivifying or life-inspiring process. Digestion is descension, nutrition is ascension.
2. _Functions of the Respiratory System._
2639. The branchiæ and lungs are the air-organ of the animal, the foliage. The animal, like the vegetable foliage, is oxydized from water or air, by which means the animal sap, which hitherto is only a root-sap, becomes differenced into an aerial sap.
2640. No animal can live without oxygen gas, because the air is the condition of the galvanic process.
2641. The oxygen passes over materially into the blood or the chyle. Beyond this it is an indifferent matter for physiology, whether the blood simply derives the positive tension from the air, or combines the positive oxygen materially with itself. In both cases the same heterogeneity originates. Were oxygen, however, not to enter the body through the lung, it could not then be seen, whence its ingress might be effected. In other respects, every change of matters is established with material combinations and separations.
2642. The carbonic acid of the respiratory process may originate accidentally, as when it is formed on the integument. In so far is the expiration one and the same with the evaporation.
2643. The chyle ascends directly from the small intestine into the leaf-fabric, or into the lung. Both organs are to be compared with an entire plant, whereof the intestine is the root, the lymphatic vessels the stem, the lung the leaf-fabric or foliage.
2644. The chyle moves in the lymphatic vessels like the vegetable sap, and thus by polar tension between lung and intestine. Lung and intestine have been diametrically opposed. The conductors of the antagonism are the lymphatic vessels.
2645. The lymphatic vessels do not absorb by virtue of their own contractility, nor by open ends; but by physical pores like the cells of plants.
2646. It is the chyle, which is oxydized in the lung; the venous blood is a subordinate object.
2647. Through the oxydation it becomes coloured like the vegetable sap does in a leaf. But the latter sap obtains only the colour of the terrestrial oxydation, of the terrestrial oxyde or the water, and becomes green; while the chyle gains the colour of the cosmic oxyde, or the fire, and becomes red. The red, fire-coloured sap is the blood. The function of the lungs is consequently a formation of blood.
2648. The blood is an infusorial, a green mucous sap, which has been elevated to what is aerial. The blood consists of aerated Infusoria--blood-globules. These are the red substance of the blood. In the lower animals only, where the blood, like the body, contains but few blood-globules, is it colourless.
2649. Through the respiration an electrical difference enters the blood, whereby it becomes separated into several substances.
2650. The _water_ in the chyle is the inorganic menstruum, wherein the mucus, as being a fundamental matter of the vegetable sap, has been dissolved. Through the digestion the mucus becomes animal--_gelatine_.
2651. The gelatine is the basi-constituent part of the chyle, being a product of the digestion or the root-process. Gelatine is the vegetable mucus repeated and ennobled in an animal. This ennobling took place through the digestion.
2652. "En route" too, the chyle mixes with the nervous blood, and already undergoes a degree of oxydation, whereby the gelatine becomes converted into albuminous matter.
2653. Through the respiration gelatine and albumen are elevated to aerial importance, and still more oxydized; by this means the albuminous matter becomes elevated, and passes over into _fibrine_. The fibrine is the last product of respiration. Fibrine is the starch-meal in an animal.
2654. The electric duplicity, induced in the blood by the process of respiration, exists accordingly between fibrine and gelatine, while the albumen constitutes or forms the indifference. The fibrine is repelled from the lung, because they are both homonymous; it is the oxydized gelatine. From its communicating its polarity to the whole mass of blood, this also is repelled from the lungs.
3. _Functions of the Vascular System._
2655. The capillary vessels of the body range opposite to those of the lungs, just as the biliary does to the splenic stomach, as alkali to acid, as precipitant and secernent to what is non-separated.
2656. The capillary vessels therefore attract the pulmonary blood, separate it, secrete and form new constituent parts; and then, after it has become homonymous, they repel it back again towards the lungs.
2657. The circulation only subsists through the polarity which exists between lung and capillary vessels, between lung and body, between oxydation and reduction, combination and separation.
2658. If the intestine and body be the root, the lung the foliage, so is the lymphatic and vascular system the stem, in which the most perfect substances have been formed. In the circulation the matters for the animal systems must have been fully formed, such as for the bones, the lime; for the muscles, the iron; for the nerve, the albumen.
2659. The liver is the principal organ of the circulation. There also must one principal preparation of the blood take place. On account of its parallelism with the brain, the condition probably resides in it that conduces to formation of the albumen, namely, of the Indifferent, which is the nutritive matter of the nervous mass.
2660. In the vegetable sphere of life there are, in addition to the liver, but two mutually distinct excretory organs, namely, the intestine and the skin.
2661. In the mucous secretion of the intestine the condition, requisite to the formation of phosphate of _lime_ for the bones, probably resides.
2662. In like manner in the formation of the carbonic acid and the water in the integument, the basis for the formation of phosphate of iron for the muscles may reside. In the circulation the intestine would thus be the lime-, the skin the iron-, and the liver the medulla-forming organ.
2663. The two extremes of the circulation, or intestine and lung, form gelatine and fibrine; the circulation itself forms the purely animal matter. Out of the integument and lung grows the muscle, from the intestine the bone and gelatine, out of the liver the nerve. Muscle is integument and air, bone is intestine and chyle, brain is liver and blood. Thus each has a function that is peculiar to it; each organ has its business to perform in the diffuse fabric of the animal body.
2664. Through this variety or change of the offices, the circulation first becomes possible.
2665. If the secernent process be therefore suppressed, the animal then dies as rapidly as if it had been suffocated. It is a suffocation of the opposed pole. Query? does not many a fit of apoplexy depend upon this?
2666. The circulation has consequently two factors, the lung as oxygen-pole, the capillary vessels of the body as hydrogen-pole, the blood as the indifferent water. The circulation is a galvanic process.
2667. In all extremities of the body the arteriose blood becomes deoxydized, decomposed; it is therefore basic and homonymous with the capillary vessels, so that it is consequently repelled, and driven back into the veins.
2668. It can, however, flow nowhere else than to the lung, because there resides its opposite pole. Being again oxydized in, it becomes homonymous to, the lung, is repelled by it, and again attracted by the capillary vessels of the body.
2669. The circulation is therefore a result of dynamic forces, not of mechanical functions. It would occur, were the vessels to be glass tubes.
2670. The pulsation of the heart is not a cause of the circulation, but inversely rather, its consequence or effect.
2671. In the circulation the whole organism, or intestine, lung and integument, is combined. It is therefore the fundamental system, which includes the whole mass of the body.
b. FUNCTIONS OF THE ANIMAL SYSTEMS.
1. _Of the Osseous System._
2672. The functions of the osseous system are simply mechanical relations, such as solidity, form and motion.
2673. The motion of the joints presents to our notice interesting relations, especially in reference to the motion of the vertebræ, ribs, limbs and jaws upon each other, but these could not be here treated of in detail.
2674. The acts of swimming, creeping, standing, walking, running, leaping, climbing, and flying, do not exhibit simply mechanical, but truly philosophical moments.
a. _Bodily Motion._
2675. Swimming can take place through simple contraction of the body, without locomotive members. It is the continuation of the vesicle's first process of origination; as in the Infusoria and Polyps. In the Worms and Serpents it is effected by an undulating motion of the body, whereby the water is struck with oblique surfaces; there it is fibrous motion. In the Holothuriæ and many aquatic larvæ, it is a propulsion, effected by expelling water from the anus, and thus by squirts, there being consequently contraction of the body or arteriose straitening of the sphincter muscles. In Fishes, swimming is a rowing or remigial stroke, produced by lever-motion, together with the oblique slap made by their tail.
2676. Creeping is either a shortening of the body by fibrous motion, as in the Snails, or an undulatory motion, as in the Serpents.
b. _Pedal Motion._
2677. Standing is the position of a lever "in equilibrio," the creation of the proper centre of gravity.
2678. Walking or running is an exchange or alternation of the equilibrium, a combination of standing and creeping.
2679. Leaping or hopping is a flight with the feet.
2680. Climbing is an use of the feet as hands.
c. _Alary Motion._
2681. Flying is lever-swimming in the air. It takes place by means of aerial branchiæ, namely, the wings in Insects, and by the thoracic extremities in Birds, which are also none other than animal branchiæ.
2682. Soaring is creeping in the air.
2683. Hovering is standing in the air.
2684. Pouncing is hopping in the air.
2685. Diving is hopping in the water.
2. _Functions of the Muscular System._
2686. The muscular system performs, in an active sense, what the osseous system does in a passive. The strength or power of the muscles, and their leverage attachment, is here especially to be regarded. The contraction of the fibres is a _charging_, by nerves and blood, of the two fibre-poles.
2687. The fibres are charged by the air. It is, in the most general sense, the respiratory vessels by which the muscle is charged. This is strictly the case in Insects, where the tracheæ traverse all the limbs, and directly conduct to the flesh the polarity of the air. In animals, however, with a closed circulation, the arteries undertake the conveyance of air upon the blood, and it is then the latter fluid which streams into the muscles in order to charge them.
2688. Thus if an artery be ligatured, the limb is crippled or lamed. The artery, however, imparts only the positive pole, and consequently of itself produces no shortening or contraction of the fibres. The oxydation takes place at the lower end of the muscle; here, therefore, the latter passes over into tendon.
2689. The nerve is the second condition of the muscular contraction, since it evokes in the fibre the negative pole. Thus if a nerve be ligatured, the limb is likewise motionless.
2690. If the poles be brought by contraction into close approximation, the fibres must re-extend, so soon as the influence of the blood or the nerve ceases.
2691. Now since the blood is constantly streaming in, the reason for the muscular rest must reside in the nerves. The rationale of voluntary motion is consequently the nerve. The relaxation or extension is an unloading of the fibres.
2692. The muscular motion is an electrical process, a motion of blood in the Solid.
2693. Through the polarization of the fibres the muscle is formed from the arteries. The muscle is therefore an individual biconical piece of fibre, having unequal cones. Oxydation takes place at the muscular extremity; here, therefore, originates the sinew or tendon.
2694. A fleshy cyst--or heart--which includes an osseous cyst, must subdivide into several fibrous cones or muscles. One reason of this is the "fore and aft," another is the quantity of the essential vascular branches.
2695. The muscle contracts upon application only of a stimulus.
2696. Every stimulus induces motion only as a result of polar excitation. Every stimulus polarizes; for even the gentlest contact is like the friction, and produces electrical antagonism. It therefore amounts to the same, whatever stimuli, whether mechanical, chemical, or spiritual, have been applied to the muscle. One acts like the other.
2697. If no motion supervene upon, or is even suppressed by, the contact of a body, the nature of the body must then be _indifferencing_.
2698. Relaxing, laming, life-destroying matters, are indifferencing, or cause a suppression of the poles.
2699. Overcharging principles, e. g. lightning or strong electric sparks may also produce relaxation. These destroy the function of the fibres, and act therefore worse than the in differencing matters.
3. _Functions of the Nervous System._
2700. The function of the point-substance is also that of the nervous system, for this is only the point-substance, fashioned and arranged into stalk and branches.
2701. But even on this account the nervous tension proceeds only according to a determinate line, while before it penetrated through the whole mass.
2702. The nervous tension takes place in a nervous system only between a special organ and the nervous centre.
2703. In itself the nervous system is an Indifference, and such then are all the organs upon which it acts, when regarded in reference to this operation; they might, through other functions, be polar.
2704. The nervous system becomes differential either through its two kinds of substances, or through extraneous influence; in the first case it thinks or moves, in the second it feels.
2705. Sensation is in the nervous what motion is in the fibrous system, namely, a polar condition, in which the two ends have a tendency to come together, in order to discharge or unload.
2706. In the sensation the extremity of the nerves strives to approach that of the brain; it is therefore a contractile effort in the nerves, like motion is in the muscular fibre. Juxta-posited granules could not, however, shorten; therefore, the already calculated polarity courses over them.
2707. All external stimuli act upon the nerves and polarize them, but not the muscular fibres.
2708. The fibre becomes directly polarized only through the internal stimuli, or by the blood, galvanic tension, &c.
2709. The most general function of the nervous system consists in its assuming polarity from the world, and imparting this to other systems, apart namely, from the alternate operation of its two substances.
_Nervous and Motor System._
2710. The first system is that of motion. The tranquil presence of the nerve in the fibre produces indifference in the latter; so that it has no susceptibility for the polarization that is effected by the artery. Doubtless the arterial blood passes, for the most part, close by the fibre through other capillary vessels.
2711. But if the nerve be polarized it is then basic, negative, and enters into opposition with the arterial blood, which now streams into the capillary vessels of the fibre, and renders the two ends of the latter heterogeneous.
2712. Thus, if the nerve be cut across, it remains always indifferent, and no motion can any longer ensue. But this does ensue so soon as the nerve is galvanized. A proof that the nervous influence is homologous to the galvanic tension.
2713. If the nerve becomes involuntarily negative, then originates spasm.
2714. If the nerve continue morbidly indifferent, then paralysis originates.
_Nervous and Vegetative Systems._
2715. The second great system is the tegumentary formation. In this too its ordinary functions occur, such as secretion, evaporation, nutrition, elevation or depression of heat without nervous influence; or they occur while the nerve only acts indifferently.
2716. But if it acts by polarizing, as in the muscular motion, then the tegumentary processes are at once changed. What is material becomes extinguished, and the simply Irritable is manifested in the cells, capillary vessels, and so on.
2717. The process of temperature becomes thereby instantly changed, because the decomposition is changed. The heat is increased by the rapid suppression of the evaporation; it is diminished by the rapid increase of the latter.
2718. The art and manner in which the nerves act upon what is vegetable, is consequently an elevation of this Vegetable to what is animal. It must resign its processes, and simply undertake those of motion. Exactly the same takes place in the secernent organs. They perform their offices themselves through the galvanic tension of the vessels. But if the nerve act upon them they secrete more powerfully, and exactly from the same cause whereby the muscle is self-moved. Thus, if the nerve operate feebly, whereby it is in an irritable state, or if its action be wholly withdrawn, then the secretion will become less.
2719. The more irritable the nerves are, by so much the more animal does the vegetable organism, or the splanchnic system, therefore become; so much the less is it produced on the mass.
2720. Irritable animals and men are therefore meagre or lean. The two cases are uniformly related to each other. Lean men are irritable, not perhaps because the nervous ends are not covered with fat, such as there might be over the integument, the ear, the tongue, nose, &c., but, because where the nutritive process is inactive, what is animal necessarily preponderates. There are consequently three kinds of nervous actions, or phenomena, though all nervous activity in entire systems is similar, or but one. The difference is only derived from the organ upon which the nerves act. There are, consequently, no special nerves of sensation, motion and secretion; or such, forsooth, as have only one of these offices to take care of. Thus, were a nerve of sensation to be prolonged or drawn over a muscle, it would excite motion, and in the case of the liver, secrete bile, &c.
_Mesmerism._
2721. When the sensibility mounts to the highest degree, each mass-function will then almost cease, and the organs of sense feel the weakest operation of the stimulus.
2722. Since every stimulation is a polarizing act, and each body is in polar activity towards the other at every conceivable distance, so may an extremely irritable nervous system also perceive the feeblest polarizations.
2723. The eye perceives the polarization at a remote distance from the body whence it proceeds.
2724. In hearing the vibrating body still indeed acts directly upon the ear through the vibrations of air. But a finer, i. e. more irritable ear, hears farther than one that is more dull.
2725. With an elevated sensibility the other systems also may therefore perceive the polarization of the bodies, without coming into contact with them.
2726. To perceive objects in the distance, i. e. merely their polar influence, is called _Mesmerism_, or animal magnetism.
2727. Now, if it is once possible for other senses besides that of sight to extend their perception into remote space, it no longer matters as to the magnitude of the latter. A feebly charged electrical machine only attracts bodies that are near, one strongly loaded, those which are more remote; such is the case too with weak and strong magnets.
2728. The integument at once perceives electrified surfaces at certain distances; now, as every surface appears electric to the integument, so must the latter, if its sensibility is very much elevated, perceive something of that sort in every proportionate distance.
2729. But homologous polarities only act upon each other, and therefore traverse thoroughly through heterologous bodies. Thus the magnet attracts the iron filings through the table-board, unhindered by the interposed wood, and without any regard being paid to this, or its being even perceived.
2730. The senses may therefore perceive their homologous polarities through other bodies, walls, and such like. By virtue of their perception, they stand in relation to them.
2731. To the very sensible nervous system the vegetable system and its impulse is a foreign object, which detaches itself from the above system, just as the objects of sense have done from the sensorial organs. The vegetable, or in general the material body, appears therefore to the mesmerized like a strange world--they see their own organs--are _clair-voyants_. Mesmerism therefore comprises nothing which could contradict physiology.
_Vegetative Nerves._
2732. The splanchnic or visceral nerves are also distinguished from the animal by their being in a constant state of tension, and hence keeping the processes of their system in constant repair.
2733. The rationale of this resides in the two nervous substances having separated into ganglia and plexuses.
2734. This also serves to explain another phenomenon, viz. that the nerves indeed, but never the brain, attain to perfect rest; because the former is without ganglionic or cortical substance, the latter is throughout surrounded, and obviously interwoven, by it.
2735. Therein lies the reason why the viscera do not sleep.
_Sleep._
2736. The condition of the nervous system, as hitherto represented, is called that of being awake. It is the interlude played by the nerves with the world, and with the animal body.
2737. When in a state of health the first interlude ceases, then the other also is over or past. The world-nerves, however, operate only upon the animal systems, upon the senses and the motion of the muscles; it could therefore be these only wherein the nervous function, in compliance with this cessation, is suppressed. Now, muscular rest originates through suppression of the tension between nerve and muscle. In sleep this rest is also derived from the same means.
2738. The tension between nerve and muscle can only cease, if the tension also between the termination of the brain and nerves be suppressed. We are now reduced simply to the consideration of the nervous system, and may, in treating of sleep, pay no attention to the muscular system.
2739. But whence comes the tension in the motor nerves? Obviously only from the cerebral tension. This can originate only in two ways; either through the special organization of the brain, one or the other substance being preponderant, or through the influence of external stimuli.
2740. The encephalic substance becomes more potent than usual, if by rest the cortical substance becomes more arteriose in character. This tension is communicated to all the nerves, sensitive as well as motor, and continues in their interlude with the world and the motor system.
2741. If this encephalic tension is not too potent, it remains only in the brain, without the ability to polarize the nerves also. It then only produces cerebral phenomena, thoughts or _dreams_.
2742. Dreaming is an encephalic tension excited by the organization, not by the world.
2743. Dreaming is the first step in the liberation of the animal from the vegetable system--it is the first step towards mesmerism.
2744. In a perfect or middling state of health, where the nervous is not very much separated from the tegumentary system, we do not dream.
2745. In a healthy condition an external stimulus would be, consequently, the only cause of waking, did not the long repose bestow a preponderating influence upon the cortical substance. Dreams therefore happen in the morning.
2746. Waking is the intercourse with the world, not with self. If one wakes also from intercourse with self, still the former is synchronous and coexciting.
2747. Thus, if intercourse with the world ceases, sleep originates. If the vegetable intercourse with it also ceases, then death originates. Waking is "consensus" with the world.
2748. Sleep is a death of the _animal_ systems.
2749. Every awaking is a resurrection from death, a new sympathizing with the vegetal body, from which the animal body again originates.
2750. As the animal originally took its rise from, and only through the plant, so also is this repeated in an individual. The plant is the ever-living, ever-verdant, or green, out of which the animal daily sprouts forth as a blossom.
2751. The animal intercourse with the world is also interrupted in two ways, and there are therefore two modes of falling asleep.
2752. The first cause resides in the want of stimulus. The nerves of the senses are not polar, do not therefore excite the brain, nor does this again affect the motor system. The muscle therefore arrives at a state of non-tension; it becomes relaxed, and along with it necessarily the organs of sense, which are thrown into activity by muscular motion. The arms and fingers, whose business is to touch, sink down; the feet which move, and thereby warm and animate the body, are slackened and bent together; the body is in the recumbent posture; the eyelids drop, the light no longer plays upon the visual organs, the external and internal auditory muscles flag also, and the sound is no longer borne upon the ear. Now also does the tension of the senses with the brain cease, and with it the sensation--there is sleep.
2753. This sleep arising from want of stimulus is a faint sleep, and rendered useless by dreams. For there is actually no cause present why the encephalic tension should entirely cease. Men, who do not fall asleep through fatigue, but from want of work, sleep restlessly, awake easily, and again readily fall asleep. Their life is dreaming.
2754. The other cause of the polar suppression in the nerves is like that of the extension of the muscles, or their falling to sleep; it is thus the discharge of the too strongly excited poles. With too high a degree of fibrous tension, which also originates through too long a continuance of the tension, the fibre is placed in a state of activity, which consists in the antagonism being balanced by approximation of the ends. Were nerves, when greatly tensed, capable of being shortened, they would also discharge themselves, and come at least for one instant to rest--they would _sleep_.
2755. The falling asleep of the fibre is its sleep, though it also does not last long. So is the expansion of the heart its sleep, so is expiration the sleep of the thorax.
2756. In all polarizable organs there is a change or alternation of waking and sleeping, which endures a longer and shorter time. This periodicity depends upon the energy of the polar influence, and upon the size and susceptibility of the substance.
2757. Every substance has its own periods of waking and sleeping, of action and repose. The pulse sleeps shorter than the breathing; this again shorter than being hungry; this again for a briefer time than the sexual function.
2758. There are organs, or systems, which are nearly always in a state of slumber, e. g. the osseous system, because in it the polarity is extinguished. It is only in states of inflammation that it wakes up. Others scarcely ever slumber, e. g. the cellular system, because in it indeed no pole is yet fixed, and in the alternation of poles its life consists.
2759. There is a similar change of poles in a nervous system, and it indeed halts or stops for a middling time. Through the persistent influence of the external world, the nerves of the senses are thrown into such a state of tension with the brain, that blood cannot flow thither in sufficient quantity, in order to maintain the two cerebral substances in mutual antagonism. Brain and nerves become therefore indifferent; muscles and sensorial organs lose their polarizability, and their intercourse ceases with the nerves as well as the world. The brain, and everything else, has now been discharged, and a deep sleep without dreams, an animal death, ensues.
2760. It has been arbitrarily asserted, that no sleep is possible without dreams, but for this statement there are no existing grounds. Whence should the dream come if there is no tension in the brain, or if it has previously undergone sufficient self-exhaustion?
_Periodicity._
2761. The sleep of the nerves ranges parallel with that of the planet. It might be said that such was the case from habit or custom, but it is, properly speaking, dependent upon a parallel process of organizing that occurs at the origination of the animal. The matter stands thus: the germ originates in the morning; until evening stimuli act upon and polarize it: in the evening they cease, and with them the tension. The muscles relax upon every movement, and rest necessarily follows in the manner above delineated. With to-morrow's morn the world again acts until evening upon the germ; it wakes up, and the same course of events happens as on the first day. Finally the substance becomes organized according to this periodicity; it becomes, forsooth, not more energetic than is necessary in order to admit or receive a charge of one day; towards evening it is exhausted, is neutral, and rejoices with the relaxation of the muscles, that the world no longer acts upon its frame.
2762. Thus, we may attribute this phenomenon both to custom and synchronous formation; nevertheless one ought not to forget that the organic formation is a metatype, or has been imitated from something that has gone before, and that consequently the law of periodicity has not been bestowed upon both at the same time, but first of all unto nature, and through this to the body, which is its image or likeness.
2763. The nervous periodicity ranges in accordance with the periodicity of the light, or is thus parallel with its archetype, and consequently with the day and night. Day is the waking, night the sleeping of nature. But the animal has originated in and by this alternating change of nature. It is spiritually, as well as corporeally, nature's likeness.
2764. The sexual function in a perfect animal, as in Man, has been adapted to the periods of the year; in other animals other natural periods exert their sway. Animals are commonly pregnant by the month, e. g. for one, two, four, five months, &c. The human species requires three quarters of a year for pregnancy, one quarter for giving suck to the babe, and then it can again conceive. Pregnancy thus lasts a year, and has been based in the sun.
2765. If the female does not become pregnant, the sexual passion, or instinct, is repeated at the expiration of a month. It sleeps about one month, and then awakes for some days. The periodicity may be divided into vegetative and animal, thus,--
a. Animal = _World-periodicity._
1. Sleep = _Rotation of the Earth._ 2. Menstruation = _Lunar revolution._ 3. Pregnancy = _Solar revolution._
b. Vegetal = _Earth-periodicity._
1. Digestion = _Water, Ebb and Flow._ 2. Respiration = _Air, Electricity._ 3. Pulse = _Earth, Magnetism._
_Awaking._
2766. The awaking takes place of itself through the origination of a new polarity in the brain during the afflux of arterial blood, whereupon the dreams follow which precede the act of waking. During sleep the plant continues to act, the unloaded cortical substance becomes again oxydized and charged; tension arises between it and the medulla, and with this come the dreams. This encephalic polarity is imparted to the sensitive and motor nerves, and the organs open. If external stimuli are superadded, all this happens somewhat earlier. The act of awaking is invariably, however, a restoration effected, or brought about, by the plant, and especially by the circulation.
2767. Man would therefore wake up had there also been no world of the senses; but then he would not continue awake, but at once relapse into slumber, and sleep the eternal death.
III. _Functions of the Organs._
2768. The functions of the organs are the functions of the system associated or combined, just as the organs are but ultimate evolutions of the systems existing under similar relations. The organic functions are always therefore in a state of concatenation with other systems, and there can be no organ which acts in an isolated manner.
2769. By this character a new field for sympathy has been opened. All the organs operate through sympathy. Sympathy is therefore the result of parallel systems, or also of antagonisms between the factors of a _single_ system. Taken in a strict sense, there are no vegetative organs. The organs are therefore limited to the _encephalic animal_, such as are those of motion and the senses, and to the _sexual animal_.
1. _Functions of the Encephalic Animal._
A. ORGANS OF MOTION.
2770. Just as the nerves have a function in themselves, and one directed towards the subordinate organs, so also has the motor system.
2771. The motor system is in the first place doomed to serve the whole body, since it flexes it, moves it forwards and backwards and upon all sides. It is related principally to the movements of the vertebral column, and serves in numberless animals to effect the act of crawling or creeping.
2772. Then again it will serve individual parts of the body, such as the belly in its evacuations, or the sexual animal in emitting urine, &c. It ministers unto the thorax in the act of respiration, which is a very complicated process. The thoracic muscles are to a certain degree co-ordinated with the constantly polar nervous system, and become thereby and in part involuntary. But one main reason of this appears to be the air that is constantly renewed in their cavity.
2773. In the act of respiration two orders of muscles are active, the proper-pectoral muscles, and the abdominal muscle, which has been displaced from the thorax, or the _diaphragm_.
2774. As originally the thorax took its rise at the expense of the abdomen, so also is every inspiration an elevation of the thorax and a displacement of the abdomen. Every breath or in-draught of air expands and produces the thorax, but narrows and arrests the abdomen. The diaphragm expresses this contest. Its contraction being a result of the respiratory tension, expresses consequently a preponderance of the thorax, and in obedience to this, narrows and diminishes the abdomen. It is supplied, in conformity with its origin, by nerves from the upper cervical vertebræ, since prior to this the abdomen extended as far as the head, and the branchiæ were appended to it as _lateral strips of integument_--as in Fishes.
2775. As the thoracic cavity is drawn by the diaphragm's descent towards the abdomen, so by the pectoral muscles is it raised upwards to the head. The latter movements are what is animal in the respiratory process, for they lift the thorax towards the head. Pectoral muscles and diaphragm stand opposite to each other, like limbs and trunk; the ribs are the limbs, the diaphragm the upper abdominal tunic or covering. Through this antagonism what is limbed, as belonging to the thoracic cavity, is drawn upwards, what is abdominal in its character, downwards; the result is expansion, and through this pumping in of the air.
2776. The air is in part voluntarily swallowed like the food, as e. g. by the movement of the ribs, in part involuntarily by the diaphragm. The diaphragm may be termed the heart of the abdominal cavity.
2777. In the lower organized animals, where merely branchiæ are present, the animal motion encroaches but little upon the act of respiration. In Molluscs and Snails, the oxygen is almost always on the branchiæ, as is the case also in Insects. In Fishes the water is still taken in like the food through the mouth, and driven out or expelled by the pharyngeal muscles from between the branchiæ. In them the air is in both respects swallowed. In many Reptiles the air is indeed drawn in through the nostrils, but conveyed into the lungs by a true act of deglutition. These kinds of functions are necessary, because as yet the whole trunk is abdomen, and the thoracic cavity has not as yet separated from it; hence the diaphragm is wanting.
2778. The respiratory originates therefore from the digestive organ; it is freely developed only from the abdomen, and is at length entirely liberated from the latter as a self-substantial cavity, e. g. first of all in the Mammalia. Respiration is originally but an act of deglutition, which has gradually become perfected, by the animal trunk being associated with it to a greater extent.
2779. The pumping in of air has therefore become in the highest degree a process of suction, and in this resembles the digestive function when it has become animal.
2780. Then also the air passes no longer through the mouth, but through the nose, as being the peculiar opening of the thoracic cavity in the head. Even the Fishes have as yet no nostrils opening into the mouth.
2781. The thoracic motion is a limb-motion, and would become locomotion, were the ribs not conjoined. In many of the lower animals the branchiæ are at once organs of motion, such as fins or rudders.
2782. Every inspiration is a self-manning towards the animal; every expiration a retro-depression unto the plant; the abdomen in this case regains the upper hand from the diaphragm reverting to its usual position, and the thoracic cavity narrowing. Respiration is a becoming animal.
2783. The motions in themselves, without reference to the trunk, are the motions of the limbs, as standing, walking, &c. The movements of the arms and feet are sympathetic, because their muscles are of equal signification.
B. FUNCTIONS OF THE NERVE-ORGANS.
2784. These functions have relation only to the nervous system itself, because all nerve-organs are elevated above the trunk, and live in themselves. They are simply the functions of the sensorial organs.
a. _Functions of the Vegetable Sensorial Organs._
2785. These must be regarded as those that still encroach upon the inferior organs. They are not, however, the inferior processes themselves, but their ascensive formations into the nervous system. This therefore works henceforward only in and by itself, but yet in relation to the inferior processes.
1. _Function of the Sense of Feeling._
2786. To constitute the sense of feeling the integument, in other words the nutritive or vascular system, has assumed a nervose character, and consequently that which is in communication with the materiality of the external world. The function of this sense will therefore have materiality only for its object.
2787. The integument is the organ by whose means the animal is absolved or liberated from the world. The sensation belonging to it, is none other than the perception of this diversity subsisting between the two.
2788. Through the tegumentary sense, the world becomes a something external in relation to the nervous function; while previously it was such through the medium of the skin for the lower organs only, viz. as an object of absorption. The discrimination of materiality is called _Feeling_. The sense of feeling is the earth-sense.
2789. The sense of feeling perceives materiality, like the nerves perceive all objects or all stimuli, through polar excitation. Every pressure, every contact is polar excitation.
2790. The sense of feeling is characterized by poles only being excited in it by absolute proximity or immediate contact. Just because it is the first sense, through which the animal is set free, so must that which is liberated be at once perceived in the moment of liberation, and thus in immediate contact. The sense of feeling is a polarity of contact, a polarity without distance. The stronger the contact, by so much the stronger is the polar excitation--there is increased pressure. The gravity acts simply by pressure. The perception is therefore resolvable into one of pressure or contact.
2791. Different degrees of pressure necessarily impart different amounts of feeling. Perception of the different degrees of pressure made by an object betrays its inequalities of surface. The sense of feeling is also the sense for determining inequalities, for the Soft and Hard, for the Solid, Fluid, and Gaseous; all these feelings, however, are referable to the contact.
2792. Through diseased conditions, the polarizability of the sensitive nerves may become exceedingly elevated, and they then perceive the polarity of contact prior to the contact having taken place. For the two bodies invariably excite poles that are antagonized to each other. Were other bodies not to approach them more closely, or else act upon them more energetically and so extinguish the polarity; they would remain at an infinite distance in a state of polar relation towards each other. Feeling can therefore be extended to an indefinite distance. Hysterical, mesmerized, and even healthy human beings, feel further than they grasp or touch with the hand.
2793. Homogeneous polarities, or those of the same kind, are throughout nature to be found also by means of others; e. g. electrical polarities are not disturbed by the intervention of magnetic ones. Such is the case also in feeling. That which is related to oneself is felt, although it is more remote from us than other objects, upon which we do not bestow any attention, or towards which we do not turn our poles.
2794. The sense of feeling differs according to the diversity of certain points in the integument, and is thus nobler in character, the higher the rank which these may hold. Thus it is most feebly developed in vegetable situations, where hairs, nails, claws, and scales are placed. It must attain the highest grade of perfection in the animal organs, and thus in the limbs or their parallels, the lips.
2795. In the limbs, by reason of their mobility, the feeling becomes voluntary. It is then wholly in our power to strengthen or weaken the contact, to press gently or firmly, and allow these periods of feeling to succeed each other rapidly or slowly.
2796. Feeling associated with motion is called _touch_; this condition of the organ the _tactile sense_. The tactile sense is by no means different from that of feeling; it is only the combination of feeling with motion.
2797. The fingers are the most perfect organs of feeling, because they are the most moveable parts of the body, and therefore they are _organs of touch_.
2798. As simple feeling perceives the asperities of bodies, so does touch the _forms_. The perception of forms is based upon the form that resides in the _tactile organ_ itself.
2799. All possible forms reside in the motion of the fingers.
2800. Each hand is a semi-ellipse, in which the four fingers are the periphery, the thumb the radius. Both hands together form a perfect ellipse with two radii.
2801. Now, in the ellipse are involved all geometrical figures. The hands include in their movements the whole of geometry.
2802. We can only perceive the forms of nature, because they all reside in ourselves, because we can create them. This, and none other, is the meaning or sense of the doctrine of pre-established harmony.
2803. The sense of touch is also the sense of form. The fingers constitute so perfect an organ, that it is scarcely requisite for us to estimate all its value. In it the whole body is repeated simply in forms, in spiritual motions.
2804. Through the hand the whole planet becomes an object unto us. It is the hand which instructs us how to know this terrestrial world.
2805. The greatest perfection is attained by the greatest variety or multiplied diversity of the organs. The limbs are simply destined for motion, yet feeling also resides in them, because they are invested with integument. Could the limbs therefore divide into motor and tactile members, the conceivable sum total of perfection must be attained. This division is present alone in Man. The feet have become simply locomotive members, because they are those of the sex; but the hands have become tactile members, because they are those of the encephalic animal.
2806. It is not, as has been imagined, the hands, as hands, which confer nobility upon our species, for by their means an essential half of the animality, or the power of locomotion, is lost; but it is the preservation of all possible functions in an animal, and in such a manner that each stands upon its highest grade of perfection. The highest perfection cannot, however, be attained where two functions are attached to one organ. Should both hands and feet exercise the sense of touch, the motion is impaired; should both move the body, then the sense suffers.
2807. The four hands of the Apes are therefore an imperfection, which we have no need to envy them. They can, properly speaking, only climb and grasp with them, but not run. Each gait of theirs becomes therefore ungainly, the horizontal as well as the upright or perpendicular, and they make use of both alternately; because too the act of handling, i. e. of climbing, is their only proper movement. Now, in the act of climbing all the limbs are brought into requisition, and consequently a free voluntary touch and a free progression disappear.
2808. The feet support the body and stand in its service.
2809. The hands are, on the contrary, supported by the body, are free.
2810. The wings also support the body.
2811. Feet and hands define man. Through the two only does he become _free_.
_Tegumental Covering._
2812. The skin, as being originally a branchia, is provided also with its branchial operculum. This is the epidermis.
2813. Scales are plications or folds of the epidermis, which accord with the situation of the branchial vessels; they are therefore arranged generally in a circular form around the body, and are more definitely varied upon the dorsal than ventral aspect of the body. Large scales, or those which may be regarded as formed by the confluence of several scales, are called scutes.
2814. It is merely the epidermis which is concerned in the production of the squamous covering; but if the former branchial vessels emerge themselves so as to project above the integument and become dried, then hairs originate.
2815. If these hairs ramify, then there are feathers.
2816. It is only therefore the Mammalia and Birds who possess a proper external covering or garment.
2817. The claws or nails are scales upon the extremity of the branchial arches, which have become animal in character, or in other words, are digital scales; they are animal branchial opercula.
2818. The nails are demi-claws, and therefore leave the points of the digits free. Free digital apices or points constitute the most perfect organ of touch, because this is then divided into two parts and because the nail increases the amount of resistance.
2819. The organs of defence are therefore an appendage of the sense of feeling, as are the bones of the motor system.
_Splanchnic or Visceral Senses._
2820. These senses will not be found to disown their predecessors; and just as the latter extracted the qualities from terrestrial matter, so also do they. But while the former had to deal with the material, the present set will have to transact business with the spiritual qualities.
2. _Function of the Gustatory Sense._
2821. Digestion is a chemical process, and one indeed wherein actual mixture and decomposition take place; it is therefore also, and "par excellence," an aqueous process. For matters that are actually decomposible are alone submitted to digestion, since it is of too coarse a character to perceive the proneness of such bodies to decomposition.
2822. It belongs only to a higher grade of perfection, or to a nervose condition of the digestive process, to perceive the rationale of the decomposition, or the spiritual conflict, which prevails between the matters, when they are about to separate.
2823. Now, the organ which only perceives the qualities of matters, without reference to their actual separation, is a sense. Upon the highest grade of perfection the digestive passes over into a sensorial function.
2824. _Tasting_ is the first commencement of the digestion in the nervous system, where the aliments are felt just before the analysis into their polar quantities has taken place. The gustatory is a water-sense.
2825. For the exercise of the sense of taste the same conditions are requisite as for digestion, viz. solution and capacity for decomposition. Without the capacity for being dissolved, and without the occurrence of actual solution, nothing can be tasted, any more than digested. The saliva is the gastric juice for the tongue.
2826. If water be the basic element in the process of digestion, so in gustation must the higher water, or the _salt_, be the basis of taste. Salt alone is sapid, and every thing in order to be tasted, must possess saline properties.
2827. The tongue, by means of the saliva, passes over gradually into salt. Salt is the last extremity of the tongue. The salt-formation is a member of the gustatory formation. Hence tasting is only an ascension of the inorganic to the animal tongue. The salt is the gustatory sense of the earth.
2828. The general object of taste is the salt of the sea. It alone can and must be converted into a pleasing taste or relish. What is general in nature, is the antetype of that which is equivalent in an organism. Sea-salt and the tongue or saliva are one in kind.
2829. Every thing admits only of being tasted, in so far as it is salt; every thing is but savoury, in so far as it is marine salt.
2830. As the component parts of sea-salt are acid and alkali, so also are both these the extremes of tastes. Tastes are divisible accordingly into acid and alkaline savours.
2831. As salt is a product of what is inorganic, so will the sapid bodies of this kind be pleasant objects for gustation, provided they do not act chemically nor in excess. Therefore, what is saline, or acid and alkali combined, is agreeable to the taste, even if its action should last a long time.
2832. On the contrary, the proper organic savours, which admit with difficulty of being reduced to the former or inorganic kinds, if they do not prove at once nauseous or unpleasant, yet become so by lengthened operation, e. g. the Sweet, the Bitter.
2833. According to these savours has the organ of taste been regulated. It hath in itself also polar relations. The point or tip of the tongue tastes what is acid, its root more readily what is bitter, its dorsum or back what is poignant or moist.
2834. The matters are not mechanically analyzed upon the tongue; it does not therefore taste the several component parts, but only their chemical behaviour or relation in the water, in other words, their reaction.
3. _Function of the Olfactory Sense._
2835. In the lungs the air is materially analyzed and deprived of its oxygen; but, when these organs have assumed a nervose character, the tension only of the air unto the analysis will be perceived. Now, the action of the air is the electrism. The nose perceives only the electrical condition of the air.
2836. The sensation of the electrical relations is called _Smelling_. The olfactory sense is an air-sense. We smell nothing but the electricity, and neither the contact, nor the impressions, &c. of the particles that find their way into the nose. These particles would have no effect upon the nasal organ, if they did not stand in an electrical relation unto it.
2837. Now, the electrical bodies in nature are the resins or Inflammables. What salt is for the gustatory, that is resin for the olfactory sense. The nose is an electrical, a resinous organ.
2838. In like manner the solubility of bodies in the air is as requisite for smell, as that in water is for taste. The water is the menstruum of the sapid, as the air is of odorous, bodies; and this indeed of necessity, because water and air are the antetypes of these mineral classes.
2839. In order to be an odorous body or object of smell, the resin must resolve itself into air, or become aeriform. Aeriform resin is æthereal oil. Bodies such as those which part rapidly with their electricity, substances containing hydrogen, æthereal oils and burnt spirits, are the usual objects appreciated by the olfactory sense.
2840. The hydrogenous body is therefore provided with a sweet scent. Most bodies which are evolved by fermentation, in so far as they are electrical, are odoriferous. Most blossoms smell agreeably, because they secrete aerial matters.
2841. The products of putrefaction emit a fetid odour, because they indicate the presence not of aerial, but aqueous and terrestrial matters. Nearly all animal bodies stink, besides many secretions of the sexual parts, because they belong to the vegetable nature.
2842. The objects of taste have their residence in what is inorganic, but those of smell, as being objects of a higher sense, have it in the vegetable kingdom. The succeeding or auditory sense has the animal kingdom for its object, while the eye scans the universe.
2843. The nose is in every respect an electrical organ; it is an electrophore, or rather a battery consisting of numerous plates. Of the truth of this, its numerous tortuous passages and laminated bones are striking proofs.
2844. That the nose consists of a great number of blood-vessels, as well as of arteriose olfactory nerves, is quite commensurate with its character or signification, as being a higher pulmonary organ.
2845. The objects of the three vegetative senses are the three elements of the planet, earth, water and air; in the first of these reside the relation of the gravity, rest, and crystallization; in the last the relation of the electricity; in the water that of the chemism. The sense of feeling is an earth-sense, that of taste a salt-sense, that of smell a resin-sense.
b. _Functions of the Animal Senses._
2846. The objects of the animal senses are no longer matter, nor its chemical qualities, but the higher relations of the solar system, and the highest organizations, the animals themselves. Throughout the supra-planetary solar system there is naught to be conceived but motion and light in action. Wherever there is æther, it is in motion; the corresponding organs of sense must therefore perceive these two relations. Now, since the animal is also motion and light, and this alone, so by means of these senses will the innermost of animality be at once perceived. Animals become acquainted with themselves only through these senses, and by them only enter, in so far as they are animals, into communication with each other. In so far as they are mere masses of matter, they are capable of self-perception through other senses. These senses may therefore also be called cosmical, while the three former are terrestrial.
2847. They correspond thus with each other. The tactile sense is a precursor of the sense of motion, and represents the motion, gravity, and pressure, after a terrestrial manner; the two splanchnic senses are the precursors of the light-sense, since they dwell upon the qualities of the matter, while the light also is only a quality of the æther. The sense of smell, being as it were an air-sense, will in particular border most closely upon the light-sense.
2848. Through the two cosmic senses the universe is translated into the animal, like the planet is through the terrestrial senses; through the former also the animal spirit, which is a transcript of the universal spirit, passes over into other animals. They are the senses of the highest instruction, of freedom.
4. _Function of the Auditory Sense._
In the æther resides the movement of the world.
2849. To the motor system that only which is its equal, and thus the movement of nature, can of necessity become an object. The motor system represented as a sense, cannot, however, perceive the borrowed or derived motion, not the planetary or massive motion, but the primary motion of the æther. The planetary motion is related to the primary motion as the oxydation is to the electrism, as chemical analysis to chemical affinity, and consequently also as respiration to smelling, as digestion to tasting, in short, as the material metatype to the spiritual antetype.
2850. The limbs are the planetary motion organized, and therefore perceive only this material motion--pressure. Touch is related to the animal sense of motion, as digestion is to the tasting.
2851. Smell and taste no longer perceive the bodies in the very act of decomposition, but their laws or their spiritual operations; so will the motor sense not perceive, like the sense of touch, the mass when in motion, but only the _motor laws of the mass_.
2852. These laws of motion are those of the primary motion. This is, however, a product of the light in the æther, an effect of polarity, and that indeed the first polarity which was manifested in the universe. The motor sense therefore perceives only a motion which has originated through primary polarity.
2853. Such motion is not relative in kind, i. e. it does not affect several portions of the matter in reference to some other matter; but it affects the whole matter internally, or its atoms, so that all matter may remain in its place and yet every atom of it be moved.
2854. This motion is like the motion of heat in the matter. By it heat is excited. For internal motion of the atoms, when aroused by polarity, so that every atom enters into a state of motion against the other, is a discharging of the poles, and consequently development of heat.
2855. This internal motion has, however, been produced by an external; for the external motion acts by contact, and this is a process of polarization. Now, the interior of a mass is only moved by repeated contact, or through the restlessness of polarization and by a proper amount of force, or one which is proportional to the mechanical resistance of the mass to be excited. The last of these is the stroke or blow, the first the vibration of the body. By vibration or oscillation only can a body be internally polarized; for, if it does not oscillate upon the shock being applied, it is still indeed set in motion, but "en masse," so that the internal parts remain in a state of rest.
2856. Oscillation is distinguished from continuous or progressive motion by its affecting the _atoms_ of the body, while the latter acts upon the body itself. Through the vibration heat is engendered, because the poles are free and the matter passes over into æther.
2857. Vibration must endure the longest in solid bodies, and thus in that which belongs to the earth. Among these the hard bodies must take precedence, because the soft are of an aqueous nature. Among the hard bodies again the heaviest must vibrate most effectually, because they offer a longer resistance, and do not yield so soon as light bodies to the effort made at separation. The purest representative of the earth-element or the _metal_ is thus the best instrument or means of vibration, and consequently the object of the motor sense.
2858. Thus as the salt of the earth-element is the object of taste, and as the resin of the earth-element is the object of smell, so would the metal be the object of this motor sense.
2859. But no sense-object is without its medium for transmission, except in the case of the sense of feeling or touch. The salt is only tasted by means of the water, the Inflammable only smelt by means of the air; the metal's primary motion could not therefore be perceived directly by the auditory sense. It must be propagated through the medium which ranks next to the heat, and whose atoms insinuate themselves most easily into those of the vibrating body--thus through the air. Man perceives the primary motion, in which things tend to resolve themselves into æther, through the air. By the metal, or by every vibrating body, the vibration of the air is communicated.
2860. This vibration is not, however, a general movement to and fro, but a dissolution of the material bands. This dissolution can only take place according to the laws of the primary motion. They are rigidified in the solid masses as crystalline forms. Every law of motion is a crystalline form which has become free or spiritually manifested. Through the vibration forms are engendered in bodies, which are commensurate with the substance and form of the mass and the degree of vibration. These forms, being as it were the ghosts or phantoms of crystals, are called _sonorous figures_.
2861. If the air be displaced when in a state of covibration, it is not thrown into undulatory circles or waves, like water into which a stone has been cast, but in each of its parts the sonorous figure of the rigid body is repeatedly represented. The vibration of air is a progressive motion of sonorous figures.
2862. If the sonorous figures are not incommensurable, several may be at one and the same time in a _single_ portion of air, without interfering with each other. They harmonize, because they have originated according to concordant laws. But if they are products of different laws, they are then confused, and an indeterminate _offensive_ vibration originates, just as savours become loathsome if they depart from their laws.
2863. These figures of the air are only perceived by the _Ear_. The ear is the only sense in which the motor system is represented in a _pure_ state, devoid of any vegetal signification, and simply endowed with nervose nobility. The ear is therefore the only organ which can perceive the primary motion of the matter; for like acts only in or upon its like.
2864. The metals are the ear of nature, the salt her tongue, the resin her nose, the earth her hand.
2865. The power or capacity excited by sonorous figures of covibrating according to the same laws, constitutes _Hearing_; the phenomenon is called _Sound_. Hearing is a primary motion in the musculo-osseous system of the ear, which is communicated to the auditory nerve. The auditory sense is æther-sense, metal-sense. Hearing is magnetizing.
2866. The sonorous figures are formed in the auditory organ, and even in the auditory nerves, just as they have been represented upon an infinitely small scale in the air. The nerve becomes in hearing a sonorous figure.
2867. It is not the mere motion in the auditory organs which produces the sensation of sound; the nerve certainly perceives each movement in the ears, because none is possible apart from or without primary motion; only such a motion is no sound, but only a noise. What has been written in the tingling metal according to eternal laws, is transcribed or copied in the auditory nerve; it is only this writing, but no massive motion of the air, which is legible by the nerve.
2868. Melody is a retrogression of the matter into æther, of the formed world into the primary world; through melody is the spirit of the world revealed. The ear is the first liberation of the animal from all terrestrial matter; through the ear the animal becomes for the first time spiritual.
2869. Melody is the voice of the universe, whereby it proclaims its scheme, or its innermost essence. Hence the wondrous, mysterious action of harmony, the secret sovereignty of music. Music is the expression of the ardent desire to revert to the primary idea. It makes man unconsciously yearn after a condition which he knoweth not; it transports him unconsciously into this condition of divine repose and godly bliss.
_Speech._
2870. That which melodizes proclaims its spirit: the melody of animals displays their internal law.
2871. _The musical system of all animal laws is Speech._
2872. Speech is the representation of all nature's sonorous figures in the human organ of sound.
2873. Through speech Man delineates himself in spiritual outlines or sketches, which, devoid of matter (or body), he sets down before himself. Such sketches are easily seen through, since to them every material covering is wanting, and they lie purely before the sensation, as the law, the will of nature.
2874. Through speech Man appears as a double essence. He is a corporeal essence; and the spoken word appears before him in the same outlines, but without body. When conversing, Man is a _self-manifestation unto self_.
2875. Previous to speech no self-consciousness originates.
2876. Without an auditory organ there is no self-consciousness.
2877. To the organ of hearing, however, belongs also the auditory nerve and the cerebellum. Without a cerebellum there is no self-consciousness.
2878. While Man appears unto himself, he appears also to change. Nature is gloomy, incomprehensible; the spirit is clear, and enlightens her.
2879. Manifestation is only possible through self-manifestation; through a doubling of itself, through its expression.
2880. Animals appear only, in so far as they are individual self-manifestations of Man.
2881. With speech Man creates unto himself his world. Without speech there is no world. For the Apes there is no world, but only tree-fruits, female and male.
2882. Through speech Man becomes acquainted with or learns to know himself; through it he becomes a self-substantial essence, which resembles God, because it creates for itself its world, and recognizes itself, i. e. speaks.
2883. Words are forms of our body mathematically laid down.
2884. A single world is dead, so also are many.
2885. Words, which are connected together according to organic laws, form an organic system, are at once alive, and have a meaning.
2886. Speech originates gradually like the organs, like Man. Speech grows like a plant; at first it is only root, next it puts forth a stem, then leaves, and finally blossoms, when it is the perfect expression of the animal body.
2887. The organ of speech is composed of the three terrestrial organs of sense, the air-, water-, and earth-sense.
2888. The air-organs are the principal medium, because they must produce the sonorous figures; the tongue imparts to them the specific modification; but the lips and jaws, as being motor members, afford the articulation or the _movement_ proper. The lungs and nose breathe out the tones; the tongue digests them; the lips move them, and fashion them into perfect bodies--words.
2889. A word is at once for itself a regularly inter-articulated body. The sounds are its members, its organs or fundamental formations.
2890. Speaking is a gentle respiration, carried on by the mouth, nose, and limbs or jaws.
2891. As respiration has a special thorax, so also has speaking. The speech-(or voice-) thorax is the larynx.
2892. The larynx represents the ribs and arms, which all move in order to form a sound. The tongue is, so to speak, the head upon this thorax.
2893. The nose imparts euphony to the sounds. It tests their fragrancy. The tongue gives them a special quality, their chemical character or taste; the teeth and lips furnish the cadence as a kind of joint to the sounds, or in other terms the words.
2894. Four organs of sense belong to speech, viz:
Touch in the Jaws. Taste in the Tongue. Smell in the Nose. Hearing in the Ear.
2895. The ear receives the products of the three vegetative organs of sense. It is a synthetic sense.
2896. The tongue gives the vowels; the jaws the consonants.
2897. In accordance with this the vowels are the _body_ of speech, and the consonants the _limbs_ or members, whereby it effects its movements.
2898. Vowels express time, consonants space; the one the chemical import, the other the form.
2899. The vowel E expresses the present, A that which has just past, O that which has quite passed, U that which has passed long ago, I the future.
2900. The more consonants there are in the words, so much the richer is the language; the more vowels, the poorer it is. Such is the speech of savages or wild men.
2901. The speech of animals is a vocal or vowel-speech.
5. _Function of the Optic Sense._
2902. As the primary motion of the world is manifested to the animal through the ear; so does the primitive cause of the motion, that of every activity and every phenomenon, i. e. light, appear unto the nerve-sense.
2903. The light-sense is similarly formed or modelled according to the light of nature, and kindles also the light within itself, just as the light originated in the æther; viz. through primary antagonism taking place in its own substance.
2904. Light is the binary division of the æther-mass, not an antagonism between it and some other matter; in like manner sight is a binary division of the nerve-mass in itself, without antagonism to other organs.
2905. Sight is the tension of the æther directly continued into the animal æther, just as taste was a chemical action continued into the animal chemism, and smell an electrical process continued into the animal electrism.
2906. In sight the nervous mass is completely self-antagonized, is a phenomenon unto itself; the eye is the brain placed opposite to the brain.
2907. Sight is thus a tension between optic and central brain; as illumination is tension between planetary and solar æther.
2908. Illumination and vision are of one kind, only occurring in two different sorts of worlds. The planet sees by means of the illumination, the animal illuminates or gives out light by means of seeing. Sight is a light-sense.
2909. Now, the illumination is a fixation of æther, a coloration, and thus a descent of the æther into what is terrestrial. In seeing we perceive the æther, as to how it becomes world; in hearing we perceived the world, as to how it became æther.
2910. Seeing and hearing are opposite functions; the first indicates the creation, the latter the return of the creation into chaos.
2911. Through sight we become acquainted with the universe, through hearing with the miniature universe, or man. Sight passes out of us, hearing into us; or through sight Man is posited in the world, through hearing among his fellow-men. Sight is the speech of the world, hearing that of the planet.
2912. Sight is the speech or language of the universe, hearing the language of Man. Through sight the world reveals unto us its spirit or its thoughts; but through hearing man only discloses what are his own. As words are but the representations of the disintegrated body of Man, so are the world-forms the representatives of the disintegrated body of the primary spirit. The word is a rigidified, crystallized thought of Man; a natural body is a rigidified, crystalline thought of the primary act--a word of God.
2913. Through hearing self-consciousness originates; through sight consciousness of the world, universal consciousness. By means of the former we only become acquainted with human relations--which is _understanding_; by the latter with those that are universal--this being _reason_.
2914. Without the ear there would be no understanding, without the eye no reason.
2915. Understanding is microcosm, reason macrocosm. From what is intelligent we expect human wisdom, from what is rational world-wisdom.
2916. That the light also has a medium, whereby it acts upon us, is at once evident from our existing in such a medium; but it could also act directly upon us, were it not inevitably broken up beforehand into colours in traversing the media. All terrestrial elements may be a medium for the transmission of light, such as the gaseous, fluid, and solid, i. e. if they are transparent.
2917. We perceive only coloured light, because our organ of light is only a rigidified colour, a material light. There is no pure light for us; there is none also in general.
2918. Sight is thus a terrestrial light-tension, a colour-becoming.
2919. This happens only through refraction. The eye is a refracting medium. It is distinguished from the brain by being a translucent, refractive, encephalic substance.
2920. Light does not stream into the eye like water into the sponge, but it progresses gradually into and operates upon it.
2921. The eye, in order to experience the sensation of light, is placed in a similar tension to the air, water, or crystal. This tension between it and the brain is perceived by the latter as illumination. The eye is a prism, in which the brain sees the world, in which the brain observes its own tension, or the production of colour. Sight is a deoxydation of the eye.
2922. The optic nerve is an organized ray of light, the brain an organized sun, the eye an organized chromatic sun or rainbow.
2923. Just as the sonorous figures are delineated in the ears, and as the nerve perceives these, but not any concussion of the air; so also does the optic nerve perceive not the light in general, but its terrestrial formation or the chromatic image, which has been propagated into the eye.
2924. In an eye, while seeing, the world is depicted; as in the ear, when hearing, the crystalline forms of the air are delineated.
2925. The eye does not on that account see two worlds. For the chromatic image is nothing else than that which is external to or without the eye. It is verily one and the same influence of light, which acts continuously in a straight line between the chromatic image and the object apparent or beheld.
2926. As a stick thrusts us from the side whither it comes; so does the chromatic image from the side whence the light comes. The exit and entrance of the light are not distinct from each other. The objects could not therefore appear reversed, because we do not _see_ the image in the eye, but _feel_ its process of deoxydation along with its direction.
2927. The objects of the eye are colours. Like as they are related in nature, so also must they be in sight; for they are only the eye extended, or this again is only the formed colour.
2928. We see nothing else but colours, no bodies. For the eye there is no material world. It directly perceives the spirit, and indeed its own spirit, or the world of light.
2929. There is no pre-established harmony, but complete conformity between the world and organ of sense. (Vid. Oken's Essays, "Ueber das Universum als fort-gesetztes System der Sinne," and "Erste Ideen zur Theorie des Lichts." Jena bei Fromann, 1808.)
II. _Functions of the Sexual Animal._
A. VEGETAL SEXUAL ORGANS.
1. _Of the Sexual Intestine._
2930. As the sexual animal is in every respect the encephalic animal reversed, so also is this the case in respect to its functions. The sexual intestine gives out through its mouth or anus, while the other or animal gut takes in. As being the intestine of the vegetal animal it receives the fermented product of digestion, or the excrement, and conveys it backwards towards the sexual orifice or mouth.
2931. The intestinal function of the sexual animal is an act of vomition. The evacuation is an act of vomition, seeing that the intestinal contents are moved backwards.
2932. The sexual stomach is the rectum. In it the excrementitious matter is accumulated, so that it may be ready for being broken away in the process of evacuation.
2933. The commencement of the sexual intestine is the cæcum, its extremity the anus.
2. _Functions of the Sexual Lung._
2934. We may distinguish two circulations, the splanchnic or visceral, which takes place between the lung, intestine, and liver, and the great circulation, which instead of going to the viscera passes to the other organs, and is called the bodily or systemic circulation. The liver excretes the product which results from the splanchnic circulation; the systemic circulation has its organ also, but one which does not secrete a special, but general product.
2935. The general secernent organ of the whole body together with all its systems is the sexual system, which having been elevated by virtue of this general character to the rank even of an animal, is a true sexual animal. That which is a general and not a partial excretion, is imparted by the sexual animal. It is the animal reversed.
2936. Thus the secernent organ of the general circulation must belong to the sexual system, and perform in it that which the liver, or in other words the reverse of the lungs, has done in the splanchnic circulation. The _kidneys_ are the lungs reversed.
2937. If the bile be an extract from the visceral blood, so is the urine an extract from the body's blood, and is consequently the purest reflex or pattern of the former.
2938. The urine is sexual blood, just as the excrement is a product of the sexual digestion. The urine is reversed blood.
2939. The formation of urine is a retro-formation of the blood into digestive fluid or sap. The urine is blood of the sexual animal which has become chyle. It has both properties in itself. It is discoloured blood, and consists for the greatest part of water and salts, all of which are characters belonging unto chyle. It, however, contains urea, which corresponds to the noblest parts of the blood. This substance, like fibrine, consists for the greatest part of nitrogen; it may be called dissolved or decomposed fibrine. It imparts colour to the urine; it is converted by oxydation into lithic or uric acid, and is precipitated of a red colour analogous to that of the blood-globules. In addition to this, albumen, gelatine, carbonate of lime, and phosphorus, consequently the whole blood, are present in the urine.
2940. In urea the muscle flows or runs out of the animal, in albumen the nerve, in lime and phosphorus the bone, in gelatine the tegumentary together with the visceral system, lastly, in water the menstruum of the digestion and respiration.
2941. Thus the urine, just like the blood, is the whole body rendered fluid, but only in a sexual manner, namely, as being half decomposed.
2942. So the bile, from its not representing the whole body, does not contain the latter in itself. It properly contains only the excretion of the intestinal process.
2943. The kidneys stand accordingly opposed without distinction to all the organs, in so far as all of them are affected by the circulation. Their remote sympathy, or if we please, their antagonism, is with the animal systems, or with bone, muscle, and nerve. With the osseous, as being the profoundest system, there is of necessity also a close sympathy. In diseases of bone the bones, as well as the _morbid matter_, flow away principally through the urine. Their most intimate sympathy must be with the organs of circulation, with the lung, liver, intestine, and skin. As the skin is also an organ of evaporation, so is the antagonism between it and the kidneys of a direct or immediate kind. The skin is the kidneys expanded into a large cyst. These are in turn, just as the lung is, the inverted skin.
2944. A lung in the reversed animal can do nothing else but expire. It only expels the evaporated matter of the sanguinary system, but takes in none, so as to alter or support the blood. The sexual animal aims at the destruction of the animal. The urinary cyst, as being the remnant of the allantois, and of the primordial kidneys or sexual branchiæ, is simply destined to purposes of expulsion. It is the larynx reversed. Micturition takes place through contraction of the cyst, as does expiration by that of the lungs in Reptilia. It is a cough.
B. FUNCTIONS OF THE ANIMAL SEXUAL ORGANS.
2945. The sexual functions proper correspond to the sensorial functions, though to these upon an inferior stage. They are sensorial functions, which are simply occupied with the materials of the senses, so that they are vegetative senses. They are a prefiguration of the sense of feeling, taste, and smell.
1. _Functions of the Male Organs._
2946. The testes secrete semen in the same manner that the salivary glands do their fluid or juice.
2947. The semen is sexual saliva, and is thus sexual virus or poison. Like the saliva destroys that which is living, so does the semen. The saliva, however, destroys it in order to form a new animal out of the food; with the same motive the semen destroys it. But both differ in this, that the saliva takes care of the body to which it belongs, while the semen attends to another body, the foetus or fruit.
2948. The saliva is only the highest condition of the digestive fluid, and is thus a totality merely of the intestinal system; the urine is the total product of the vascular system in its antagonism with the lung; but the semen is the product of the whole body. Through the semen the whole body, rendered fluid or reduced to the primary form, runs away. The semen is the chyle already prepared for all parts; but because it is in a sexual animal, it thus takes the reverse direction and passes out.
2949. A fluid, in which the whole animal has been dissolved, is parallel to the nerve-or point-mass. The semen is a fluid point-or nerve-mass, the fluid brain.
2950. Even what is spiritual directly resides in the semen; it need only assume a form and the cerebral functions commence.
2951. The penis, as being the sexual tongue, has only retained the sensibility of the sense of touch and the function of ingestion.
2. _Functions of the Female Parts._
2952. The female aperture is the pharynx for the ingestion.
2953. It is by means of the female parts that the whole sexual system becomes for the first time equal to the perfected animal; through them for the first time the male tongue obtains an oral cavity.
2954. In the total representation of the sexual animal the female parts environ and include the male. This moment is called _copulation_.
2955. Copulation is the representation of the entire animal out of two incomplete ones. The sexual animal is only an entire beast in copulation, and is only then to be considered as equal to the encephalic animal. Copulation is a representation of the hermaphrodite.
2956. This propensity for bringing about the sexual animal's completion is sexual passion or lust.
2957. In copulation the male parts are "par excellence" the sensorial organ, the female only the recipient mouth. Both are properly organs of sense, but the one is operative or active, the other patient or passive.
2958. Previous to copulation the female parts are consequently inactive, just as digestion is before taste. As digestion first commences after taste has given up the food and excited the stomach to activity, so also the sexual function first commences in female animals, after the act of tasting is past.
2959. Through copulation what is female becomes masculine. It now secretes for the first time self-substantial semen. Through impregnation the female ovaria are first excited to secrete the saliva, which the whole animal contains in a state of solution.
2960. As the chyle becomes what it is, or is derived from the saliva and food; so from the semen and vitellus proceeds the foetus, but in such a manner that the female substance gives it the mass, while upon this the male only bestows the polarity.
2961. Thus, were the male semen to actually solidify into the foetus; it is still not its mass which comes into consideration in the latter, but only its polarizing strength. It supplies the place of the nervous system. This power appears to reside principally in the (seminal) Infusoria or animalcules, just as that of the blood does in the blood-globules. Meanwhile both are only signs of the maturity of their respective juices, as the Infusoria in sea-water are a proof also that the sea can produce other animals from its mucus or slime. The Infusoria are the primary mass of the Organic. Its life is only a manifestation of the seminal polarity. The Infusoria are semen which has been poured out over the earth. Propagation is only possible through reduction taking place to the infusorial primary mass.
2962. The semen and the ovum first meet or come together in the uterus.
2963. The ovum is the mean between vegetable and animal semen. As the former is distinctly formed and at once represents, upon a small scale within itself, the principal parts of the future plant, so does the ovum; but only in parts, from which the animal organs first of all grow forth, upon which having commenced, the former or vegetal parts are cast aside.
2964. The ovum is the entire animal in idea, or in design, but not yet in structure; it is the thought unto the animal; it is related to the animal as the thought is to the word.
2965. The ovum has therefore no organ of the animal preformed within itself, but only the materials requisite thereunto. But the materials are not so general in character, that like as from the infusorial mass, everything could become or derive its existence from everything else. But they are at once destined for definite organs, as the vitellus for the intestine, the albumen probably for the integument.
2966. In an ovum therefore the animal resides preformed only in a spectral or phantom-like manner. There are principal masses present in it, from which the principal organs originate.
_Mammæ._
2967. In the oviparous animals the secretion of the vitellus is distinct from that of the albumen; the one takes place in the ovarium, the other in the oviduct or uterus.
2968. By degrees the albumen-secreting vessels advance further outwards upon the orifice of the sexual parts, and are then called milk-organs--_Mammæ_.
2969. Mammæ are only the vascular bundles of the oviduct placed in the direction outwards, or albumen-glands of the integument.
2970. Mammalia are those animals where the ovarium has completely separated into albumen-and vitellus-organ.
2971. Those mammæ which have scarcely been detached from the oviduct, and become free, are necessarily more incomplete, and are situated in the neighbourhood of the sexual parts--as udders.
2972. As the separation of the substances composing the ovum is an advancing-step towards their improvement, so also is the removal of the albumen-glands from the vitelline sac a nobler condition. They cannot, however, remove further off than to the thorax, because this is the highest post or station of the vegetative parts.
2973. Milk is a vegetable product of the animal.
2974. Numerous mammæ are a lower development.
2975. The milk is albumen, which has been secreted by tegumentary glands; it is animal albumen. The lacteal or milk-organs therefore belong to the sexual system.
2976. As the male parts are only the female otherwise developed, it is to be comprehended why the male animals also have mammæ. They are probably the embryo's principal organ of absorption.
_Functions of the Uterus._
2977. Now, the uterus obtains the sexual food in a living state, and from its being such is affected by it.
2978. The uterus must be thus a world for the living germ. Two processes are, however, indispensable for the germ, namely _nutrition_ and _respiration_; both of which are furnished by the uterus.
2979. The uterus is to be regarded as the water, the sea in which the germ or embryo is developed. The water decomposes into basic nutritive matter and oxygenous matter of respiration. The water of the uterus is the blood; this is separated through the antagonism of the foetus into mucus and oxygen. The mucus penetrates into the amnion, the oxygen into the chorion, or the placenta.
2980. The fundus of the uterus is more arteriose than the os uteri, and stands therefore in opposition with it.
3. _Development of the Foetus._
a. _Anatomy._
2981. The germ may be regarded as a vesicle, full of nutritive matter or albumen, situated within the cavity of the uterus, the walls of which act upon it.
2982. As the fundus of the uterus is the arterial pole, so it oxydizes the vesicle and repels that part which lies close upon it. Through this originates a saccular inversion, as in the mesentery of the peritoneum, and the vesicle separates into three divisions. In itself it is amnion, the inverted part is the integument of the embryo, the tube which unites these the umbilical cord.
2983. The amnion is thus the root or primary cyst of the integument.
2984. Through continuous oxydation blood-vessels are developed upon the surface of the amnion, which finally withdraw to constitute a special integument, which is called chorion. Its vessels are in like manner repelled from the fundus uteri, and prolonged into the inversion of the umbilical cord and the embryo. The chorion is thus the root or primary cyst of the vascular system.
2985. These two bladders, sacs, or cysts are the only general ones which circumscribe or invest the entire embryo, because there are only two general vegetative systems, namely, the tegumentary and vascular.
2986. The embryo has not originated freely in these shut sacs, but only through their introversion; it is itself a portion of these sacs.
2987. The embryo properly lies external to its envelopes, as the intestine does in respect to the mesentery.
2988. Just as the two general vegetative systems have been developed from primary cysts, so also are there sacs for the two special vegetative systems, or the intestinal and sexual; but, on that very account, these cannot be general sacs, nor any longer envelop the embryo.
2989. At the entrance of the inversion of the umbilical cord is situated a small vesicle, which divides and is prolonged into the two intestines. It is therefore the root or primary sac of the intestinal system, and is called in Man the 'vesicula umbilicalis,' in the Mammalia the 'tunica erythroides,' and in oviparous animals the 'vitellus.'
2990. In the same situation is placed another cyst, which is prolonged into what has been called the 'urachus' and the urinary cyst, and from which the primordial kidneys, the true kidneys, and the sexual parts are developed by sacciform eversion. This sac is called the 'allantois,' 'tunica allantoides,' and is consequently the root or primary sac of the sexual system.
2991. These cysts or sacs are consequently not envelopes that serve for the protection of the foetus, but its developmental organs, which disappear, so soon as their prolongations into the foetus itself enable them to exercise their functions.
2992. There are thus as many developmental sacs or cysts, as there are vegetative systems present, viz.:
a. Two General Cysts. 1. The Vascular cyst--Chorion. 2. Tegumental cyst--Amnion.
b. Two Special Cysts. 3. The Intestinal cyst--Vitellus. 4. Sexual cyst--Allantois.
2993. It is only the vegetative systems which take root in the foetal envelopes, but not the animal systems. There is no developmental cyst for the nervous, muscular, and osseous systems.
2994. The persistent vegetative systems are the developmental organs for the animal systems; as the intestine for the bones, the veins for the muscles, the tegument or branchial sac for the nerves.
2995. The foetus consists of three floors or stories like a house, whereof one has been based upon, or rather developed out of the other, viz.:
a. Of the Developmental cysts. b. " Vegetative systems. c. " Animal systems.
2996. According to time the sacs are developed in the following series. The first sac is that of the vitellus or the intestine, which is also the first that is present in the development of the animal kingdom. Upon this vitelline membrane the blood-vessels ('vasa omphalomesenterica') are developed, are prolonged into the body with the intestine, turn again outwards and form the chorion. From this the amnion next separates into the envelopes, and the integument upon the embryo. Lastly, the allantois appears, and in its prolongation the sexual parts.
2997. Originally the whole chorion is replete all around with vessels; but as the process of oxydation occurs most powerfully upon the fundus uteri, so are the vessels developed most abundantly in that very situation, and form the placenta.
2998. The placenta is no peculiar organ, but only the more energetic part of the chorion.
2999. It must necessarily be placed around the insertion of the umbilical cord, because at this spot the inversion takes place, on account of the strong oxydation.
3000. The placenta is always situated upon the fundus uteri, because it originates only through its influence. It cannot therefore begin to suck, like the mouth of a leech, in a fortuitous or voluntary manner anywhere. If it is found occupying some other situation, it is a proof that the oxydizing process of the uterus has been displaced. This is consequently an abnormal situation.
3001. Opposite to the foetal vascular system is first of all developed the general system of animal life, namely, the nervous system, and indeed the spinal chord, or what has been called the 'carina' or primary streak.
3002. The development of all other systems oscillates in this antagonism of blood and nerves.
3003. In the antagonism of the placenta is formed the liver, which in the embryo is one of the largest organs, and in its antagonism the brain is developed.
3004. In the antagonism of the amnion the integument is evolved, and in its antagonism the branchiæ and lungs.
3005. In obedience to the antagonism of the arteries and veins, the vitelline sac divides into small and large intestine. The one pursues a course towards the arterial extremity of the body, the brain, the other towards the venous or sexual parts; mouth and anus.
3006. Finally, from the antagonism of the allantois the sexual parts emerge at the very extremity of the body, which is placed opposite to the mouth.
3007. The osseous and muscular system first makes its appearance, when the vegetative parts are present.
3008. The intestines, the vessels, and the commencement of the sexual parts are originally situated in the umbilical cord, which is itself surrounded by the integument. The umbilical cord is therefore nothing else than the posterior extremity of the body or abdomen, through which the embryo respires and is nourished.
3009. The first respiration and deglutition is therefore a respiration and deglutition effected by the sexual parts, as in the lowest animals.
b. _Functions of the Foetus._
1. _Nutrition._
3010. The juices which are contained in the developmental sacs are nutritive juices or chyle; they contain principally albumen. The chyle of the intestinal vesicle depends upon the vitellus.
3011. The foetal water in the amnion is derived from albumen in the ovum; it is here secreted by the oviduct, but in the Mammalia by the internal wall of the uterus, and absorbed by the general envelopes.
3012. The cause of this secretion resides in the decomposition of the blood through the influence of the chorion. Thus if the maternal blood become deoxydized, it must necessarily revert to the condition of chyle. This chyle is the foetal water.
3013. The foetal water is absorbed by the embryo through the integument.
3014. No blood is transferred from the mother directly to the foetus.
3015. The blood-vessels of the uterus and placenta do not open into or communicate with each other.
3016. The foetal water corresponds to the albumen or the white of eggs, not to the vitellus. This becomes also during incubation consumed in the formation of the body of the chick, and not the vitelline mass, which is first of all destined for the intestine.
3017. Towards the termination of pregnancy, when the foetus is endowed with muscular motion, the foetal water is also absorbed. Nutrition is therefore in the commencement an absorption carried on by the integument, and lastly by the intestine.
2. _Respiration._
3018. The respiratory organ of the foetus is the chorion, and in particular the placenta. Its tissue is like that of the branchiæ or spleen.
3019. The arterial blood transmitted through the umbilical vein is conducted through the foramen ovale into the left ventricle of the heart, and from thence directly to the principal organ of the foetus, the brain and spinal cord. From hence it returns, venous in quality, to the right side of the heart, and from thence through the ductus arteriosus into the inferior or descending aorta, from which, pursuing its course through the umbilical arteries, it again reaches the placenta, in which it undergoes a renewed oxydation.
3020. If therefore the umbilical cord be compressed, the foetus dies suddenly, as happens indeed accidentally in certain cases, which completely resemble those of death produced by suffocation. The chick when in the egg dies, if the egg-shell be coated over with varnish, or if the egg be submitted to the noxious influence of gases which are devoid of oxygen.
3021. The respiration effected through means of the placenta, admits also of being proved by the sudden change which occurs in the circulation after birth. No arterial blood being then any longer brought to the heart through the umbilical vein, the left heart is no longer stimulated, and the foramen ovale collapses and is closed. Thus, all the blood enters the right side of the heart, and finding no thoroughfare into the ductus arteriosus, it is driven forcibly into the lungs, which now expand, and thereby leave an empty space between the pulmonary vessels, into which the air rushes in.
3022. The first act of respiration is therefore the result of the lungs being injected with venous blood, and thus of necessity ensues. When, on the contrary, cases arise causing pulmonary suffocation, the blood then regurgitates to the umbilical vessels, in order to reach the original branchia or placenta.
3023. Another process of respiration is found to take place in the vessels of the allantois. Its fluid becomes oxydized and penetrates through the urinary cyst into what have been called primordial kidneys. This kind of respiration through the anus is persistent in many Worms and larvæ of Insects.
3024. Finally, there is a respiratory process on the body of the embryo itself through the branchial apertures on its neck; these in the Frogs and Salamanders still continue visible after their exclusion from the egg. This oxygen must be derived from the foetal water.
3025. Without doubt also the vitelline vessels respire, and thus each vegetative system has its own process of respiration. The intestine respires through the vitelline vessels, the vascular system through the vessels of the chorion, the sexual system through those of the allantois, the integument through the branchial apertures. The lungs belong to, and perform respiration for the whole body.
_Decay of the Developmental Organs._
3026. When all the organs are developed, the intestines are drawn together with the vitellus into the abdomen. The chick is still nourished several days after exclusion from the egg by the vitelline mass, which passes through the vitelline canal into the intestine. The vitelline membrane subsequently becomes flaccid, and finally disappears through maceration. In the Mammalia the umbilical vesicle separates at an earlier period from the intestine, and continues to lie in the umbilical cord.
3027. The secernent point of the vitelline membrane or of the umbilical vesicle is the coecum.
3028. There are therefore two intestinal systems, which branch off in a bifurcated manner from the coecum, namely, the sexual intestine and the small or truncal intestine.
3029. All embryos have originally umbilical herniæ, which do not originate through protrusion of the intestines out of the abdominal cavity, but through their entrance into the latter being retarded.
3030. The umbilical herniæ therefore indicate an earlier condition of the animal, which has originated through arrest of the development.
3031. At birth all the enveloping membranes die, and their point of liberation is called the navel or umbilicus.
3032. The animal has originated through the umbilicus, and through this has it respired.
3033. All animals which breathe by the anus, do so properly through the navel; animals of this kind are fundamentally umbilical animals.
_Parallelism of the Foetus with the Animal Classes._
3034. During its development the animal passes through all stages of the animal kingdom. The foetus is a representation of all animal classes in time.
3035. At first it is a simple vesicle, stomach, or vitellus, as in the Infusoria.
3036. Then the vesicle is doubled through the albumen and shell, and obtains an intestine, as in the Corals.
3037. It obtains a vascular system in the vitelline vessels, or absorbents, like as in the Acalephæ.
3038. With the blood-system, liver, and ovarium, the embryo enters the class of bivalved Mollusca.
3039. With the muscular heart, the testicle, and the penis, into the class of Snails.
3040. With the venous and arteriose hearts, and the urinary apparatus, into the class of Cephalopods or Cuttle-fish.
3041. With the absorption of the integument, into the class of Worms.
3042. With the formation of branchial fissures, into the class Crustacea.
3043. With the germination or budding forth of limbs, into the class of Insects.
3044. With the appearance of the osseous system, into the class of Fishes.
3045. With the evolution of muscles, into the class of Reptiles.
3046. With the ingress of respiration through the lungs, into the class of Birds. The foetus, when born, is actually like them, edentulous.
3047. After birth it is suckled or fed. The milk is the nutrition continued by means of albumen; for the mammæ are verily only the albuminous vessels of the Bird, which are placed free and external in the Mammiferous animal. After the time for sucking is past the young one obtains teeth; and thereby becomes for the first time independent of the mother, and passes over into the class Mammalia. Now, should the sketch here afforded of these parallels be not in all respects correct or justifiable, still sufficient proof remains, that a perfect parallelism is found to take place between the development of the foetus and that of the animal kingdom.
3048. Animals are only the persistent foetal stages or conditions of man.
3049. Malformations are only persistent foetal conditions, or animal formations in individual animal bodies.
3050. Diseases are vital processes in animals. Pathology is the physiology of the animal kingdom. A human foetus is a whole animal kingdom. (Vid. Oken's 'Die Zeugung,' 'Beyträge zur vergl. Anatomie,' and 'Ueber die Nabelbrüche.')
_Periods of Life._
3051. If the young in the ovum or in the mother's body resembles the aquatic animals, and has passed through their organization; so after birth does it belong to the air-breathing animals and traverse their organization.
3052. One period is that of sucking; the edentulous condition of the Bird. This is called the suckling age, baby-hood or infancy.
3053. One is that of the eruption and persistence of the milk-teeth; the condition of the Rodentia, a repetition of the gelatinous animals. This is the age of childhood.
3054. One is the eruption of the permanent set of teeth until the attainment of puberty, or the development of the sexual functions; the condition of the Marsupialia; repetition of the Conchozoa or Shell-animals. This is boyhood.
3055. From the state of puberty until the development of the faculty of understanding; the condition of the Shrews and Bats, the repetition of Insects. The age of youth.
3056. The period of the understanding passes through the Ungulata, and is a repetition of Fishes, Reptiles, and Birds. This is the first stage of manhood.
3057. After the understanding the reason matures; this is the condition of the higher Mammalia up to Man; it is a repetition of the Mammalia. This is adult age or mature manhood.
3058. Then follows the decadence or dying off of the sexual functions; a retrogression through the animal classes. Gray old age.
3059. Finally, the reason and understanding die; childhood returns and terminates with the death of the vegetable in the animal.
3060. Death results through the sexual animal.
3061. Death is only a continuous growth through retrogression into the organic primary matter or Infusoria.
3062. Death is an organized decomposition.
3063. Decomposition is a forming of seeds, ova, and foetuses.
3064. Dying is a multiplication of self.
III. ZOOLOGY.
3065. Zoology is Zoogeny divided and self-substantially represented. What in Zoogeny was the organ of a single indivisible animal here becomes the organ of a separated animal, or becomes a self-substantial animal.
3066. The self-substantial animals are only parts of the Great animal, which is the Animal Kingdom.
3067. The animal kingdom is only _one_ animal, i. e. the representation of animality with all its organs, each of which is a whole for itself.
3068. A single animal originates, if a single organ frees itself from the general animal body, and yet exercises the essential animal functions.
3069. The animal kingdom is only a dismemberment of the highest animal, i. e. of _Man_.
3070. Animals become nobler in rank, the greater the number of organs which are collectively liberated or severed from the Grand animal, and which enter into combination. An animal, which e. g. lived only as intestine, would be, doubtless, inferior to one which with the intestine were to combine a skin; and that animal again must be regarded as higher than the latter, which should present, in addition to these organs, vessels, liver, branchiæ, tracheæ, and lastly bones, &c.
3071. Animals are gradually perfected, entirely like the single animal body, by adding organ unto organ. The animal kingdom is developed through the multiplication of the organs.
3072. Each animal ranks therefore above the other; two of them never stand upon an equal plane or level. Animals are distinguished by their position of stages or degrees from each other, by the number of their _different_ organs, but not by the division of a single organ.
3073. The animal system cannot be arbitrarily disposed according to this or that organ, just as it may chance to meet the eye; but only in accordance with the rigid prescripts of the animal body's genesis.
3074. The animal body separates into two series of organs, which, corresponding with, pursue a proximal course in relation to, each other; into the Anatomical systems and the Sensorial organs, unto which the sexual parts appertain.
3075. The number of the sensorial organs is 5, and they thus stand according to their genetic development one above the other:
Tactile sense or Skin. Gustatory sense or Tongue. Olfactory sense or Nose. Auditory sense or Ear. Optical sense or Eye.
3076. In animals, which are characterized by the sense of feeling or touch, the other sensorial organs must be either still wanting, or if present but imperfectly conditioned, i. e. not constituted like those of man, who is the type, pattern, or paragon for every formation.
3077. Their sensations are limited to those of general touch or feeling, and of those derived through the medium of the other senses we meet with but feeble manifestations.
3078. Their body itself will only be a tegumentary body, with the organs subordinated to the integument, namely, the _viscera_. They are therefore devoid of a true tongue, of a nose, and of ears and eyes consummated after the fashion of these organs in man; they are devoid of an osseous, muscular, and myelonal (spinal chord) system, and therefore of the nose "in toto," as being the anterior extremity of the myelon.
3079. Such are what have been called the Invertebrate animals, which are consequently, in accordance with their physiological signification, Splanchnic or _Tegumentary animals_.
3080. The tongue exhibits for the first time in Fishes a resemblance to the human structure, while their nose, ears, and eyes have not yet attained the latter grade of perfection. To the nose are wanting the posterior nasal foramina, to the ears the external auditory meatus, to the eyes the palpebræ and power of motion.
3081. In the Reptiles the nose opens for the first time into the mouth, and serves for the thorough passage of air. It is thus developed as in Man, while to the ears the external auditory meatus and cochlea are wanting, the eyes being barely endowed with lids and motion.
3082. In Birds, for the first time, the external auditory meatus, as well as the cochlea, is exhibited in its perfection, while the eyes are scarcely gifted with motion, and have only the inferior lid perfect; the tongue and nose, with the limbs also, have again become retrograde in character.
3083. For the first time, in the Mammalia, the eyes are moveable and covered with two perfect lids, without the other organs of sense having suffered degradation through this completion of the eyes.
3084. Thus in respect to the Senses there are only 5 animal divisions of equal value or worth. They should properly be called classes; but, as the lowest division, from comprising within itself the viscera or the vegetative systems, is very rich in contents; were we to call these divisions classes, many inequalities in rank, and hence also in the number of the orders and families, would originate--
1. Dermatozoa _Invertebrata._ 2. Glossozoa _Pisces._ 3. Rhinozoa _Reptilia._ 4. Otozoa _Aves._ 5. Ophthalmozoa _Mammalia._
3085. Now, with the sense of feeling, or the integument, the sexual system is associated or conjoined, and that indeed as the first or lowest development of the tegumentary system. Nevertheless, the sexual system divides into two groups, into the sexual organs, which are impressed with a true sensorial signification, and into their product, or the sexual juices, and the ovum or foetus. With these two divisions the development of the integument proceeds "pari passu."
In the ovum the tegument and its contents are not as yet separated. Both consist of a transparent mucous or gelatinous mass, as is exemplified by the vitellus and albumen; such is the case also in the Infusoria, Polyps, and Acalephæ or Sea-nettles.
In the sexual organs, however, both parts separate into membranous capsules or cysts, and glandular contents, as in the roe or ovary; milt or testes; kidneys, with furthermore the oviduct, penis, and urinary cyst. The latter are sentient membranes. Such is the case in the Bivalve Mollusca, and Snails.
In the next place the tegument becomes, for the first time, a self-substantial organ of sensation by appearing as an envelope of the body; the vesical form is then repeated, and by this means the annular character originates as a series of cysts arranged one behind the other; it is thus a veritable skin, from which, finally, the members sprout forth, as in Worms and Insects.
The Dermatozoa will accordingly range in three stages.
1. Blasto- or Oozoa. 2. Sexual animals. 3. Cutaneous "
An annulated tegument, cutis or true skin, appears for the first time in the _Worms_, with here and there lateral filaments and tentacula. True feet and antennæ appear in the _Crustacea_ or _Crabs_. Lastly, feet and wings in the _Insecta_ or _Flies_.
3086. The external sexual parts, especially the male, first make their appearance, and that indeed with a very striking amount of development, in the Snails, and in like manner the body of the Bivalve Mollusca or Mussels has become almost a complete mass of ova or roe. In the Cuttle-fishes the first traces or rudiments of urinary organs appear. The animals which belong to this group are accordingly the _Conchozoa_ or Shell-animals.
3087. Animals, which are directly resolvable into sexual fluids, or that represent parts of the ovum, are the gelatinous Infusoria, Polyps, and Acalephæ. Unto this category belong the _Protozoa_ or _Mucus-animals_.
3088. The complete subdivision of animals, according to the organs of sense, would consequently stand thus:
I. Dermatozoa Invertebrata. 1. Oozoa _Protozoa._ 2. Glandular animals _Conchozoa._ 3. Cutaneous " _Ancyliozoa._
II. Glossozoa _Pisces._ III. Rhinozoa _Reptilia._ IV. Otozoa _Aves._ V. Ophthalmozoa _Thricozoa._
3089. Unto these organs of sense the anatomical, or internal parts, are subordinated, and range parallel to them in a striking manner. The following is their order of succession in accordance with that of their origin:
1. Intestinal system. 2. Vascular " 3. Respiratory " 4. Osseous " 5. Muscular " 6. Nervous "
3090. That the vegetative systems are correctly arranged after this manner, is proved chiefly by their order of development in the animal series.
3091. The animals, occupying the lowest grade, are nothing but an intestine, which is in many instances scarcely separated or distinct from the tegument; they are devoid of vessels and branchiæ, and are barely provided with self-substantial ovisacs. Their body consists of one or two concentric cysts of an homogeneous and transparent substance--_Intestinal animals, Protozoa_.
3092. When the intestine is freed from the mass of the body, both then obtain the form and substance of tegumental cysts, whereof the external being only an intestinal tunic, thus represents the peritoneum. They are now, however, united by means of a vascular system, which is again surrounded by a cyst, and thus by a pleura. Their body consists of three concentric cysts; intestine, peritoneum, and pleura. It contains too a liver and self-substantial sexual parts--_Vascular animals, Conchozoa_.
If these cysts be repeated in the direction of the axis, the tegument becomes an annulated skin. An annulose animal is a multiplied cystic animal. Associated with this character, the respiratory organs are gradually evolved to form vascular plexuses, branchiæ, feet, tracheæ, and wings, while the sexual parts are for the most part separated--_Ancyliozoa, Respiratory animals_.
3093. The osseous system first appears in the Fishes, along with imperfect, mostly tendonless, and only white muscles, as also a myelon that is only developed into a stunted encephalon, in which the brain-organs of the Thricozoon are in great part wanting.
3094. Typical, or true muscles, provided with tendons, and of a red colour, are first exhibited in the Reptilia.
3095. A complete nervous system, pretty similar to that of the Thricozoa or Pilose animals, having a cerebrum and cerebellum, with similarly distributed and delicate nerves, is first displayed in Birds.
3096. According to the anatomical systems there are, therefore, six divisions of animals:
A. Splanchnozoa. 1. Intestinal animals _Protozoa._ 2. Vascular " _Conchozoa._ 3. Respiratory " _Ancyliozoa._
B. Sarcozoa. 4. Osseous animals _Pisces._ 5. Muscular " _Reptilia._ 6. Nervose " _Aves._
3097. The Pilose animals first originate through completion and combination of all the organs of sense. They are _Æsthetic_ or _Sensorial_ animals.
3098. The arrangement of animals, according to the organs of sense, coincides consequently with that derived from the anatomical systems, and each animal division is therefore determined by two principal organs, by a vegetative and an animal. Every animal is at once a vegetable and animal body, the inferior kinds being partly so, but the highest or the Pilose animals entirely so, or in every respect, i. e. in them are found all the anatomical systems, and all the sexual and sensorial organs.
3099. The signification of animals is accordingly as follows:
I. _Anatomical Systems._
A. _Vegetative Systems._
1. Intestinal animals. 2. Vascular " 3. Respiratory "
B. _Animal Systems._
4. Osseous animals. 5. Muscular " 6. Nervose " 7. Sensorial "
II. _Organs of Sense._
A. _Tegumental Sense._
1. Oozoa _Protozoa._ 2. Glandular animals _Conchozoa._ 3. Cutaneous " _Ancyliozoa._
B. _Cephalic Senses._
4. Glossozoa _Pisces._ 5. Rhinozoa _Reptilia._ 6. Otozoa _Aves._ 7. Ophthalmozoa _Thricozoa._
The ovum divides into vitellus, and albumen, along with the calcareous shell, and into germ or envelopes; the intestine into pharynx or stomach, bowels and absorbent vessels, thus:
1. Gastric animals Vitelline animals _Infusoria._ 2. Intestinal " Albuminous " _Polyps._ 3. Absorbent " Involucral " _Acalephæ._
The sexual parts divide into female, male, and urinary organs; the vessels into veins, arteries, and hearts, thus:
1. Ovarial animals Venous animals _Mussels._ 2. Orchitic " Arterial " _Snails_. 3. Renal " Cardiac " _Kracken._
The annulated tegument divides into papillæ, feet, and wings; the respiratory organs into tegumental network or rete, into branchiæ and tracheæ, thus:
1. Papillary animals Reticular animals _Worms._ 2. Pedal " Branchial " _Crabs._ 3. Alary " Tracheal " _Flies._
When parallelized with the organs of plants, the following remarkable relationships between them are rendered apparent:
1. Cells | Stomach | Vitellus _Infusoria._ 2. Bark | Intestine | Albumen _Polyps._ 3. Root | Absorbents | Envelopes _Acalephæ._ 4. Ducts | Veins | Ovary _Mussels._ 5. Liber | Arteries | Testes _Snails._ 6. Stalk | Hearts | Kidneys _Kracken._ 7. Tracheæ | Retia | Papillæ _Worms._ 8. Wood | Branchiæ | Feet _Crabs._ 9. Foliage | Lungs | Wings _Flies._ 10. Seeds | Bones | Tongue _Fishes._ 11. Pistil | Muscles | Nose _Reptiles._ 12. Corolla | Nerves | Ears _Birds._ 13. Fruit | Senses | Eyes _Thricozoa._
A. DIVISION INTO PROVINCES.
3100. The animal body divides first of all into the vegetative and animal. There will therefore be animals in which the former, and others in which the latter, systems predominate. The kingdom consequently separates into a vegetative and into an animal province. The vegetative parts are all tegumental developments, and thus the creatures in whom they prevail are _Splanchnic_ or _Visceral animals_, but the animal parts are developments of the flesh, and constitute the _Sarcose animals_.
_First Province. Splanchnozoa._
3101. Unto the Splanchnic or Tegumental animals are wanting bones, muscles with the nerves belonging to them, and thus the neural axis or encephalon; they are consequently devoid of bone, muscle, and brain, being in a word _asarcose_ or _fleshless_ animals.
The tegument is, however, the general organ of sensation or feeling; they are thus _Sensitive animals_.
3102. In them, developments only of the sense of feeling can occur, such as sensitive papillæ, tentacula, feet, and wings. All the remaining organs of sense can only be exhibited as rudiments of a very feeble or stunted character. They do not possess a true tongue, nose, ears, and eyes, i. e. constructed after the type of these organs in Man. The eyes only, from their being the sense of the animal system proper, can assume a definite kind of development.
3103. But, these organs of sense are the sensorial organs of the head, or rather are the head itself; the Tegumental animals are therefore devoid of a true head. They only possess such an one, in so far as it is determined by the tegument and nervous sense, by the mouth and the eyes.
These animals are what have been called _Invertebrata_, a name which is, however, defective, from its indicating the absence of one part only or of a single animal system, while the word _flesh_ comprehends bones, muscles, and nervous mass; they are _asarcose animals_. But even this last appellation is not correct, because it is negative. Their positive system, or that under which they actually exist, is the tegument; so that the name of Dermatozoa or Sensitive animals, is the only proper one.
As the tegument includes the viscera and forms therefore the trunk alone, they may thus also be designated _Trunk-animals_.
_Second Province. Sarcozoa._
3104. As in the animal body, bones, muscles, and encephalic system are suddenly associated with the tegumental system; so also does a second series of animals, having these systems, suddenly originate. Now, as the first formation of the osseous system is the vertebra, so all these animals have, as is generally understood, a vertebral column, and are on that account indeed Vertebrata; but, they are much more than this, and hence the title is of too limited a character. Besides there are among them animals, in which only the chorda dorsalis is present, without the ring of the vertebral body being formed around it.
Along with the animal systems the head is for the first time developed with its organs of sense--_Cephalic animals_.
These animals have consequently, in addition to the general sense of feeling, a true tongue, nose, ears, and eyes--_Cephalæsthetic animals_.
B. ANIMAL CIRCLES.
3105. Animal Circles are representations of entire anatomical systems as self-substantial bodies.
3106. Now, the vegetative body divides into three principal systems, into the Intestinal, Vascular, and Respiratory system, with their functions, or the digestive, nutritive, and respiratory processes. There are therefore Intestinal, Vascular, and Respiratory animals.
_Circle I. Intestinal Animals._
3107. The intestinal system is the first form of body, or that from which the other systems have not yet separated. The body of these animals consists therefore of the homogeneous primary mass--the animal protoplasma, mucus or slime--_Mucus-animals_.
Now, the protoplasma is a hollow globule. The intestinal system is therefore nothing else than the original cystic form. Thus, there are _Cystic animals_ like the Infusoria.
Again, cysts can increase in no other way, than by dividing into their like, or engendering cysts within themselves. The first kind of increase or multiplication of species is thus fissiparous, or produced by division.
The newly-engendered cysts are to be compared with the vitellus, and when they have attained perfection, to the ovum. They are therefore Oozoa or _Ovum-animals_.
In these animals consequently there are no separated sexual parts, namely, in addition to the vitelli, neither testes nor renal organs, or at least only obscure indications of them. In these Cystic animals the lowest feeling only, that of the _sex_ or _general sensation_, can occur.
The Oozoa or Cystic animals, when compared with the plants, are the first flower that has been set free, or in other words, a flower which no longer stands polarwise upon a stem, because it is not developed in the differencing air, but in the indifferent water. It may be said that nature, having brought matters as far as the development of the sexual parts, then quits or passes out of the vegetable world; while these parts, or even the entire plant, requiring now no longer the stem and root, become a root themselves, and in behalf of this enter the water.
Animals, which have the form of flowers, are round or radiiform. There are _Radiata_ or Radiated animals.
3108. These _Anthoidal_ or Flower-like animals are Infusoria, Polyps, Acalephæ; being single or double concentric cysts.
3109. We may regard the Flower-animals as the fundamental mass of the sexual parts, which has attained unto free motion. They are sex throughout, or nothing but sex; it cannot therefore be said that they possess sexual parts like the plants, but that they are sexual parts. They are sexual parts that swim.
3110. Formerly, most of these Anthoidal animals were actually taken for real plants, on account of their floral and ramular form, and even their very substance; so slightly withdrawn are they from the vegetable kingdom. The whole difference between the two is effected by the water. Could we transpose them into the air, they would then become real plants.
3111. Now, as the vegetable flower is not a mere sexual system, but is also stock or trunk; so also is the animal flower at once an organ of digestion, respiration, and nutrition. The lowest condition of these organs is, however, only one of absorption, evaporation, and rigidification; these processes will therefore be present also, though only upon the lower stage--they are Intestinal animals; for in a simple intestine the same processes could occur, only within each other, whereas they are mutually dissevered or set apart in intestine, lung, and capillary system or parenchyma.
3112. The sexual parts are themselves viscera, or the viscera themselves are sexual parts, just as the fungus is at one and the same time root and seed-capsule. The sexual parts themselves absorb, respire, and nourish.
The floral sac is not therefore a mere sexual sac, but also an absorbent sac; upon a somewhat higher grade it is even a digestive sac, the sac-wall itself being a respiratory and nutritive paries or wall.
The sexual function has at once become an ingestive function, tending unto nutrition, or the deglutition of the food is itself an act of copulation.
The sexual capsule in these animal flowers can as well be termed stomach as uterus, and its wall ovary as well as branchia.
3113. As being of a sentient, mucous nature, they are point-substance or nervous mass. The tentacula are higher organized stamina, and thus occur as cilia, surrounding the oral aperture or mouth, as in the Infusoria. These palpi or feelers are, from being organs of ingestion, both male penes as well as digits or tongues, as in the Polyps. Their structure is still wholly tubular, while their elongation appears to be for the most part effected by injection with water--they are absorbents, as in the Acalephæ. The Oozoon brings forth young in the same cavity, in which it digests and by which it respires, and impregnates itself with the same filaments, whereby it seizes, swallows, and tastes its food.
In the bottom of the cavity of the Germ-animals, granules develop, which are born or escape through the floral opening--pharynx, and again become similar beings. In others the granules also sprout forth attached to the walls of the cavity, remain there some time united with the parent animal, and thus completely represent the mode of propagation in plants by means of gemmules or buds. Among the Polyps and Acalephæ it is well known that the ova issue from apertures near the mouth; while in the Actiniæ this is stated to take place from the stomach. The ovaria are, as is well known, situated between the stomach and parietes of the body.
3114. The intestinal animals are an entire animal organism, but only in the chaotic condition. They are the fundamental tissue, the cellular system of the animal, and the higher animals are only separated cells.
3115. The propagation is in every respect similar to that of plants. Now, as the seeds are the whole plant upon a small scale, so is the granule or ovum the entire animal; it is liberated through the pharynx and continues to grow merely by increasing in size. But if the young animal protrude through the tegument, that is a true gemmiparous propagation.
3116. Those Oozoa, which like plants can develop buds, consist of several animals, and may be cut in pieces like plants; when each piece becomes again an entire animal.
3117. The Oozoa represent those products of nature which are prior or antecedent to the animal world; namely, first of all plants, and further still the inorganic kingdom also, or the earth, since they have originated in the water and can be as well developed from the stones as the Lichens. There are therefore Lithozoa or Stone-animals, and Phytozoa or Plant-animals, among the Oozoa.
3118. Would we compare these animals with the parts in plants, they then represent their cell-development, or cells, bark and root. They are themselves either vesicles, as the Infusoria, or barks, as the Corallines, or a root-like fascicle of tubes, as the Acalephæ.
3119. In the animals, however, the cells have become stomach, the bark intestine, the root absorbents. The Oozoa represent therefore the aggregate of the intestinal system, the primary mass of the animal body.
Their whole body is a digestive body, or parenchyma, traversed in many species in all directions by tubes or absorbents, as in the Acalephæ.
As yet no nervous filament, nor any muscular fibre, &c. has been separated from their mass, as in like manner a tegument has barely been freed or separated from the intestine. These animals are nerveless and sine-tegumental, precisely because they are wholly made up of nervous mass and tegument.
_Circle II. Vascular Animals._
3120. In the next place the digestive separates completely from the tegumental function, and each constitutes a function for itself, but which now being separated from its fellow could no longer subsist. Between the two therefore is formed the nutritive function in the _vascular system_.
Now, the vascular system of the self-substantial intestine is the liver. This organ will therefore for the first time make its appearance in this animal circle--_Hepatic animals_. Upon a higher stage, salivary glands are also developed on the intestine, and will here likewise begin for the first time to appear--_Salivary animals, Snails_.
3121. Through the separation of the viscera from the remaining substance, this must necessarily remain behind as a hollow cyst or tegument investing the former. The true free tegumental formation is therefore by no means accidental, but is necessarily bestowed with the viscera in the course of animal development. This tegument is _peritoneum_. There are animals which are invariably bicystic, but consisting of concentric cysts--_abdomens_.
Around the ventral integument, however, the vascular system also forms its tunic; this is the branchial membrane or operculum--_pleura_, mantle. There are tri-cystic animals--intestine, abdomen, and thorax disposed concentrically around each other--Mussels. Their body is therefore not articulated, but its parts are still inserted within each other.
The Vascular animals are consequently multiplied Protozoa; or Mucus-animals of the second power--_Ovum_².
The sense of feeling ascends upon its second stage, when its organ, having been freed from the mass of the body, surrounds the viscera as a self-substantial tegument. The feeling is then no longer one of a merely general character, but is a definite perception of external objects, a passive power of feeling.
3122. True muscles could not as yet originate in this skin, and that for obvious reasons, although fibres are present; for the former are to be brought under the signification of arterial fibres.
3123. Cilia, when furnished with fibres by whose agency they are rendered moveable and susceptible of inversion, are called _tentacula_, which here occur under every variety of form.
3124. Would we compare the Vascular animals with the plants, they must then represent their ducts, liber, and stalk. The heart is the stalk, the arteries liber, the veins tubes or ducts. These animals have assumed also upon the whole the cauline or cylindrical form--Cauline animals.
3125. From their having obtained in addition to the intestine only the vascular system, they are still governed by the water, and live therefore for the greatest part in this element. They have the first mode of respiration and thus an aqueo-respiratory process--possess branchiæ but not tracheal tubes.
3126. The sexual parts, which in the Germ-animals were still coalesced for the most part with the body, here become self-substantial through the separation of the teguments, make their appearance as a repetition of the digestive system under the condition of a free or separate system, and are evolved into true ovaria and even male parts--_Sexual, Glandular animals_.
3127. The first step towards the evolution of male parts is, however, only half achieved. Only one testis originates, while the other remains behind as ovarium--_Androgynous_ or _Bisexual animals_.
3128. These animals, which are characterized by the vascular system, and by the first self-substantial or external sexual parts, which indicate organs of sense, are the _Conchozoa_, such as the Mussels, Snails, and Kracken.
_Circle III. Respiratory Animals._
3129. When once the intestinal and vascular system, through perfection of their individual parts, such as the liver and branchia, and through separation of the sexual parts, are completed; then the individualization of the tegument steps into view, and it becomes an independent _respiratory system_.
3130. Through the increased process of oxydation the tegument hardens and is converted into _horn_. All induration, however, only takes place in opposition to soft places. The tegument therefore separates into hard and soft rings--_Annulate animals_.
3131. The annulate tegument is a tracheal tube wholly converted into a body. To distinguish it from the general tegument it may be called the cutis or _skin_--_Cutaneous animals_. The annular tegument may be regarded as a series of cysts placed one behind the other. The Annulate animals are therefore multiplied Malacozoa; Mucus-animals of the third power--_Ovum_³.
The respiratory organs, being in their lowest condition, will not as yet be freed from the tegument; the vessels simply form a network or projecting filaments and lamellæ--_reticular branchiæ_; as in the _Worms_. The tentacular organs, from being yet soft and thus scarcely moveable, are still very imperfect. Upon the lowest stage the tegument or _skin_ simply feels; in the next place papillæ, and finally filaments, originate, especially about the mouth--_Cutaneous_, _Papillary_ and _Filamentary animals_.
3132. If the tegument, as being the original branchial membrane, is converted into horn; then the branchiæ cannot continue as retia, but must elongate above the tegument into filaments, ramules or lamellæ.
With this, these elongated branchiæ separate into two organs, one part of them becoming indurated in like manner with the general tegument, and supporting the other as gill. Horny branchial filaments, which contain vessels, nerves, and fibres, are called _feet--Pedal animals_.
3133. The limbs or members of these animals are simply hollow tegument, hollow hair, and are therefore thoroughly different from the bones or the animal system.
The tegument thus hardens around the soft parts and the viscera. A horny coat of mail originates, and thus we have horny or mailed animals, in opposition to the Malaco-or Conchozoa.
3134. Beneath the horn, however, there must still be soft skin, and this becomes fibrous by the strong oxydation which it undergoes. Fibrous fascicles are attached to the coat of mail and to the hollow limbs, and are consequently within the tubes.
3135. These fibrous bundles are not flesh, but a fibre-drawn _tegument_, so that there are also no true muscles. They must on that account too be numberless.
3136. The articulations or joints are external not internal; they thus consist of tegumental tubes, not of bones, abutting against each other, and are not surrounded by flesh. Hence, like all the preceding groups they also are devoid of flesh--asarcose animals like all the preceding ones.
3137. In the branchiæ, however, it is only the cauline portions which become horny, while their ramules continue to perform the respiratory function. The branchiæ therefore are appended to the extremity or roots of the feet; or rather these latter form the branchial arches.
Annulate animals, having true or hornified and likewise annulated feet, are called Crustacea or _Crabs_.
Thus, in these animals the sense of feeling obtains special and moveable organs; they are _Tactile animals_.
Tactile organs are prolongations of tegument moved by muscular fibres, which in some degree adapt themselves by pressure unto the forms of objects, or have the power of seizing and retaining them--such as feet, antennæ, maxillæ, palpi.
3138. But if the tegument be entirely converted into horn, and the respiratory vessels have in this way disappeared within it, then the internal respiratory organs are formed by inversion of that part of the tegument which is between the rings, and through these openings the air penetrates to the internal parts--stigmata, tracheæ.
3139. The tracheæ can first of all originate when the respiratory process has attained its highest development, or, in other words, when the creature breathes air.
3140. Lastly, in the air-breathing Ancyliozoa even the external branchial lamellæ harden and are converted into _wings_--_Pterozoa_, _Insects_ or _Flies_.
The wings of Insects do not correspond to the wings of birds; they are not feet, but pedal appendages or _branchiæ_, and thus are no new or unknown organ.
_Circle IV. Sarcose Animals._
3141. The second animal province may be regarded as the fourth stage in the self-substantial development of the anatomical systems, although, properly speaking, it ranks as regards its value or worth upon a level with the three former circles, and resolves itself directly into three stages, which accord with its three systems. But since these stages are at the same time also classes; they should, for uniformity's sake, retain the latter name.
3142. The animal kingdom accordingly divides into _four_ great districts or circles.
CIRCLE I. _Intestinal Animals_, Oozoa--Protozoa. II. _Vascular_ " Sexual animals--Conchozoa. III. _Respiratory_ " Cutaneous " --Ancyliozoa. IV. _Sarcose_ " Cephalic " --Vertebrata.
C. ANIMAL CLASSES.
3143. The animal Classes may be designated as the self-substantial representation of a stage in the development of an anatomical system or of the inferior organ of sense, and in the Sarcose animals of these systems themselves or of the cephalic senses.
3144. There are then as many classes as there are stages of development or systems. Thus the intestinal system separates into pharynx or stomach, intestine and absorbents.
3145. The vascular system into veins, arteries, and hearts.
3146. The respiratory system into branchial membrane or skin, into branchiæ, and into lungs or tracheæ, i. e. air-tubes. Taken in a strict sense, such divisions do not constitute classes, as has been already remarked.
3147. It is only the animal systems, which do not divide into several functions, but remaining upon a level with each other, are simply repeated in the higher organs of sense.
_First Province. Splanchnozoa, Dermatozoa._
FIRST CIRCLE. INTESTINAL ANIMALS, OOZOA.
3148. The intestinal animals are nothing but depressed cysts. They rank therefore upon the lowest stage of development, and consist of mucus or granular nervous mass--_Protozoa, Gelatinose animals_.
3149. It is an established fact that, with animals as well as plants, the first function consists in imbibition or absorption, and the body must consequently be an absorbent cyst or a pharynx, which nevertheless takes up the food in the same way that the tegumental lymphatic vessels absorb. We can therefore style these animals pharyngeal or gastric, although the name is not perfectly correct--_Infusoria_. Then the intestine associates with the pharynx or stomach, so that they are Intestinal animals--_Polyps_ or _Corals_.
Lastly the intestine sends out absorbents, and then the animal consists of pharynx or stomach, intestine and absorbents; such may be termed an Absorbent animal--_Acalephæ_.
The Intestinal animals therefore divide according to the developmental stages into three Classes.
_Class 1. Gastric, Vitelline Animals._
3150. The lowest animals commence with the water, which has scarcely become mucus; they are nothing but drops, vesicles, which swim about independently--_Protozoa_ or _Primitive animals_.
3151. The Protozoa correspond to the vitellus or the male semen, which is nothing but vitellus in a state of solution. They are the animal semen of the planet, the animal dissolved. Animal generation cannot take a deeper commencement or origin. The stone, which is decomposed into carbon mixed with water, can become nothing less than a point. They are the animal germinal powder. The fungus is a tissue of vesicles, which dissolves directly into seeds--fungus-powder or dust. Thus, they are ovaria or testes which have dissolved into seed, fluid testes--_Vitelline, Seminal animals_.
3152. The vitellus or semen is point-or nerve-mass dissolved. The Vitelline animals are sentient or nerve-points, which have combined all other processes in this identical mass. The divided point-mass ranks, however, in the signification of vesicular or cellular tissue. These animals are nervose cells.
3153. Nerve-cells must originate in every water, because every water is in a state of tension with the earth and the air; thus, dissolves the former and absorbs the latter. The water itself is a digesting and respiring mucus.
3154. The nerve-cells have an internal cavity, from the surface becoming oxydized and consequently converting itself into a denser layer, or into tegument. This, however, can only take place at the expense of the internal mass, as being that alone which abuts against the external parietes and becomes rigid.
3155. As the animal life is not simply a single act of rigidification, but a repetition of the same, alternating with solution, so must the Protozoon again replace the mucus-granules which have been disposed of from its interior; it must eat.
3156. It is a matter of indifference for the philosophy, whether the reception of food is effected by one or several mouths. There are Acalephæ and even Intestinal worms (Entozoa), which absorb by several mouths, almost like plants.
3157. But in the animal the mouth or mouths is or are definite organic openings, not merely interspaces or pores as in the plants; for they rank in the signification of the blossoms, or of composite parts.
3158. It can be therefore said that every animal hath a mouth or mouths, and consequently a stomach or stomachs.
3159. Their motions consist in abbreviations or straitenings of the cyst. Indications can scarcely be present of the secretions of higher organs, such as the intestine, vessel, branchiæ, liver, and such like parts.
3160. On the contrary, developments of the tegument and nerves may occur, the former as cilia which serve also as organs of motion and as branchiæ, the latter as ocular points or rudimental eyes; for both are none other than nervose tegument. The mouth of these animals is still passive, and subordinated to the water. It is surrounded only by cilia, which by the vortices produced by their movements in the water, impel or draw the latter into the mouth and with it the food. Such animals are called _Infusoria_.
3161. As the Infusoria are the seed or vitellus itself, so also are they the very ovum, and require no special sexual parts for propagation. They absorb, and are so nourished, and if there is a sufficient quantity of the mass to admit again of its division into several points, they thus divide. A magnified Infusorial animalcule has become, as it were, an ovarium or a testis, which then produces seed by dissolving itself into the latter. They are a constant conflict of the organ and its product, of the Solid and Fluid, a vitelline and orchitic process.
_Class 2. Intestinal, Albuminous Animals._
3162. With the separation of the cyst into an internal and external layer, or into intestine and tegument, the animal must necessarily ascend a stage higher, since it now contains two systems different from each other, and is consequently a double Infusorium.
3163. The form of the _Intestinal animals_ passes gradually from that of the globe into the tube--Tubular animals.
3164. They are tubular nerves surrounded by a tegument.
The cilia also are perfected and lengthen into filaments, which no longer perform simply vortical movements, but now actually seize and convey the food self-substantially into the mouth. Such animals are called _Polyps_. Their multiplication takes place no longer by division or fissure, but by ova and shoots, or ramification.
The oviducts or egg-tubes lie between the intestine and tegument, and open upon the margin of the mouth between the tentacula. In many instances the ovisacs, or cysts, hang also freely from other parts of the body, as in the Sertulariæ.
The sprouts or offshoots detach themselves from the parent and become independent animals; but they frequently continue to stand as ramules upon the maternal animal, though they nourish themselves independently of the latter.
3165. If the process of oxydation be augmented, then, the tube's external wall indurates, becomes coriaceous, and, lastly, ceratoid or horn-like in texture. The nerve-tubes or the animal proper can now swim no longer, since one kind only of motion is left it, namely, to protrude itself out of, and then retract itself within, the tube. It consequently falls to the ground, and while the external mucus hardens, it clings to the former; such are the Sessile or fixed Polyps.
3166. Sessile Polyps having coriaceous or horny tubes are called _Plant-animals_, Zoophyta, Phytozoa.
3167. The adherent, dried and dead external tegument of the Polyp is called _stem_. Such a ramified stem completely resembles a plant.
3168. These woody or herbaceous stems are not rooted in the earth, but have the power of adhering firmly to every substance, to stone, glass, shells, and such like bodies. They do not therefore draw in nourishment through any root.
3169. The ramification is often wholly plant-like in character, resembling that of a shrub with separate ramules, which even assume too the form of leaves, and the animal tubes that of flowers.
But frequently the ramules grow together also by their extremities, giving rise to a trellis-work, the production of which in plants is impossible. The soft animalcules, which come in contact, cleave unto each other, and grow together like wounded parts in the Sarcose animals.
3170. Upon the surface of the ramules or the leaves are apertures, out of which the mucous substance protrudes the radiated mouth. But these mouths are frequently, especially in the Cystic corallines, of two different modes of formation. The one kind are cysts without filaments, and contain ova, which are developed and fall out. The others have filaments, which move and do not produce ova. The former look like seed-capsules, the latter like flowers with stamina, while the entire trunk resembles a monoecious plant.
3171. With increased oxydation calcareous earth is deposited in the external tegument or rind, and the stem is converted into stone--_Lithozoa_, Lithophyta, _Corals_.
This calcareous earth contains the most general acid, or the carbonic, and thus oxygen combined with the inorganic carbon, while the bones contain phosphoric acid, oxydized gelatine.
3172. As the calcareous earth is, properly speaking, only a granular deposit in the tegument, like it is in the cartilages of the higher animals; so is it not to be regarded as a free worm-tube, but as the body itself. Meanwhile it forms a tube open above, from which the mouth of the animal projects.
3173. As the animal ramifies, so also do the stony tubes increase, and there originates a phytoidal or plant-like stem, but one consisting of a stony mass.
3174. Thus the Coral is the earth-animal, and indicates the globular or osseous mass under its first grade of formation in the animal kingdom.
But there are also Polyps, whose stem only originates through saccular inversion of the upper portion of the animal's body; yet this is only distinct in the soft stems. In most of the species, where a separate intestine is to be found, it is probably only the upper part of the body which is so inverted. Meanwhile there are some, whose intestine forms a circle by returning upon itself, and opens into an anus.
3175. If then the Infusoria are the vitellus or seed of the animal kingdom; in like manner the Corals are its ova. The carbonate of lime is the shell surrounding the albumen, while the animal or intestine is the vitellus--_Albumen-animals_.
3176. Nature forms these living ova, when she takes vitellus and albumen out of the sea-slime, invests them with a shell derived from the earth, and hatches them, after they have been vivified by sun and air.
_Class 3. Absorbent, Involucral Animals._
3177. Did the former animals remain in the condition of ova, from want of a perfect vascular system; they are next developed, so soon as vessels appear and form a vascular plexus, into foetal involucra or envelopes. These animals are vitellus with the vascular membrane.
3178. The Absorbent animals are no longer simple vesicles, but large cysts or capsules like the developmental envelopes of the foetus, along with a choroid plexus--_Involucral, Foetal-animals_.
This choroid plexus does not, however, consist of arteries and veins; but is only a ramification of the intestine, so that the vessels are of a lacteal character--_Absorbent animals_.
3179. In these animals there is no longer any egg-shell, but everything has been taken up into the galvanic circle; the shell has itself become organic and life-imbued. Their substance is still mucous or albuminous; they are still vitellus, though converted into a vascular tissue.
3180. They therefore cling firmly nowhere; but swim about freely, like brain-masses converted into radiated cysts.
3181. Free Mucus-animals, traversed by vascular plexuses, are _Acalephæ_.
3182. There are Acalephæ which are simply air-sacs, like the air-sacs of ova, to which hang ramified vessels as absorbent tubes--Cystic, Tubular Acalephæ.
Others represent hemispheres with numerous absorbent tubes, which concur in the middle of the animal to form a kind of stomach, from which again other tubes pass towards the border, in order to elongate into tentacula. Thus the absorbent vessels have become motor and sensitive organs.
Besides this, many have around the mouth four large lobes, which must be viewed as the antetypes of the sentient lobes of the Bivalve Mollusca.
Lastly, others have a true mouth, which leads to a similar gastric cavity, out of which the same vessels emerge and ramify. Both kinds are called Hutquallen or Acalephæ.
There are yet others having the same structure, but oviform, with respiratory lamellæ upon the absorbent tubes, Rippenquallen or Beroes.
3183. An Acalephan is a brood-egg, which swims freely about without a shell.
3184. The vessels are quaternary in their distribution, and form a cross, like the involucral vessels of the chick.
3185. In these animals the ova first begin to be detached and agglomerated together in definite situations so as to constitute ovaria. The number here is also four.
They lie usually within four cavities surrounding the stomach, and into these wide apertures situated near to the mouth lead; they are at the same time regarded as respiratory cavities.
In other species vesicles, wherein seminal animalcules are developed, occupy the same situation. Here therefore is recognised for the first time a separated sex.
In the Röhrenquallen or Physaliæ the ovisacs mostly depend externally in the form of bells.
Besides this we find in the Acalephæ all sorts of laminæ, which are probably organs of respiration.
It is not known what is the meaning of the air-sacs and of what the air consists.
Most of the species emit light like globes of fire, just as many Infusoria do also. It is probably a phosphorescence given out by the mucus when passing over into a state of decomposition.
Very many have also the stinging property of the nettle; but, whether the cause of this is chemical or mechanical, is not yet exactly known.
SECOND CIRCLE. VASCULAR, SEXUAL ANIMALS.
3186. So far, or in the ascending scale up to the Acalephæ, the animal is only viscus with an absorbent canal, which is at the same time a canal of evacuation, without a distinct intestine being set apart for that purpose; such is the general rule.
3187. After the Acalephæ the formation undergoes a change; the distinction between the exterior and interior is prominently displayed, and the internal wall becomes freed as a separate and perfect intestine along with a mouth and anus; the external wall appears as a free tegument. But, seeing that two concentric and separated cysts could not subsist without combination by means of the nutritive system or the vessels; a perfect vascular system is formed, divided into veins, arteries, and hearts.
The tegument, wherein the vessels become self-substantial, is the branchial membrane. There is therefore placed around the intestinal body a vascular body also, or branchial membrane, which is consequently pleura or mantle (pallium), as in the Mussels. The body of the intestine consists of intestine and peritoneum; that of the vessels of branchiæ and pleura or mantle.
These animals are hence bisystemic, being both Intestinal and Vascular animals; but, since the vascular system is a new addition, it is thus characteristic of the circle, and its members must be therefore called _Vascular animals_. With the vascular system, however, all its further developments have been bestowed; thus especially the complication of the vessels with the intestinal ramifications or, in other words, the liver--_Hepatic animals_. The salivary glands also are a similar complication, and in this series therefore they make their appearance.
Lastly, the renal glands are such a kind of vascular organ, or branchiæ of the sexual parts; they also begin to be astir in this series.
With the separation of the systems into separate teguments or membranes, the sexual parts also separate. The ovarium becomes an independent organ furnished with its own excretory ducts; the male parts are individualized to form veritable testes furnished with excretory ducts, or even with a penis. Still this is all effected only by degrees, or as yet within the confines of this circle.
These animals divide according to the viscera into Venous, Arterial, and Cardiac animals; according to the sexual parts, into Ovarial, Orchitic, and Renal animals.
_Class 4. Venous, Ovarial Animals._
3188. With the protrusion into view of the vascular system, the veins are chiefly developed along with their principal organ, the liver, as being the bond of union between the circulation and the intestine.
3189. The animals, which first bring to bear in addition to the intestine a liver, are the Bivalve Mollusca.
3190. With the veins arteries also originate, but with a preponderance of venosity or the venous character. The blood is lymphatic, colourless.
3191. The cardinal venous organ or the liver evokes also into existence a corresponding organ of respiration, namely, free branchiæ with the tegumental form--branchial laminæ or leaves.
3192. In the middle between the branchial laminæ and the liver, the heart is evolved; this organ comprising always a ventricle with an auricle, but consisting of a venous, membranous tissue almost devoid of fibres.
3193. The first heart is in other respects arteriose; it receives the blood from the branchiæ and transmits it to the liver as well as to the remaining parts of the body, from which it proceeds directly to the branchiæ without entering any venous or right heart.
3194. In these animals, as is well known, four branchial leaves lie externally on the belly, which includes the intestine with a large liver, and hangs together with the branchiæ as a separate purse within the mantle.
3195. In the Mussel a structure originates for the first time, which can be compared with a thoracic or pectoral cavity. What covers the branchiæ, must stand in the signification of the thorax or chest. The pallium or mantle of the Mussels is pleura.
3196. Their shells are branchial opercula (as in the Fishes). They are secretions from the mantle, and everywhere accompany the branchiæ.
3197. The locket or hinge of the shell-valves corresponds to the rachis or spina dorsi, as is especially distinct in the Teredines or "ship-worms."
The shells of Bivalve Mollusca are a calcareo-thoracic box, open in front, inverted behind, and moveable like ribs.
3198. The two sphincter or occlusor muscles signify shoulder and haunch.
3199. These animals begin for the first time to exhibit bilaterality or symmetry, because in them there is stirring the idea of the osseous formation. From the branchiæ being situated symmetrically upon both sides, the cardiac chambers are also symmetrical.
3200. The pectoral tunic (mantle) usually elongates at the anal extremity into two tubes, respiratory tubes, through which the water is drawn in and thrust out or expelled. Such is the case in the highest Worms or Holothuriæ, only there the respiratory tube leads into the body itself. A similar arrangement is found in the Echini. Many aquatic larvæ from all kinds of Insects or out of different classes breathe through anal tubes. All these animals consequently repeat the Mussels, and this formation admits of being followed out up to Man, where it is left as allantois and primordial kidneys. The thorax of the Mussel thus opens by the anus. But, since the thorax is that which is here preponderating and is nearly the whole animal, so does the anus open into the posterior respiratory tube.
3201. The Mussel can be regarded as an animal consisting of three cysts or sacs inclosed within each other, namely, of intestine, around this the ventral pouch, and around this again the thoracic sac or mantle. If, moreover, the shells be regarded as a cyst, the animal then consists of four sacs. The heart and branchiæ lie within the thoracic cavity; intestine, liver, and ovarium within the abdominal or ventral cavity. The Mussel is thus a doubled Acalephan.
What is termed foot in the Bivalve Mollusca is nothing else than the abdominal tegument dilated in front to form a muscular ridge.
3202. The Mussels are embryos, in whom the liver originates, and whose chorion acquires a placenta. Then again, as the embryo is nearly all liver, and hangs surrounded by a watery fluid within the widely-expanded chorion and amnion, so does the abdominal pouch hang within the pallial cavity, or in the wide, water-full chest.
3203. In the abdomen there is still only the ovarium, and that indeed very large. There are properly two ovaria, each of which, according to my observations (vid. Göttinger Gelehrte Anzeigen, 1806. Stück 148), opens laterally under the shoulder-muscle and gives exit to the ova, which then enter within or between the branchial fringes, in order to be there developed.
3204. Here the respiratory organs are still at the same time a kind of uterus. The ova may be oxydized therein, like the embryo is in the uterus. These branchiæ are probably still to be compared with, or designated as, sexual branchiæ.
3205. In the back of the shoulder these animals have a highly vascular organ with two excretory passages, which open near to the orifices of the oviducts. I formerly regarded it as a kidney, but, according to more recent observations, it should be a testis. In the anterior part of the foot there is frequently situated a gland which ejects a gelatinoid moisture, which hardens into glutinous filaments--the byssus or beard. It is probably a memento of the tentacula of the Acalephæ. There is besides also an organ in the feet of many Bivalve Mollusca, which occasionally squirts a watery juice out to a great distance. I have occasionally found this organ in our Teichmuschel. Its signification is probably the same as that of the preceding organ.
3206. The organization of the Bivalve Mollusc can be thus most distinctly described; it is an abdomen, wherein is an intestine with mouth and anus, a liver and a double ovarium; upon the sides of this abdomen are situated the branchiæ in the form of four laminæ; around the branchiæ and the abdomen is the pleura or mantle, which is always open posteriorly.
3207. The mouth is placed directly at the anterior extremity of the abdomen, is devoid of neck and head, as also salivary glands; it is consequently not a true mouth, but only a pharyngeal aperture. Thereupon, however, are situated four sensitive lobes, which are in structure exactly like the branchiæ--cephalic or pharyngeal branchiæ. They are the further formations of the four arms of the Acalephæ.
3208. The Mussel has a perfect splanchno-neural system with ganglia and a pharyngeal ring, which probably corresponds to the nerves that sweep around it.
3209. The Mussel has no other organ of sense than that of the passive sense of feeling, the tegument. It cannot once move its sensitive lobes voluntarily; it has no lips.
3210. The abdomen only elongates in most of them to form a moveable, variously constructed foot or _keel_, which cannot, however, creep but only push. The progression of the Bivalve Mollusca is backwards, as in the Acalephæ. In the Snails the ventral surface first becomes a creeping _sole_.
3211. The Mussels repeat the Infusoria; are Infusoria with a bivalve calcareous testa or shell.
_Class 5. Arterial, Orchitic Animals._
3212. In the preceding class of Bivalve Mollusca, it is in truth the abdominal viscera only, such as the intestine, liver and ovarium, which have been perfected; and then the veins and arteries with a membranous heart. The cephalic organs, eyes, maxillæ, salivary glands, and even moveable lips with tentacula, are wanting, as well as the muscular heart. Lastly, the arterial sexual system or a self-substantial testis and the penis were absent.
Now, bivalve Mollusca, which have eyes, maxillæ, a muscular heart and a ventral sole or foot, with salivary glands and a penis, are Gasteropoda or _Snails_.
3213. The Snails possess salivary glands, a trace of the tongue and of maxillæ, moveable lips and tentacula, with thus an approximation to the head, unto which the eyes are rarely wanting--Salivary animals.
3214. With the development of the head or rather of its inferior organs of sense, the antagonism makes its appearance also in the ovarium. A half of the ovarium is converted into the testis.
The Snail is therefore a Mollusc, which is female upon one side, male upon the other.
3215. The androgynous or bisexual animal is, as a general rule, asymmetrical.
3216. The mantle also or the branchial cavity obeys this want of symmetry. The branchiæ of one side dwindle down; those of the other turn with the mantle towards the head, and the respiratory aperture occurs upon the back.
3217. With the one-sided evolution of the mantle, one shell also is only developed, while the other is stunted or placed under arrest. The Snail's shell is one of the Bivalve Mollusc's shells, its operculum is the other. This last is stony, horny, and finally is entirely wanting.
It is remarkable that the right shell has been pretty generally perfected, while the left dwindles down into the operculum; all the Snail's openings are therefore upon the right side, such as the anus, with the orifices for the escape of the ova and semen.
Male animals are situated on the right side, female on the left; or where the right side was predominating, there originated the male sex, where the left, the female.
3218. As the orifice of the mantle and the shell is properly the aperture of the branchial foramina; so can it be said, that the Snail were a Mussel, which does not simply extend the foot, but also the mouth or head to the branchial opening; it is a Mussel reversed.
3219. Every thing else in the Snails is arranged in accordance with these fundamental organs and forms.
The cephalic portion of the intestinal canal is indicated also by muscular fibres. The pharynx and the mouth can contract and expand, seize and bite off; the former frequently admits of being protruded and retracted as a fleshy proboscis with perforating maxillæ.
3220. Since the muscular fibres are only tegumental fibres, and therefore lie within cavities; they thus act like as in the feet of Insects.
The tentacula of many Snails are moved like the feet of Insects; but as they are not horny, but soft, they are turned either inside out or "vice versâ." The oviduct and seminal duct or penis follow the same mode of formation. They are likewise everted and inverted.
3221. These members of the Snail are true Insect-limbs that have remained soft, and are thereupon susceptible of inversion and eversion. Were an Insect's foot soft, every one will admit that it would then yield so as to become inverted, if the fibres pulled upon it. The limbs of Insects have thus only become stiff, and are thereby Snail's horns that resist inversion. All these members are teguments, and give the lie only unto limbs; for it belongs to the essence of a limb, that it be dense.
3222. That which suffers eversion or its converse is no limb, but only a sheath, a prepuce. Nearly the whole Snail is but a prepuce, a "membrum virile" or "verge."
3223. There is no class of animals, in which the testes and penis are found so disproportionately developed as in the Snails--_Orchitic, Penes-animals_.
3224. The vascular and nervous systems are related pretty nearly as in the Mussels. But the heart is fleshy and has, by reason of its unilateral or single branchia, only one auricle also.
3225. The Snails repeat the Corals in the cylindrical form of the body, the tubular-shaped shell, and the return of the intestine upon itself towards the mouth, as in many Corallines.
3226. In them also the organ, which virtually corresponds to the kidney, appears to be astir, namely, what has been called the _calcareous gland_, situated within the branchial cavity, and which opens not far from the anus.
_Class 6. Cardiac, Nephritic Animals._
3227. Hitherto there has been found only a single "cardia," namely, the left or arterial heart, which, receiving the oxygenated blood from the branchiæ, propels it to all parts of the body--for the purposes of nutrition. But now also the right heart suddenly makes its appearance, which drives the blood into the branchiæ--for oxygenation. This must doubtless be regarded as a higher development, especially since, as I have shown, in the higher animals as well as in the embryo, the right heart is first perfected subsequently to the left.
The Conchozoa with a double heart are the Sepiæ or Cuttle-fishes. These are in consequence correctly called _Cardiac animals_, and must be regarded as the fundamental form of this stage.
With this completion of the heart other structural changes, which probably stand in intimate connexion with it, appear. In the Snails indeed a kind of renal organ, or what has been called the calcareous gland in the branchial cavity, has been already shown to exist. Whether the shoulder-gland in the other Mollusca belongs also to the members of the present class, may be left undetermined. This kidney, like other vegetative organs, pours out its secretion quite involuntarily. In the Cuttle-fishes it is, however, combined with an organ, by means of which it can discharge its fluid, or the ink, voluntarily as in the higher animals. The kidney or ink-gland with the ink-sac is therefore characteristic likewise of the Sepiæ, and admits of our calling these creatures _Nephritic_ or _Renal animals_.
At the same time the whole form of the body is altered; it becomes cylindrical, and is provided upon the ventral aspect with neither a muscular keel nor sole, whereby it might glide or shove itself along; thus herein also a resemblance is manifested to the higher animals.
This motionless body is on the contrary endowed with independent organs of locomotion, namely, with fins or arms, which are wanting in the Snails and Bivalve Mollusca. The labour of motion is consequently removed by the organs which minister unto it from the body, and thus we trace another similitude to the higher animals, which transport their body from place to place by wings or feet. These animals can be therefore called _Cylindrical Snails_, out of contrast to the _Sole_ and _Carinate Snails_ or the Bivalve Mollusca.
Closely allied now to these cylindrimorphous Conchozoa are all those Mollusca in whom the keel (carina) or sole is wanting; i. e. such snail or mussel-like animals, which either cannot transport themselves at all, or effect this by means of fins or podoidal appendages, in short all that have been called Snails, having no sole, and all supposed to be Mussels, but without a keel.
Now, the Heteropoda or Pterotracheæ and the Pteropoda or Clionæ have projecting processes which act like fins.
The Brachiopoda or Terebratulæ, though unendowed with the power of locomotion, have, as their first name implies, arm-like organs.
The Cirripedia or Lepades, being likewise without the power of continuous motion, possess podoidal or foot-like appendages. In both these families, however, these appendages are nevertheless true motor organs; for by means of them they seize their food, an act which hitherto has been seen to take place neither in the Snails nor Bivalve Mollusca.
Lastly, the prehensile organs dwindle down into mere filaments or small lobes, but the body always retains its cylindrical form without keel and sole, as in the Ascidiæ or Meerscheiden.
In the Salpæ only the cylindrical body is left, unto which appendages are still, and not rarely either, attached, serving them wherewith to lay hold of each other. In accordance consequently with their external form all these animals appertain to one class along with the Cuttle-fishes. I call them _Kracken_.
The renal organs are still not to be found in all of them; the structure alone of the heart agrees essentially, despite of its simplicity, with that of the Cuttle-fishes. In the Ascidiæ, simple as well as compound species, it is indeed but a single, though muscular pouch. Now, this pouch drives the blood alternately, at one time into the branchiæ, at another backwards into the body, and is thus in the first act a venous or right heart, in the latter an arteriose or left; consequently, according to function this single pouch is compounded or made up of the two hearts.
The branchiæ deviate entirely from those of the Mussels and Snails, and exhibit a very complicated structure; in the mussel-like families they present the form of a trellis-shaped sac, as in the Ascidiæ, or that of filiform appendages to the feet, as in the Cirripedia, or they are funiform as in the Brachiopoda. In the Pteropoda they are very varied; in the snail-like Heteropoda they are mostly pectiniform or tuft-shaped; in the Sepiæ or Cephalopoda phylliform or fin-shaped.
The external character ccommon to the whole class is the _cylindrical body_, unto which may be aptly conjoined the possession of special organs of motion; whether such organs be prehensile in function, media of support, or veritable instruments of progression.
The snail-like Pteropoda, Heteropoda, and Cephalopoda, have a head, but this is wanting in the mussel-like Salpæ, Ascidiæ, Cirripedia, and Brachiopoda.
THIRD CIRCLE. RESPIRATORY, CUTANEOUS ANIMALS.
3228. Respiratory animals are Dermatozoa with a predominating system of respiration.
Now, the respiratory system is the tegument, which here then must attain the highest pitch of perfection.
This again takes place through the increased process of oxydation, which produces induration of the parts.
The vessels, which surround the tegument, must nourish this portion of it to a greater degree and render it harder or more compact than any other, whereby alternating expansions and contractions thereof originate, in other words, the structure of the trachea.
3229. The whole body of the Respiratory animals becomes a trachea, a series of rings. The Respiratory are therefore the _Annulate animals_.
The rings are to be regarded as cysts, which are mutually juxtaposited or repeated, not being inserted within each other as in the preceding animals, but behind each other, so that the present circle consists of multiplied Cystic animals.
3230. The Annulate animals must represent the developmental stages of the tegument, while the viscera make a retrograde step--_Tactile animals_.
3231. The tegument has two functions: it is an organ of respiration and of general feeling or sensation. In both cases it passes through three stages of development.
It is either entirely a branchia, i. e. _reticular_ in character; or the branchiæ become individualized in certain situations and partly horny, constituting _branchiæ_ proper; or finally, the tegument is converted into an air-breathing organ, _tracheæ_ or _air-tubes_.
3232. It either feels with its whole surface, or by _papillæ_; or the branchiæ change into horny _feet_; or lastly, into alary appendages or _wings_. The Ancyliozoa consequently divide into Reticular or Papillary animals; into Branchial or Pedal, and into Tracheal or Winged animals.
3233. As the arterial character preponderates in these animals, so do the venous organs, especially the liver and renal structures, retrograde.
3234. The body is now an intestinal and tegumental body with predominating respiratory and sentient organs; here therefore sensitive papillæ, antennæ, feet and wings, appear in abundance.
3235. With the retrogression of the viscera, the glandular structure also and large proportional bulk of the sexual parts disappears. They assume the form of the intestine and tegument, i. e. become tubes.
The spawn like the milt consists, as a general rule, of only two long membranous tubes, running near the intestine.
3236. The external sexual parts belong to the sense of feeling and follow the developments of the tegument; they make their appearance as antennæ, and are usually accompanied by foot-like accessory organs.
3237. The nervous system in like manner accompanies the tegument. It consists of two gangliated cords passing along the ventral surface.
3238. As the maxillæ are only repeated feet, so they are rarely wanting, and are, like the latter, disposed after the fashion of scissor-blades. The same holds good of the antennæ; they are mostly in pairs, and consist of a series of rings projecting from the head.
3239. The eyes are usually present, but only as tegumental organs, or placed upon the points of the antennæ.
_Class 7. Reticular or Papillary Animals._
3240. The repetition of the Intestinal or Vascular animal, taking place under the dominion of the respiratory system, must be accompanied by the vascular system. The tegument is a vascular, a branchial membrane, a _branchial network_ or _skin_.
3241. A body with predominating tegumental and intestinal system, is cylindrical. It can only become gross or lump-shaped, if the glandular viscera, such as liver, ovarium and testes, get the upper hand or prevail.
3242. A tegument which appears as a branchial membrane, is annulate or ringed.
3243. An animal, having an annulate respiratory membrane without annulated feet, is a _Worm_.
Upon the lowest stage the vascular system is still similar to that of the Vascular animals; its blood is a colourless fluid--_White Worms, Entozoa_.
In order to be a Worm, it is sufficient to have an annulate tegument. Even if the intestine be wanting and the tegument supplies its place; the character is nevertheless complete. Here it is the skin, which digests, while in the Intestinal animals it is the intestine, which digests and breathes. Since it is here the tegument, which undertakes the offices of the whole body; the vessels, liver, and salivary glands are wanting unto the intestine.
The sexual parts are also frequently arrested and the ova appear to originate in the tegument.
Many undergo division without injury to themselves; yet the detached portions do not become again entire animals.
In most of them, however, the sexual parts of both sexes are present, being associated with, though separate and distinct from, each other; those of the female mostly opening upon the sides of the body.
As being Dermatozoa, they have for the most part special organs of sensation around the mouth, under the form of papillæ, filaments, spines and cups. The nervous system, wherever it is to be met with, consists of a ring surrounding the pharynx, and of a double ganglionic cord lying along the ventral surface of the body.
3244. Upon a higher stage of development the arterial system gains the preponderance, and the blood becomes mostly of a red colour--_Red-blooded Worms_.
3245. With the arterial the fibrous system also makes a more decisive appearance. The tegument is a fibrous tunic--it is itself an artery. All Annulate animals with a fibrous tunic, which can consequently contract, belong to this class, whether too they have red blood or not, like the Holothuriæ and Star-fishes.
3246. The Lumbricales or Earth-worms and the Hirudines or Leeches respire obviously through the whole tegument, though a special organ of respiration does begin to be evolved, in the first family in the "saddle," in the second in the lateral vesicles or cysts.
3247. In others the branchial vessels make their appearance above the level of the tegument as filaments or ramules, and are arranged in two rows, as in the common Sea-worm and in the Nereides.
3248. Lastly, they accumulate about the neck or head, as in the Amphitrites and Serpulariæ.
3249. There are also Worms which respire only through the intestine, its vascular system being bathed on all sides by water, as in the Aphroditæ. This water becomes, as it would appear, simply imbibed by the tegument in Thalassema, but through an opening at the anal extremity of the body in the Holothuriæ.
3250. These Worms have also no liver, or at the very best only traces of that organ, by reason of the preponderance of the arteries.
3251. Papillæ or sensitive filaments gradually sprout from the external branchial filaments, lie along the sides of the body, and are the prelude of the feet; as in the Nereides--_Papillary, Filamentary animals_.
3252. Others of these filaments become horny and appear as setæ or bristles, placed somewhat similarly to the above, as in the Earth-worm.
3253. Mouth and head are more perfectly developed than in the Entozoa. The first can readily expand and contract, frequently protrude the pharynx like a proboscis or tube, and not unfrequently has manducatory pincers, like as in Insects.
3254. On the head there are mostly annulate tentacula with muscular fibres, and frequently simple eyes.
3255. In those species which draw the water itself into the body and use it for the purposes of respiration, the formation of the mouth has risen higher, and the pharynx been provided with maxillæ, in number five or ten--Echini, Holothuriæ. These maxillæ form in themselves a peculiar skeleton around the pharynx, which is ranged circularly instead of by pairs.
3256. The nervous system is directed according to the relations of the tegument and intestine. It forms two ganglionic cords running along the ventral side of the body, and, where it makes with the nerves going to the maxillæ a ring around the pharynx, corresponds to the pharyngeal or pneumogastric nerves.
3257. In reference to the sense of feeling it may be said, that the Worms were those among the Sensitive animals, which feel by the whole tegument or body. Their body itself is a tentaculum.
3258. The sexual parts are likewise intestini-and dermiform, not glandular in figure like the ovarium and testis of Mussels and Snails, but tubular, as in the Entozoa. There are usually two oviducts and two seminal tubes to be found.
3259. So far as we are acquainted with the Red-blooded worms, they are androgynous, at least as regards the Earth-worms and Leeches, and their sexual parts are indeed tolerably symmetrical; yet they do not open posteriorly, but far forwards on the ventral side of the body, this being the case with even the male sex-organs.
_Class 8. Branchial, Pedal Animals._
3260. The representation of the Cutaneous animals is not yet the entire completion of the tegument. It attains a higher grade by separating in accordance with the visceral systems, and in the conversion of the branchiæ into sensorial organs or feet.
The branchiæ endeavour to separate self-substantially, and form a body of their own. The branchial body is the thorax.
By this attempt, the abdomen originates "per se" as the body of the intestinal and sexual system. The branchiæ upon it are arrested or stunted. In like manner the body of the nerves separates itself from the thorax and becomes head.
But meanwhile this separation is only imperfect; the head and thorax, if otherwise distinguished by their size and the feet, are for the most part connate; abdomen and tail are usually fused together, and only to be recognized as distinct by the limbs or members.
With this greater division of the body each division of it obtains its articulation into rings; but they will be hardened by virtue of the separation of the branchial network, or will become horny--_Horny animals_.
3261. With the conversion into horny texture of the body, its branchial filaments or tentacula must undergo a similar change, and divide likewise into rings like the general or bodily tegument.
Annulate, horny branchial filaments, are feet--_Pedal animals_.
A part of the branchiæ remains hanging to the feet, mostly to their basal joints, and is supported by animal motor members or limbs; the respiration becomes voluntary--_Branchial animals_.
The Annulate animals, having branchiæ and feet, are the _Crustacea_ or _Crabs_.
The branchiæ and feet are, of course, most largely developed upon the thorax.
Those upon the abdomen continue small, and those upon the tail or sexual body dwindle in size and change into other organs, such as fins, sacs, filaments, pincers.
3262. The number five of the branchiæ is also exhibited in the feet. There are mostly five large pairs of feet on the thorax, and as many small ones on the abdomen, as in the Crabs.
On the tail they appear more dwindled in size, and are frequently reduced to a smaller number.
Each part of the body has properly five rings; and these parts being the thorax, abdomen and tail, there are thus fifteen rings in all.
In the Crabs it is distinctly demonstrated that the maxillæ are nothing else but feet; there are therefore for the most part, after taking into account sundry arrests and coalescences, five pairs of maxillæ also.
The Crabs have usually two pairs of antennæ, whose signification is unknown; one pair is probably the elongation of the ear.
All Branchial animals have eyes, frequently too supported upon articulated pedicles.
They have a double nervous cord upon the ventral surface of the body.
They have, like the Mussels, a heart upon the dorsal aspect, and arteries with veins.
The intestine opens into the apex of the tail, and is surrounded by a liver.
The sexual parts still open for the most part on the thorax, and indeed by two orifices, as in the Worms.
There are no more androgynous or bisexual beings in the present class.
In some few the branchiæ already enter the body and become air-tubes, as in the Scolopendræ, Spiders and Scorpions.
The Branchial animals repeat, as being the second class of their circle, the Corals and Snails; their "quasi" coat of mail is therefore harder, often richer in calcareous ingredients, and, in addition to this, lies frequently upon the thorax or branchia as a special testa, called the scute or shield.
_Class 9. Tracheal, Alary Animals._
Annulate animals, whose branchiæ have undergone a partial conversion into tracheæ and into wings, are the _Insecta_ proper or _Flies_.
3263. A Worm with feet, tracheæ and wings, is an _Entomon_ or _Insect_.
3264. The first separation takes place in respect to the three tegumental segments of the body, the abdomen, thorax, and head. All three are, in the Insects, more separated from each other than in the Branchial animals, and united together usually by a narrow tube; even in cases also, where they are connate with each other, they are still easily recognized by respect being had to size, form, or appendages.
Every Insect is divided into three segments. In the abdomen are the organs belonging to the Worm, such as the intestine, and a fatty body which appears to be an analogue of the liver, a dorsal vessel, tubular sexual parts and air-tubes (tracheæ), but nothing else.
The abdominal feet now disappear entirely, and even the number of thoracic feet diminishes, owing, doubtless, to the production of wings.
3265. The thorax alone is reserved for the limbs or locomotive members. It never carries more than three pairs of legs and two pairs of wings. Of the viscera, it contains nothing but the oesophagus, while in the Crabs, important intestinal organs and even the liver, are situated within its cavity. It is in Insects therefore nothing else than a medium of support to the respiratory organs which have become limbs. Hence the thorax never has more than three rings, namely, one for each pair of legs. The wings invariably stand upon its two posterior rings.
3266. Since the limbs of the Insect are only the lateral filaments of the Worms, which have become hard and consequently hollow; they are as such not to be termed true feet, but are only to be compared with branchial arches or ribs; a step by which their greater number also admits of being understood. They are not to be compared with our feet, but unto the toes, which have been separated from each other as far as the rings of the body. The Crab has properly five thoracic and five abdominal toes. All its thoracic feet taken together are only equivalent to our hand.
The feet of the higher animals are only Insect-feet connate or coalesced.
3267. In other respects they already typify or prefigure true limbs, as well from their position as by the division of their joints. A perfect beetle's foot divides exactly like the limb of Man, into femur, patella, tibia, tarsus, and phalanges. These parts of the leg must not, however, be so absurdly divided and named, as has hitherto been unfortunately the case in our systems, where the femur has been called coxa, the patella trochanter, the tibia femur, while the toes have been lumped under the name of tarsus. (Ed. 1st, 1811. § 3087.) The regular number of the toes or tarsal joints is five, so that they correspond to our digital phalanges, to the metacarpal, and the anterior carpal bone.
3268. The wings are the branchiæ of the Mollusc that have been set free; they are placed therefore upon the back and are four in number. In many Insects there is still a pair of wing-like scales in front of the four wings, as in some Lepidoptera. They perhaps correspond to the shells of Molluscs, are branchial opercula.
It is only from this point of view that the structure of Insects admits of being fully understood; and apart from this it is absolutely devoid of all analogy. Thus it is possible for only six legs to take their origin from the thorax in the direction downwards, and nevertheless for there still to be wings upon the dorsum or back. The wings of Birds are by no means homologous to the alar appendages of Insects; they are, as is well known, the anterior members themselves, and there is no longer therefore in the Bird any feet attached to the thorax below, as in Insects. Besides, if the wings did not signify arms in the Bird, then it must have four legs. Thus in the Insect the wings could not also mean feet.
Their structure also speaks in favour of our view of the Insect's wings. They are known to be completely traversed by respiratory tubes, are true, or only desiccated, branchiæ--aerial gills. (Ed. 1st, 1811. § 3088.) Wings and feet are dependent from the same ring of the body, and are thus like the branchiæ and feet of the Crabs. Let the Crab's branchiæ elongate and dry, and they will thus be wings.
3269. Since the wings are newly eliminated organs of the sense of feeling, so are they here characteristic of the animal, and are consequently of greater importance for the purposes of division and arrangement than the organs of the head, which in all the lower animals is only an apparent head, and cannot therefore serve to characterize groups, &c.
3270. That the tracheæ in Insects have been developed out of the branchiæ by saccular inversion, is a fact displayed in a particularly distinct manner by the Scorpions and Spiders, who still possess at bottom internal branchial laminæ, unto which, however, air instead of water finds its way. It may be said, that with the general conversion into horny texture, the arteries of the Mussels were transmuted into internal tracheæ, and the branchial flabellæ into external organs. Would we interpret the body of the Insect in a strictly philosophic sense, the parts must then receive very different names to what they now bear. Properly speaking, in our own species the thorax has no limbs, but only the neck. The upper limbs are not lungs, but branchial organs, and it is the cervical vertebræ, from between which nerves are sent off to the arms, for it is just upon the neck also that the branchiæ have been left. What is termed therefore the thorax in Insects, would be properly their neck. Their abdomen would consist accordingly of thorax and abdomen, and it is this also, which carries on the principal share of the respiratory process. It therefore consists generally of ten rings, and has ten pairs of spiracula, namely, twice five, for both thorax and abdomen.
Or again, the abdomen may be regarded as an intestinal and sexual cavity, and the thorax may be left with its own name. In that case there would be five spiracles for the sex, five for the intestine, and perhaps only two for the thorax.
Would we proceed yet further; the head can then only be viewed as pharynx, and consequently as neck.
3271. In most Branchial animals that live in the water, a perfect circulation is present, because, by reason of their feeble amount of respiration, all the blood is not consumed. This is also the case in the young of air-breathing Insects, so long as they may have to grow. But when they have ceased to do this, so strong an amount of tension emerges in the circulation, owing to the increased respiration of air, that blood remains but in scanty quantity to be carried back by the veins, and the arteries now for the most part convey the air in a pure state, namely, uncombined with blood, as in the higher animals.
3272. As the air-tubes pass to all parts of the body, like the arteries whose place they now supply, so does the nutritive juice become everywhere oxydized and converted into parenchyma or tissue. The nutritive juice doubtless transudes at once through the intestine and penetrates to all parts, as in the Plants.
3273. Of the vascular system nothing at last remains but the dorsal vessel, whose ramifications appear to disappear entirely. According to its analogy with the Crabs, Scorpions, and Spiders, it is the aorta. It appears, as if in Insects the circulation dies off from the living body.
The whole Insect is an air-organ, an aero-vascular stem. All the organs respire directly, such as the intestine, the motor fibres, nerves, sexual parts, and wings. There is no part unto which tracheæ do not pass, and in this respect completely resemble the arteries of other animals.
3274. The intestine is invariably furnished with an anus, and that too situated quite posteriorly. It is usually expanded into several stomachs and has appendages, almost as in Fishes, which virtually correspond to the abdomino-salivary or pancreatic gland.
3275. The salivary ducts open into the mouth, and thus as in the Snails and Kracken, whose similar structures they repeat.
3276. Whether what have been called the biliary vessels are what the name implies, and really convey bile from the fatty body to the intestine, or whether they are lacteal vessels and discharge their fluid near the rectum into the dorsal vessel, does not yet admit of being determined. They have been thought to constitute an urinary apparatus; only certain cysts which occur in connexion with the sexual parts, appear rather to correspond to the latter.
3277. The nervous system consists, as in the Worms, of two ganglionic ventral chords.
3278. In the head, the feet again, and probably also wings, undergo repetition. This repetition is nowhere so distinct as in Insects. He who can still entertain any doubt of the maxillæ being arms, let him resolve to descend into the Insect-world, and he will become in place of a sceptic, a believer. (Ed. 1st, 1811. § 3095.) What have been called manducatory pincers move outwards as well as the legs, and seize like arms, are only arms. Their gripe takes place in the lateral or horizontal direction, and resembles that of scissors or shears.
3279. Where, moreover, there are three pairs of legs on the thorax, we find also three pairs of maxillæ, namely, an upper and lower maxilla, and a labium or inferior lip, which consists of the same parts as the maxillæ, only they are united by tegument.
3280. Where there are five or more pairs of legs on the thorax, as in the Crabs, there also are found as many pairs of maxillæ.
3281. Upon these maxillæ, palpi are likewise situated, which are probably nothing else than what has been termed tarsus, thus repeated in the head also. They are only arrested upon the upper maxillæ or mandibles. (Ed. 1st, 1811. § 3096.)
3282. The wings appear to be repeated upon the head as antennæ. Thus the head also is in the Insects a perfect trunk.
3283. Upon the head there is nothing more than the eyes. They have also become horny in texture, whereas before they were completely membranous, as in the Snails.
The eyes have been subordinated to the sense of feeling; they are nervous papillæ placed beneath a transparent tegument upon the apex of a tentaculum.
Insects therefore have a number of eyes; they stand either separate, as in the Worms, and are then called simple eyes, or they are crowded together, constituting compound eyes.
3284. Of the other sensorial organs a papilliform elevation in the pharynx is frequently exhibited as a tongue.
3285. Ears are found in the Crabs at the root or base of the antennæ, or in them there is only a tympanic cavity with an ossicle inclosed within; in the Insect there is nothing of the same kind. But meanwhile, since many species attract each other by means of sounds, they must thus possess the faculty of hearing. The antennæ therefore probably correspond to the auditory conch. The auditory conch of the Mammalia ranks also in the signification of the hand, and thus of the organ of feeling.
Nevertheless the antennæ may be transformations of the wings. The auditory ossicles are indeed members which have originated from branchiæ. Probably the antennæ are the auditory ossicles themselves which have emerged outwards, as in the Fishes and Reptiles.
There is no trace of a nose; nor can there be any, for it is the anterior orifice of the vertebral canal, and this is wanting in all lower animals. A noseless animal is an inferior or Tegumental animal.
3286. The sexes are separate, because they are Air-breathing animals and perfectly symmetrical in form. The ovi-and seminal ducts are likewise symmetric and in pairs. The ovipositors and penes are perfect, as in the Snails, because they can be shoved forwards, but not everted as in the latter.
3287. The position of the sexual parts at the posterior extremity of the body has been permanently established, with exceedingly few exceptions, as in some aquatic Insects.
That they correspond to the head, or are imperfect cephalic organs, is most distinctly shown in the Insect. They are usually environed by valves, pincers, and filaments, which resemble maxillæ and palpi.
3288. The Ovum-animals multiply by division, by granules or gemmæ, i. e. shoots, the Sexual animals by membranous ova, the Arthritic animals by horny ova. These last, or egg-shells, are hard, and frequently also so strung together or laid by the parent upon each other as to represent the annulate body of some Insect.
3289. In the ova of the Dermatozoa or Tegumental animals the vitellus only appears to exist, being without albumen, which first seems to make its appearance along with the animal systems. As the animal separates into higher and lower substances, so also does the ovum or microzoon. The simplicity of the Tegumental animals has been prophesied or foretold in that of their ova.
The Insecta, as being the third class of their circle, repeat the Acalephæ and Kracken. With the last they bear much resemblance in form and in their motor organs.
3290. Would we arrange the Worms along with the spiral vessels or ducts, the Crabs with the woody rings in the stem, we must term the Tracheal animals foliage or leaves. Their wings are pinnate leaves, and among the Orthoptera many occur which, as well in the form of the body as of the wings, appear as if they had been just liberated from the Papilionaceous tree. The tales or stories, about leaves changing in the torrid zones into Insects, are not without meaning; for poetry is none other than the Ideal of natural history.
_Metamorphosis._
3291. We now proceed to retrace our steps. The Insect is a Tegumental animal represented in limbs. There can be therefore present in the Insect no other development than in the Cutaneous animal, which works itself up into the Branchial and Tracheal animal.
The Insect passes through three stages prior to its attaining the adult or perfect condition. It is at first Worm, next Crab, then a perfect, volant animal with limbs, a Fly.
3292. The representative passage of the Insect through the preceding classes in the course of development, constitutes its _metamorphosis_. Thus the Insect's metamorphosis obtains a meaning and an explanation. Upon the whole, the history of every kind of pregnancy is none other than the passage which takes place through all the animal classes, as I have first represented in my book upon "Generation" (Von der Zeugung, 1805); but in no class of animals are the periods so dissevered or drawn apart as in Insects. It therefore comes to pass that these animals are the equivalent transcripts or copies of a system common to them and the preceding animals.
3293. The Worm is the first condition of the Insect. It is represented, by the larva, which, according to its diversity of size, is called maggot, caterpillar, grub, &c.
3294. The larva is only a Cutaneous and Intestinal animal. It knows of nothing else but eating like the Worm; it has no sexual function, no lust, nor pain; it can scarcely move; in many the feet are wanting, as in the larvæ of Flies, which thus resemble the Entozoa; many have a crop of lateral papillæ, like the caterpillars, which resemble the Nereids.
3295. The change into a nympha, chrysalis, or pupa, commences with the horny induration met with in the Crab, and in the higher organized kinds of Snail. The pupa is the embryonic Crab or its antetype, it is the Snail in its shell.
3296. In the third condition the Insect makes an advance above the branchial condition, and casts aside the Snail's or Crab's shell; and is then the perfect Insect, the _Fly_.
3297. The metamorphosis is accordingly the embryonic transition of the Insect, after extrusion from the egg, through the three classes of its circle. In the ovum state it only passes through that of the Intestinal and Sexual animals.
3298. This is a retrospective proof that the higher animals also pass in the ovum through the condition of the lower animals, but after birth through the classes which directly precede them.
What holds good of the Insects, does so also of the preceding groups of animals, although in a less degree; the higher class of each circle still passes after birth through one or other of the inferior classes.
Thus, the Acalephæ first appear under the form of Infusoria; next change into Polyps, and then obtain for the first time the form of the perfect Acalephæ. The same phenomenon occurs in the naked snails or Slugs. They have, when freshly hatched from the egg, a small mussel-like shell, which they lose at a later period.
The Echini or Sea-urchins, which I believe ought to be ranged with the Kracken, also pass through a metamorphosis.
Finally, it is well known that the Crabs also are subjected to a metamorphosis. Their first condition must be regarded as that of the Worm.
The law is thus universal. The second and third class of every circle traverse after birth the classes, to whose series they belong.
It would even appear that the classes, also of a lower circle, are subject to similar changes; at least many Entozoa, and thus the first class of the third circle, appear to undergo such a metamorphosis, since they first of all resemble an Infusory animal, next change as pupæ into Molluscs, and are then manifested for the first time as Worms.
3299. The Dermatozoa or Sensitive animals range in the following manner, according to the anatomical systems:
A. INTESTINAL ANIMALS. B. VASCULAR ANIMALS. C. RESPIRATORY ANIMALS.
1. _Gastric animals._ 4. _Venous animals._ 7. _Reticular animals._ Infusoria. Mussels. Worms. 2. _Intestinal animals._ 5. _Arterial animals._ 8. _Branchial animals._ Polyps. Snails. Crustacea. 3. _Absorbent animals._ 6. _Cardiac animals._ 9. _Tracheal animals._ Acalephæ. Kracken. Insects.
3300. According to the developments of the feeling-sense they stand thus:
A. OVUM-ANIMALS. B. SEXUAL ANIMALS. C. CUTANEOUS ANIMALS.
1. _Vitelline animals._ 4. _Ovarial animals._ 7. _Papillary animals._ Infusoria. Mussels. Worms. 2. _Albuminose animals._ 5. _Orchitic animals._ 8. _Pedal animals._ Polyps. Snails. Crustacea. 3. _Involucral animals._ 6. _Renal animals._ 9. _Alary animals._ Acalephæ. Kracken. Flies.
3301. From these tables the number and kind of relationships or affinities is readily deduced. There is a _Relationship of Proximity_, as that between Infusoria, Polyps, and Acalephæ.
A _Relationship of Repetition_, as of Infusoria, Mussels, Worms. There is also a _Relationship of Series_, which takes its rise from the parallelism of the classes. The two last relationships have been confounded together under the term _analogy_, which at best is but a word of random definition.
_Second Province--Sarcozoa, Cephalozoa._
FOURTH CIRCLE--SARCOSE ANIMALS.
3302. The system of motion and sensation, which broke forth so forcibly in the tegument, now passes over into other forms, into the globular form of bone, the fibrous form of flesh or muscle, and the point-form of the nerves.
3303. The nervous system when liberated from the vegetative organs is, or consists of, the myelon (spinal chord) and the encephalon, i. e. brain.
3304. These animals are therefore Osseous, Muscular, and Encephalic animals, whereupon the senses are proportionately developed, and constitute the basis of the highest animal forms--_Sarcozoa_. The Dermatozoa are _asarcose_ or _fleshless_ animals.
3305. With the sudden appearance of the animal systems all organs of the head, such as tongue, nose, ears, and eyes, are also developed; no asarcous animal has nostrils. Now these parts, especially the nostrils, as marking the terminal extremity of the vertebral column, make up the head--_Cephalozoa_, or Cephalic animals.
_Class 10. Osteozoa, Glossozoa._
3306. The animals, in which the osseous system for the first time makes its appearance are, the _Fishes_.
The Fish alone has more bones than any other animal. It has dorsal rays, which are wanting in all others.
3307. The animal systems are for the first time slightly separated from each other. Bone, muscle, and nerve are rather a gelatinous mass, which only bids fair for becoming something higher; on this account the bones are frequently but soft cartilages or tendons, the muscle composed of white fibres like those upon the intestine or vessels, the nerve thick, oleaginous, and soft in texture, while the brain is in its constituent parts hardly comparable with that of the Thricozoa.
3308. The muscles of Fishes are not yet perfect in character, since they are devoid of individualization and red colour, and their fibres run mostly parallel with each other without uniting into tendons. Their muscular body is a muscular wall.
3309. The Fishes, ranking upon the first stage of the Sarcozoa, repeat the same stage of the preceding circle, and thus the Infusoria, Mussels, and Worms; or the stomach, veins, and branchial rete or network; furthermore, the vitellus, ovary, and skin, which systems must in them accordingly predominate.
_Pelvis._
3310. In Fishes the pelvic organs, sexual parts, and tail predominate. The tail, as being an appurtenance of the pelvis, is larger and stronger than in other animals. It mostly makes up the largest part of the body, and is, properly speaking, its only motor organ.
3311. The sexual parts have still the form of the intestine, and occupy the greatest part of the abdomen. The ovaria are two sacs, like two Infusoria, in whose parietes granules are developed; even the testes are only two such sacs or seminal vesicles wherein the "milt" is contained. Fishes may be termed anorchitic, or animals devoid of testes.
The ova are small, consisting of spawn or roe without shell, but they separate into albumen and vitellus.
3312. External sexual parts are not present. Everything usually opens into a cloaca, which is thus here a true pharyngeal cavity.
3313. With the perfected formation of the head, the animal, so to speak, undergoes a sudden and entire change, and the sexual parts are developed with all their accessory organs, such as the sexual lung. Kidneys are present and mostly an urinary cyst. The kidneys are indeed still so amorphous and soft, that they more resemble coagulated blood than an organ, yet meanwhile agree in this respect with the 'milt' and branchial substance.
_Abdomen._
The abdomen is not simply confluent with the sexual body, but thrust by it completely forwards. The anus mostly lies in front of the middle of the body.
3314. The intestine separates for the first time distinctly into cephalic and sexual, or small and large, intestine.
3315. The vitelline canal is the coecum. This is very distinct in the Sharks and Rays.
3316. In Birds therefore the vitelline canal is the coecum also, and the two coeca, falsely so called, which are placed upon the side of the rectum cannot represent the coecum coli, or else the Bird must have three of these blind appendages or sacs.
3317. The spleen here appears for the first time; while the pancreas divides into a number of coecal appendages.
_Thorax._
3318. The thorax of the Fish is reduced within very narrow limits; it is like the first thoracic formation, and is thus a Mussel's thorax. The branchiæ with their opercula are similarly formed to the branchial plates and shells of the Bivalve Mollusca. The thorax is therefore attached only externally to the body, and the Fish is to be viewed as a Mussel, from between whose shells a monstrous abdomen has grown out.
3319. But this Molluscan thorax is conjoined with animal systems, and has assumed their noble or elevated type of structure. Here therefore the osseous and sarcose system blend together, and the higher formation of the thorax emerges into view.
3320. For the first time an advance is made towards the formation of a trachea, namely, by the branchial framework or skeleton, which opens into the mouth, and corresponds therefore properly to the larynx. Fishes are therefore the first animals which breathe through the mouth. In all the preceding classes the air entered the body, or the water found access to the branchiæ, by other routes.
3321. They may be termed Mouth-breathing animals, for the first formation of the trachea extends no further than to its junction or communication with the mouth; since, for it to be continued into the head and open self-substantially as a nasal organ, constitutes a second step in advance, which cannot be ventured upon in an Abdominal animal. In Fishes everything relates to the abdomen, and this is expressed by the first union of the trachea having taken place with the oesophagus or the mouth.
3322. But, at the same time that the trachea is inserted within the mouth, it opens externally and in the lateral direction upon the body, thus letting the water escape from it posteriorly after a mussel-like or sexual fashion. Thus, the trachea is not yet closed inferiorly, nor hence also the thoracic cavity. Between the head and abdomen there are still openings--branchial foramina.
3323. The trachea is itself, however, a thorax upon a small scale, consisting as it does of rings, or as it were ribs. These arches are not yet united by muscles with each other, and the water flows out between them into the apparent thoracic cavity, from which it next escapes beneath the branchial operculum.
3324. These tracheal rings are the branchial arches. The branchial vessels are tracheal and by no means pulmonary vessels.
3325. Thus, upon taking a retrospective glance at the matter, we may fairly conclude that the branchiæ of the Dermatozoa are not equivalent to the lungs of the higher animals, but are only the antetypes of the bronchi, being thus cervical organs.
3326. The trachea is thus formed prior or antecedently to the lungs, but is still membranous and devoid of any continuous connexion between its rings.
3327. The lung is an organ extraneous or foreign to the trachea, and becomes only as if accidentally associated with it.
3328. But a Sarcozoon is not devoid of lungs nor of aerial respiration; for, being the totality of all lower animals, it consequently unites in itself the respiratory apparatus of the Branchial and Tracheal animals; thus gills and lungs.
3329. In the Fish the first lung appears, that is to say, if we designate or interpret the respiratory organ in Insects only as tracheæ, which do not open into the mouth.
3330. The Fish's lung is the air-or swim-cyst.
3331. The lung is still subordinate to the abdomen and intestine. It is therefore still separated from the trachea or the branchial arches.
3332. Fishes swallow the air and drive it into the pulmonic cyst, where it is analysed or decomposed.
3333. This lung, so soon as it stands opposite to and enters into conflict with a branchia, is what serves to direct or control the action of the heart. It is only when the swim-cyst is allowed to hold good as lung, that the circulation in Fishes admits of being understood. For otherwise venous blood must flow into the heart, from this into the branchiæ, from these directly into the aorta, and so to the different organs without again entering a cardiac cavity, a structure which occurs in no other class of animals, but rather in every instance its reverse. The first heart is arteriose, not venous in all animals, even in the Mussels and Snails and in the embryos. Now, the Fish is still such an embryo, and has only an arteriose heart.
3334. Thus, the following is the "modus operandi" of the parts in question; the swim-cyst is the lung, in which, seeing that it contains air, the blood is oxydized; this oxydized blood flows into the heart and renders it _arteriose_ in character, despite the concomitant influx of venous blood into that organ. Upon this, the blood passes out of the heart through a true aorta, which is called the branchial artery. Instead then of this aorta giving off only some twigs as bronchial vessels to the gills, and next proceeding to pursue its course as a main trunk through the body and along the back, it passes wholly to the branchiæ, i. e. it entirely becomes a bronchial vessel, is slightly oxydized, and then returns upon itself to form the aorta, which should have taken its course directly from the heart.
_Senses._
3335. In accordance with the thorax, the corium or true skin is developed. It is mucous in character and muco-secernent like the intestine, from its being constantly in the water. The whole skin is undermined by mucous canals and perforated by their excretory apertures. These foramina of the lateral line are arrested, and metamorphosed branchial foramina, which have only retained the evaporative function of respiration.
3336. The remnants of the tegument's annulation are the scales. They are desiccated air-branchiæ, alary opercula or elytra, and consequently indicate that feature of the Insect, which has continued to work its way into the class of Fishes.
3337. Like the skin, so are the limbs--tegumental members. What is flesh and bone upon them, has kept quite close to the body, and that only which is designed for splitting into digits projects, constituting tegumental digits with cartilages--_fins_. These fins are something better than the lateral papillæ of the Worms, are furnished at their base with joints, and are in number only four, but crippled or stunted in every possible way.
3338. The fin-rays do not correspond to the digits, but to the nails. They are fibrillated nails, like the wing-feathers of Birds.
3339. Lastly, the head possesses all the organs of sense which belong to a head, but they are still far removed from their perfect condition.
3340. As the nervous system is the first mass, from which the remaining ones have been set free, so also is the nervous sense the first after that of the tegument, which appears as a whole, and serves as a pattern for the subsequent senses. The eye is the sense which is first developed most perfectly, not directly as regards its own completeness, but when considered in reference to the others.
As the sense of feeling has been at once manifested in the vegetative animals as a peripheric nervous sense; so antithetically or _vice versâ_ it is in the higher animals the light-sense, as being the central nervose sense, which appears.
3341. Hitherto both these senses stood altogether alone with each other upon the stage of animal existence, in order, as it were, to play their parts in mutual consent; but so soon as the sense of feeling became individualized into tactile organs, then also the eye made its appearance in a detached or isolated manner.
3342. The Fish's eye is upon the whole composed of the same structures as in the Mammalia; but it is devoid of motion and external coverings.
3343. The ear, as being the sense of motion, has scarcely withdrawn itself from the brain, has not yet become a true external organ, and that portion of it which appears externally, is subservient to inferior systems, such as the branchiæ or gills.
3344. The external auditory organ has become confluent with the branchial aperture, and the auditory ossicles have become parts of the branchial operculum.
3345. In the internal ear only the three semicircular canals have been left. The cochlea is not yet developed.
As true palpebræ or eyelids are wanting to the visual organ, so also are the conchs of the external ear.
3346. The nostrils exist, because a vertebral canal, which terminates in them, is present; stoutly-developed olfactory nerves are also present, so that the sense of smell cannot be wanting. Only this sense has not yet taken the respiratory organ into communication with itself, and both therefore live for themselves in an arrested or stunted condition. This nasal organ does not open into the mouth, admits of neither water nor air passing through it, and therefore serves not as a test-organ to the respiratory process. This is my main point for distinction of Fishes from Reptiles.
3347. Every Sarcozoon, whose nostrils do not open into the mouth, is a Fish. The Siren therefore does not belong to the present class, while the Lepidosiren, having an imperforate nasal organ, does.
3348. The tongue has remained rather an organ of touch and deglutition, than an instrument of taste. The salivary glands are scarcely developed.
3349. But the tongue here appears _for the first time_ in the animal kingdom as a perfect organ provided with muscles, and a lingual bone, as in Man; Fishes are, as regards the development of this sense the _Glossozoa_, Lingual or Tongue-animals.
3350. As Fishes are the repetition of the intestine and vitellus, so they may be termed _abdominal_ or rather _Pelvic animals_. They are an abdomen, unto which are appended branchiæ, fins and head.
They are the repetition of the Infusoria, Mussels, and Worms; in them we meet with mucus, branchial opercula, articulated filaments, and labial cirrhi or palpal organs.
_Class 11. Myozoa, Rhinozoa._
3351. Those Sarcozoa which for the first time obtain true muscles and a perforated nasal organ are the _Reptilia_.
3352. True muscles are red, have a definitely circumscribed outline, and are divisible into head, fleshy portion or belly, and tail or tendon. As such they are found for the first time in Reptiles.
_Thorax._
3353. With the osseous system in Fishes the sexual abdomen was chiefly developed; with the muscular system therefore, as in the present class, the abdomen proper or intestinal abdomen becomes perfectly developed, and the thorax more spacious in calibre.
3354. The thorax is still blended with the abdomen. In Fishes the disposition was already stirring to create for themselves an air-organ; only this was imperfectly attained, since the swim-cyst communicated with the gullet, but not with the branchial arches or larynx, nor with the nasal organ, as in the present class.
3355. Now, if the swim-cyst be symmetrically developed, if it communicates with the branchial larynx and opens through the nose, the aerial respiration is completely attained, and shares its dominion over the body along with the digestive function.
3356. A Fish, respiring through the nose into two swim-bladders, is an Amphibious animal, a _Reptile_.
3357. In the Reptile, however, the thorax is still subordinated to the abdomen. Its lungs traverse the whole abdominal cavity, and its manner of breathing is still similar in character to the abdominal function. The lungs are simply two membranous bladders, like two intestines, and admit also of being voluntarily filled with air like the intestine is with water. This filling or inflation of the Reptile's lungs takes place also by the abdomen being expanded at the same time that the mouth is closed by the muscles of the fauces, and is therefore a true process of deglutition.
3358. Their inspiration is still therefore fish-like in character, excepting that the air is drawn in by the nostrils; the respiratory process, however, and the respiratory organ is constituted as in the perfect air-breathing animals.
3359. Reptiles may therefore be called _Abdominal animals_, while the Fishes are Pelvic animals.
3360. The metamorphosis of the branchial arches into larynx is placed beyond doubt in the Reptiles. The anterior branchial arches frequently unite with the lingual bone, whereby the latter obtains several cornua.
3361. The thyroid gland also makes its appearance here for the first time, while the branchial vessels separate from the arches. Fishes have therefore no thyroid gland.
3362. The circulation is complete. The venous and also the arterial blood enters the heart. But both kinds of blood are still mingled with each other as in Fishes. Yet already, through the direction and arrangement of the cardiac orifices, provision seems to be made for their separation.
3363. The rationale of this mixture of the blood appears to reside in the fact, that many of these animals, and probably all of them in the ovum state, breathe by branchiæ. (This proposition, which was announced in the 1st Edition, 1810. § 305, and there based upon the transitional stages traversed by the animal classes during embryonic development, has since then been raised to a state of certainty.)
3364. The mixture of the blood takes place through an opening in the septum separating the two cardiac ventricles, and which corresponds to the foramen ovale of the foetus. The heart of Reptiles is therefore a persistent foetal heart.
3365. Without doubt, however, the arterial blood only enters the left heart, while the venous remains in the right, to be thence driven into the lungs.
3366. In other respects the foramen ovale is not a true opening, but a bifurcation of the vena cava, one branch of which passes into the right, the other into the left ventricle, as I have demonstrated in the 'Beyträgen zur Anatomie' (Frankfurt bei Wesche), in treating of the calf's heart.
3367. There are generally no foramina in the body, but only fissures without orifices.
_Abdomen._
3368. The digestive organs are pretty similar to those in Fishes, as is exemplified by the intestine, liver, and spleen; the digestive process is, however, more energetic, and frequently aided by venom or poison.
3369. The sexual parts are perfect. The first true testes and ovaria have originated. The tegumental formation has consequently passed over at the extremities into a glandular, because the vascular system preponderates. Even in the highest Fishes, such as the Rays and Sharks, the testes and ovaria are still not developed to the extent which they are in Reptiles.
3370. In the female the oviducts pass together as complete cornua into an uterus. In the male the seminal ducts are still indeed separate, yet the two bodies or crura of the penis are frequently developed, and continue separate in the Serpents and Lizards, but are connate in the Tortoises.
3371. The kidneys are distinct and symmetrical; their ureters usually unite into a large urinary cyst. Thus, the sexual lung is also significantly developed.
3372. Many of these urinary bladders split, as in the Tortoises, into two cornua. This is the form under which the cyst again occurs in Birds, where its two cornua have been incorrectly termed coeca, but the urinary cyst rectum, because the intestine opens into it.
3373. In Reptiles also there is a cloaca or sexual orifice, into which all the sexual apertures meet together.
The ova of Reptiles possess a vitellus and albumen, and are surrounded by a membranous or sometimes by a calcareous shell.
3374. Hence Reptiles have originally also a coecum, but it has for the most part disappeared; like its antetype the vitelline canal, which is so small in most Fishes and Birds, as to be scarcely distinguishable.
The tail projects from the body, and is rather trailed after the animal, than enabled to assist the body in its motions, as in Fishes.
_Senses._
3375. Many Fishes have only a vertebral column without lateral bones; in the Thoracic animals this deficiency can hardly occur any longer, although we are not, on the other hand, to expect any great amount of perfection to be attained by these parts. Reptiles have osseous ramules, whether such be ribs or true feet; the Serpents have indeed only ribs, but these exist in great numbers; in the Frogs, on the contrary, ribs are wanting, but they have limbs.
In no Reptile do fin-rays occur; consequently, there are no peculiar bones as in Fishes.
3376. In Reptiles the limbs themselves are in a varied or undetermined state, although they constitute a class in which the formation of true muscular limbs is decided. In Reptiles the toes appear for the first time.
3377. As regards the tegument, it quite corresponds with that of the Corals, Snails, and Crabs, of which the Reptiles are a repetition. In some cases it is slimy and naked; in others Mussel-shells originate, as in the carapace of the Tortoises; the scales, scutes, and claws of Serpents and Lizards repeat the rings and scutes of Crabs.
3378. The head has separated itself more from the thorax, the eyes and ears being much more perfect; as regards the latter organs the cranium is always perforated, in the former there are eyelids.
3379. The eyes have still no power of free motion, and the eyelids are as yet very imperfect. It is principally the lower eyelid which closes, while in Mammalia the upper one is the most developed.
3380. The external auditory meatus is notwithstanding usually covered by integument, the auditory ossicles are mostly blended into one, which projects above the skull; the cochlea is wanting.
3381. Reptiles produce sounds for the first time through the lungs; they have voice, but as yet no song.
3382. The nose, however, gives the finishing touch to the head. It has not only opened as a vertebral canal in front, but also as a thoracic cavity posteriorly into the mouth, which was not the case in Fishes. The open nose is the æsthetic character of Reptiles, just as the red muscles are the characteristics of the anatomical systems. The Reptiles are _Rhinozoa_.
3383. The tongue takes a higher rank than in Fishes. It is throughout soft, fleshy, and smooth; but in most species is still slit into two, which reminds us of a similar condition of the penis.
3384. The teeth are in these animals more like digits than in Fishes. While here they consist for the greatest part of front teeth, and are therefore associated with the intestine, in Reptiles they are mostly lateral, and thus true maxillary teeth, which are annexed to the salivary system. With this special dental formation the saliva is also more active; it is a rapidly fatal poison.
3385. The poison-teeth or fangs have likewise a groove, which can be regarded as the continuation of the salivary duct.
_Class 12. Neurozoa, Otozoa._
3386. Those Sarcozoa, whose nervous system is for the first time perfectly developed, and whose ear is open externally, are the _Birds_.
3387. The completion of the nervous system is the brain; now the brain defines or determines the head; the Bird is, properly speaking, the first Encephalic animal.
3388. In the Bird the head has for the first time, and that indeed suddenly, freed itself from the trunk, and been placed upon a long neck, far removed from the thorax. In no class are there such long necks and so many cervical vertebræ to be found as in Birds. They can be therefore called also Cervical animals, as the neck is not simply present, in order to render the head independent or self-substantial.
The caudal vertebræ, on the contrary, are lessened throughout the Birds, to a degree met with in no other class of animals.
3389. The brain separates for the first time completely into cerebrum and cerebellum, begins to exhibit convolutions, and has in general most of the individual parts analogous to those in the Thricozoa. They are therefore _Encephalic animals_, which the name of Neurozoa will properly enough imply. The nerves are in comparison with the spinal cord much thinner than in Reptiles and Fishes.
In the Bird also, all the spiritual or mental faculties make their appearance for the first time and suddenly, whereas in the preceding classes but slight traces of them were observed. Such for example are their mechanical instincts, varied modes of nidification, powers of imitation, susceptibility to instruction, knowledge of their benefactors, sentiment of joy, wheedling or coaxing manners, and so on. We have no example of Fishes and Reptiles having learnt any artificial tricks.
3390. Birds are the closest repetition of the Insects, but stand remote from the Crabs and Acalephæ, indications of which are reflected by their structure, disposition, mechanical instincts, and in their nests.
_Thorax._
As the sexual body with the tail predominates in Fishes, and the abdomen in Reptiles, so does the thorax in Birds. The whole abdomen and sexual body has been subordinated to the thorax.
The ribs are here directed in such a manner for the first time, that the thorax can act as a voluntary pump-organ. Sternum and respiratory muscles are usually large and peculiarly built.
3391. The lung is only a cluster of Insect tracheæ. They are full of foramina, out of which the air can penetrate into the whole body, exactly as in Insects. In Fishes the lung was still an actual intestine, as seen in the single air-cyst; in Reptiles it was nothing more than a double intestine; in Birds this lung is divided after the fashion of an Insect into aerial vessels or ducts.
In the Bird the intestine lies in the air, and breathes from it, as in the Insect. Birds are also animals which breathe by the intestine.
3392. The whole Bird is lung. Its body is a thoracic cavity, while in the Fish it was simply a sexual cavity, and in the Reptile had obtained the abdominal form. In Birds therefore there is a number of ribs and a strong ossification, from the air itself penetrating into the bones.
Even the intestine has passed over into a motor organ. In Birds and Insects only do we find a true muscular stomach, wherein the food is crushed.
_Members or Limbs._
3393. In these animals the formation of limbs or members must be wholly attained. Everything, that works, must work to the production of limbs. The whole body becomes limb.
3394. With the perfected nervous and respiratory system the bones and muscles make also a perfect appearance. The skeleton is hard, complete, and full of air instead of marrow; the muscles are red and separated, the movements free and complex.
3395. The elevated condition of the motor system is demonstrated in the limbs, which here appear in the greatest diversity. There are only two pairs of limbs, thoracic, and abdominal or sexual members. These two are equivalent, so long only as the sexual and abdominal cavity belonging to the trunk proper predominate, and therefore preserve the equilibrium. But the limbs, which have been given to serve the office of the trunk, are destined for progression or swimming, and are simply terrestrial or aquatic members.
3396. The higher limbs are thoracic respiratory members, which are filled with air and clothed with tracheæ, thus exercising a function conformable to the thorax. The thorax has an aerial character. The highest thoracic limbs must be aerial members.
3397. As the abdominal members move upon the earth or in the water, so do the thoracic in the air. The terrestrial limbs are feet, the aerial limbs, _wings_.
3398. The wings are in the member-formation the extremes of the thoracic limbs. It does not follow from this that they are the noblest in rank, but only that they are the uttermost unto which bodily motion can attain.
3399. The winged animal is the Bird.
3400. The Bird is an Insect with fleshy limbs.
3401. The wings of Birds repeat the alary appendages of Insects in the flesh.
3402. A Bird's wing is a strange but very instructive composition. It consists, namely, of a Reptile's foot and an Insect's wings.
3403. We saw how the branchiæ of the Insect dried up, separated from the feet, and being liberated as wings, were permeated by tracheæ. In the Bird these wings have remained standing upon the feet and been converted into feathers.
3404. A feather is an Insect's wing.
3405. As the Bird grows out at the thoracic limbs into the Insect's wings, so does it upon the whole body into dried branchial laminæ.
The whole body of the Bird is clothed with branchial laminæ or plates.
3406. The wings of the Insects might be called free tracheæ.
The Bird's feathers are Insect-tracheæ. As in the Insect the wings are a leash or net of tracheæ held together by membranes, so feathers are tracheæ dividing into fibre-like ramules.
3407. The Bird is a Reptilian or Frog's body, beset all over with Insects, like as with parasitic animals.
3408. The highest Insect only attains to the possession of four wings, which in some Moths split again into several feathers. In the Bird a vast number of such wings originates.
3409. An Insect's wing is not more than _one_ feather, and is therefore placed also directly upon the body. These wings must multiply so soon as they occur upon a membered trunk, upon arms. Thus, we need not ask why the Butterfly has four, but the Bird only two wings, seeing that the latter is the nobler animal. The discourse cannot be concerning the wings of the former, for the Butterfly has indeed none, but only four feathers.
3410. What the Bird is, it is by virtue of its feathers. It is throughout a trachea, a pair of bellows. Its bones are hollow, full of air, and stand likewise in communication with the lungs; the feather-quills are also hollow.
_Senses._
3411. The wings have all the muscles to themselves; the bone has in them gone to ruin. On the legs, on the contrary, the muscles have declined, and the bone has got the upper hand.
Hence it results, properly speaking, that only the thoracic members are perfected, because the Bird is nothing but thorax. The abdomen has, so to speak, vanished, and through this the abdominal limbs are left only, as thin and dry poles or staffs.
3412. From the same cause the muscular flesh upon the head has disappeared. Neck and head are lean, or like Insect horn, which serves only the nervous system.
3413. Beyond the excess of motion the sense of feeling has been nearly lost. The toes are simply destined for motion, and to be used as scrapers, and the digits have become the supports of feathers.
3414. The bill is an Insect's proboscis. In the Bird no teeth whatever project from the flesh, but the jaws themselves. To such an extent has the flesh been withdrawn. What has been called the cere is the only remnant of the facial flesh. Even nostrils and tongue have suffered ossification.
3415. The tongue is a feather. Saliva is scarcely present.
3416. The ears, as being the sense of motion, are far more perfectly evolved than in all the preceding classes. They have opened outwards, and possess an additional auditory part, the cochlea.
With the limbs the sense of hearing must of necessity grow perfect.
3417. The Bird is thoroughly, or "out-and-out" organized as an animal of song. In it Nature attains unto a definite hearing and speech. The Bird speaketh the language of Nature.
With the Bird, the voice, properly speaking, breaks forth for the first time, and that too in a high grade of perfection, as melody.
3418. The ear is the highest representation of the trachea in muscles and bones. The Bird is the _Otozoon_.
_Sexual Parts._
3419. The kidneys are symmetrically constructed, although not yet a perfectly coherent mass. They are of very large size. But a strange feature has come to pass in the urinary cyst, which is the sexual lung. Into this, as has been already said, the intestine opens, and thus it here again passes over into the lung, just as it has passed over into a fleshy cardioid stomach, and completely upwards into the feather-like tongue.
Into this sexual lung the seminal ducts also enter, or the penis, when one is present, along with the oviduct.
3420. The ovum consists of two completely separated substances, and these indeed are so distinct, that the vitellus is secreted at an entirely different place to the albumen, the former in the ovarium, the latter in the oviduct--whereas in Fishes both originated together.
The separation cannot extend further, or else the substances would no longer unite with each other, and the vitellus must be first mixed with the albumen, after it has been completely formed, after or when it is an embryo. In the Bird albumen and vitellus come together during their passage out of the parent's body, or in the act of being laid, and thus before the albumen has been converted into the chick. But in the Mammalia they are first completely united, subsequently to their being laid--i. e. during the period of lactation.
3421. The vitellus is directly secreted from the arteries, but the albumen from an enteroidal sac or the oviduct, which is finally converted into mammary glands.
3422. The vitellus is more a product of the thorax, and is therefore formed directly into the intestine and entire embryo.
3423. The albumen is an intestinal or digestive product, a solution of the organic mass into protoplasma or primary mucus. It is not fashioned itself into the embryo, but is only absorbed as fluid nutriment by the latter.
3424. The shell of the ovum is the last bone, which the animal deposits from the sexual blood, as being analogous to the urine. It is an aerial product, or an analogue of the feather's quill.
3425. Even the nest of the Bird is a spiritual repetition of its plumage, for in it the stalks of plants, tracheæ, or feathers are united into one body, which reminds us in the Swallows of the Acalephæ.
FIFTH CIRCLE. AISTHESEOZOA.
_Class_ 13. _Aistheseozoa._
3426. An animal with all its organs of sense perfectly developed, is a _Thricozoon_.
3427. The nervous system emerges at length freely above the other systems, and it is no longer its mass, but its _organs_ which impart character to the animal. The nervous organs are, however, simply the organs of sense. Through these therefore must the present differ from the preceding class of animals.
3428. Now too the senses first make their appearance in a self-substantial manner above the other organs, serving merely their own functions, and only by chance those of others.
3429. As in Birds the whole body was subordinated to the thorax, in Reptiles to the abdomen, in Fishes to the sex, in Insects to the tracheæ, and so on; in like manner is it here subordinated to the system of the senses or the head.
3430. As the higher senses determine or define the anterior part of the head, and are in their state of perfection provided with muscles; so here the face or visage is invested by flesh, whereby, properly speaking, a true, namely, a moveable countenance, first originates. The Aistheseozoa have a _fleshy face_.
3431. All possess moveable eyes; fleshy noses, which stand open both externally and internally; ears opening outwards, and mostly provided with a moveable flap; a fleshy tongue, free in front, and moveable lips; with at least thoracic limbs and a skin covered with hairs.
3432. In the Bird, Reptile, and Fish the face is merely invested by tegument, nearly devoid of any muscles, and therefore immoveable. They have a _tegumental face_, which is incapable of producing any expressions.
3433. In the tegumental face the eyes are motionless, and very rarely both directed so forwards that they can view an object together; the nostrils are frequently devoid of a fleshy rim; the tongue often feather-like, cartilaginous, or covered with teeth; true fleshy lips are wanting, as are frequently the teeth, with even the limbs and digits, or these are divided into a number of rays, forming either feathers, or fins; in the Aistheseozoa there are never more than five toes present, and if there be fewer, then the number admits of being referred to some crippling of the normal quantity, five.
3434. It is remarkable, and serves to the discovery of many laws, that the highest sense here appears for the first time in its state of perfection. The eye in the Aistheseozoa is present in a perfect condition throughout the class, with the exception of the eyelids; the other organs of sense are, on the contrary, exhibited in all their gradations of structure.
3435. It appears, as if the whole animal was first perfected, when the eye is present with all its investment or clothing. The eye of the Aistheseozoa has not simply all its internal chambers and all its humours, but also all its muscles; it is moveable and has perfect eyelids, with very few exceptions--_Ophthalmozoa_.
3436. The ear now begins to suffer arrest. Its completion is indeed the formation of an external concha or flap for receiving the rays of sound, the hand being repeated in the ear, and its skeleton constituted by the auditory ossicles. This auditory hand occurs only in the Aistheseozoa, and might serve as characteristic if it were not wanting in many, while the eyelids are present. As in the Whales, where the auditory passage can be, however, closed, an act which is not possible in any Bird. The Bird must hear whether it will or no.
In all Thricozoa the interior of the ear is perfect, having cochlea, semicircular canals, tympanum, and, as brachial parts, the three conjoined auditory ossicles. The concha of the ear passes besides through all stages of development, from having a simple margin to one with most varied convolutions, lobes and opercula.
3437. Still more than the ear does the nose undergo modifications. In the Whales it seems to be less adapted for the purposes of smell than of respiration. The olfactory nerves are in them very delicate, and the apex of the nose is destitute of motion.
In other beasts, on the contrary, it is elongated into a very muscular proboscis or snout, and is endowed with the power of voluntary motion.
The form also of the nostrils is very varied, they being round, narrow, patulous, and frequently capable of being closed.
3438. The tongue is indeed mostly fleshy and soft; yet in many species it is provided with horny points, in others invested by a dense coriaceous tegument, so that it appears to represent an instrument for deglutition rather than gustation.
3439. The lips also are mostly fleshy and moveable; yet in many they retrograde exceedingly and lose their mobility, as is partly the case in the Ornithorynchus.
3440. In most, however, the limbs, especially the toes, are still subject to variation. Their perfection consists in the number five, and in the difference between the two pairs of limbs, as in Man. In the Apes the posterior feet are also hands, which is an imperfection; in the Marsupials there are hands posteriorly but toes in front; in other respects there are generally toes, sometimes five, then four, finally two perfect and two dew-claws in the Ox, and at length only one in the Horse, while the posterior extremities are virtually lost in the Whales.
3441. The dental system, as being the set of maxillary claws, is alone present in its state of perfection in the Thricozoa. They alone have, in addition to the incisor teeth, all the five kinds of teeth that differ from each other in form, namely, canine, false molars, laniary, with second and third true molars, corresponding to the five fingers reckoned from the thumb.
3442. In the dentition of those animals which tear their food the greatest amount of completeness and variety is met with, since each tooth has a different form and function.
In the Bears the molars are uniform in character, and so on through the Apes up to Man.
In the Marsupials they are tolerably similar, as also in the Bats and Shrew-mice.
They are still more alike in the Pig and Horse, and the incisor teeth begin to be wanting in the Ruminantia, e. g. the Ox.
In Mice the canines are wanting, in the Sloths the front teeth, and in the Ant-eaters actually all.
3443. In opposition to the perfect eye the general sense of feeling is developed in the tegument. The tegument which is best developed will be that, which represents a self-substantial organ with all its appurtenances, and thus an organ of touch, whose nobility of rank consists in motion. A skin, which is moveable by means of muscles, must take the noblest rank. A skin with _tegumental_ muscles is an organ of feeling, which is already in some degree subjected to the influence of the will. If tegumental muscles do not occur in all these animals, they still do in most of them.
The production of the most perfect coat or covering is the second step, whereby the tegument ascends.
3444. Hitherto the outer covering of animals was pretty inorganic, consisting either of hollow tracheæ, feathers, or semicavernous scales, coat of mail, or lastly, only mucus. All these organs were only constructed after the type of individual vegetable systems, in greatest part only after the respiratory organ, and were therefore partial in aspect or deficient.
The highest covering must also bear the highest meaning. This is that, which grows out of the sanguinary system collectively, out of the capillary vessels. The capillary vessels of the covering are, however, the hairs. The hair is the most perfect covering of the animal.
3445. These animals are the _Thricozoa_.
Already the highest birds, e. g. the Ostrich and Cassowary, exhibit feathers which merge over into hairs.
3446. An integument clothed with hair, and self-substantially moveable is the perfection of this organ, it is a _fur_--_Furred animals_.
3447. The fur is the peripheric combination of the Vegetable and Animal. The hairs are the highest Vegetable, as being the vascular system grown out of or above the animal, and constituting the fundamental or basi-system of the whole body. The tegumental muscles are the lowest Animal. In the fur consequently the whole animal has been represented, but as a limit set between animal and world.
3448. The tegumental covering also varies. The hairs become scanty in respect of number; instead of them spines, horny rings or scales occur, yet there are always hairs on the belly. In many Whales they seem to dwindle into bristles. In some they have coalesced to form a kind of bark, as in the Stellerian Walrus or Sea-cow.
_Thorax._
3449. In all the respiration is aerial; in all there is a true costal structure, and a respiration effected through the motion of the ribs or a pumping action; in all the lungs are filled with cells; in all there is a diaphragm, a larynx, a trachea with cartilaginous rings, and a thyroid gland. But the air no longer penetrates from out the lungs into all the cavities of the body, as in Birds.
_Abdomen._
3450. The two intestines, namely, the large and small, are more distinct from each other than in other animals; the cæcum coli is in most distinct; the stomach expanded and membranous in texture, so that it operates simply through chemical influence. Liver, pancreas, and spleen occur in all.
3451. The sexual parts are in every respect very perfect. The penis is present in all, and all have an uterus, fallopian tubes, and separate oviducts. The penis is in many still retracted into a cloaca, and the testes frequently lie within the abdominal cavity.
3452. As in the tegument the parts separate and each becomes self-substantial, the fibre animal, the tegument vegetal, and so on even in the sexual animal. The ovarium, consisting of two parts, now also separates, since one becomes _animal_, but the other remains vegetal. The albumen-organ becomes animal, detaches itself from the sexual parts, is developed in the self-substantial tegument and called mamma.
The Aistheseozoa are thus also _Mastozoa_ or Mammalia.
3453. This separation of the sexual animal is one of the first characters. The mammæ could never be wanting, because they indicate an essential stage in the development. The albumen-organ becomes an organ of sensation. If the existence of mammary organs be doubtful, as in the Ornithorynchus, the hairs are in that case perfectly distinct. No Reptile nor Bird can have hairs, because their covering is derived only from a partial system, the respiratory; while hairs are from the general vascular system, which is the foundation of the tegument, of the sense of feeling. The Ornithorynchus is a _furred animal_, and this would suffice to bring it among the Aistheseozoa, were we even to deny it mammary organs.
3454. Both sexes, male and female, have breasts. In youth they are most readily detected in the male, because their sexual parts then rank nearer to the female. In other respects the mammæ are probably the chiefest organs of absorption for the embryo.
3455. The mammæ are nobler in rank, when, as udders they become self-substantial; and the more they remove from the belly or abdomen, and come, as breasts, to be placed upon the chest.
D. ORDERS AND FAMILIES.
3456. Had the names been given in strict accordance with rank; then the class-divisions, which correspond to the circles, would have been called _Orders_.
3457. Families would be class-divisions, which correspond to the classes themselves.
3458. In no class therefore could there be more than four orders admitted, or five, if the sensorial organs be reckoned to constitute a special circle.
From the same reason there are no more than thirteen or seventeen families.--These names cannot meanwhile be so strictly adopted, because the classes are not of equal rank, as has been observed; on this account it is necessary to shove in here and there other divisions, which should be termed at one time _Cohorts_, at another, _Alliances_.
3459. If the animals of a class differ from each other, it is only possible by their bringing to bear in addition to their characteristic organ some other organ, and consequently becoming similar to an earlier or later class.
3460. Yet nevertheless in this ascent the animals could never outstep the confines of their own circle. There can be no Dermatozoon which could have bones; for in that case it would be an Osteozoon, and belong to another province.
Every class therefore comprises as many orders only as there are circles with which it comes into contact. Thus in the first circle there is only one order, in the second two, and so on.
Each class therefore includes also as many families only as the circles which are touched by it contain classes. Thus the first to the third class has three, the fourth to the sixth, six; the seventh to the ninth, nine; the tenth and so on, thirteen.
3461. The serial arrangement of animals into families is naturally difficult; but in Physio-philosophy we have to treat not about the execution of the Systematic in detail, but concerning its principles.
FIRST CIRCLE. INTESTINAL, OOZOA--MUCUS-ANIMALS.
_First Class._
_Gastric, Vitelline Animals--Infusoria._
3462. The Infusoria admit of being reduced to three divisions: the lowest of which, such as the Infusoria proper or Monades, are provided for the most part with cilia; the next in succession, as the Rhizopoda, possess extensible processes, and are mostly covered by a multi-chambered shell; lastly, the most perfect Infusoria have all kinds of internal organs, and especially what has been called rotatory apparatus, as being the dawn of future tentacula.
3463. These three Families obviously correspond to the three classes of the present circle, and that indeed as follows:
To the _first class_, or Infusoria proper, correspond the _Monades_.
To the _second class_, or Polyps, the _Rhizopoda_.
To the _third class_, or Acalephæ, the _Rotifera_.
3464. The Monades are obviously the simplest organized creatures, being mucous vesicles, which move, obtain their food by stirring up vortical currents in the water, and emit what is undigested again by the mouth.
3465. They occur very abundantly in all infusions, and can very well originate, like Fungi, by division of the organic mass, although they are in a condition to propagate themselves, i. e. by spontaneous division.
3466. The Monades are the semen of the animal kingdom, which is dissolved in, or rather produced from, the sea.
3467. The animal body is nothing else than a compound fabric of Monads.
3468. Decomposition is a separation into Monads, a retrogression into the primary mass of the animal kingdom.
3469. All propagation, even that of the sex, commences like the animal kingdom, or with its first family. On that account the embryonic development must be a passage through the animal kingdom.
3470. The Rhizopoda usually adhere together within a multi-chambered calcareous shell, out of which they protrude mucous filaments, and are therefore the antetypes of the Corals or Polyps.
3471. The Rotifera exhibit all kinds of viscera, such as intestine and ovaries; besides what have been called rotatory organs, which remind us of the arms or tentacles of the Acalephæ.
3472. The families of these animals may be therefore aptly named as follows:
Fam. 1. Typical Infusoria--_Monades_; cilia or vibratile organs. 2. Polypary Infusoria--_Rhizopoda_; extensible processes. 3. Acalephan Infusoria--_Rotifera_; intestine and oral organs.
_Second Class._
_Intestinal, Albuminous Animals--Polypi._
3473. The Polyps also do not admit of being divided into more than three families. The first are only tubes or vesicles with _capillary tentacula_ around the mouth, as the naked Polyps, Tubulariæ, Sertulariæ, and Cellulariæ.
The others have true ciliated _tentacular rays_ surrounding the mouth, and are always condensed inferiorly into a horny, and occasionally stony axis or stem, as is seen in the Gorgoniæ, Alcyoniæ, and Isidiæ.
The third have ordinary _tentacula_, occurring in great numbers like tassels or tufts around the mouth; their integument is either hardened into stone, or becomes fleshy, as in the Star-corals and Actiniæ.
3474. There is no doubt that the _naked Polyps_ are closely allied to the Infusoria, and among them indeed to the Rotifera, so that they represent but a higher stage of these creatures, characterized by superior size and by tentacular in place of ciliary or vibratile hairs.
3475. The _Cellulariæ_ cannot be more distinctly characterized than by saying that they are cortices or ramules inhabited by Vorticellæ. Thus they are Vorticellæ surrounded by a shell, and may be compared to ova, in whose coriaceous or leathery shell calcareous granules are blended, as in the eggs of the Crocodiles and Tortoises.
3476. They increase by means of ova and by ramification, if the former mode of division is not carried too far.
3477. The tubes of the Tubulariæ appear to be nothing else than the posterior extremity of the Polyp, desiccated or dried. These tubes are not therefore the product of an excretion, but the body itself.
3478. On the contrary, the tubes of the Sertulariæ must be held as being a tegumental excretion, within which the Polyp ramifies and produces ova-cysts. If the naked Polyps resemble ova which are devoid of shells, such as the roe or spawn of Fishes, then the Sertulariæ resemble ova surrounded by a tough skin, like those of Rays and Serpents.
The _Radiated Polyps_ or Horn-corals are invariably ramified and converted towards their interior into a common horny or stony mass; so that the animals themselves appear to have coalesced upon this into a common tegument or bark. They possess a stomach and ovaries surrounding it, which open into the border of the mouth between the rays. They thus increase by ova and ramifications.
They therefore represent the class proper of the Polyps.
The tufted Polyps comprise the proper Corals or Lithozoa, are in form and substance like the Acalephæ but with this difference, that the covering is mostly lithoidal, while in many Acalephans, as the Porpitæ, it only appears as a cartilaginous disc or plate.
3479. These Corals are true ova, having a perfect calcareous shell, like that of the Bird. The gelatinous animal which adheres within a wide-mouthed Madrepore, e. g. Fungia, resembles a vitellus just hatched, and from which the foetal envelopes have been developed.
3480. The numerous tentacular filaments surrounding the wide mouth resemble the shags of the chorion, which accumulate around the orifice of the umbilical cord to constitute a placenta in the higher animals.
The Corals are brood-eggs inclosed within the uterus of Nature or in the sea. The Coral-animals are an umbilical cord, treasured up for the embryo, while the Tubulariæ are only membranous vitelli, and the Gorgoniæ, ova with desiccated albumen.
3481. The families of Polyps stand therefore in the following order of significance:
Fam. 1. Infusorial Polyps--_Tubulariæ_. 2. Typical Polyps--_Alcyoniæ_. 3. Acalephan Polyps--_Actiniæ_.
_Third Class._
_Absorbent, Involucral Animals--Acalephæ._
3482. The Acalephæ also can only be brought under three divisions, viz. Röhrenquallen, or _Physaliæ_, Rippenquallen or _Beroes_, Hutquallen or _Acalephæ_.
3483. The _Physaliæ_ are without doubt the lowest, being only giant Infusoria; the _Beroes_ are allied through their simple bodily cavity, to the Gorgoniæ; the _Acalephæ_, through their form, to the Madrepores, especially the Actiniæ.
3484. They range therefore in the following order of significance:
Fam. 1. Infusorial Acalephæ--_Physaliæ_. 2. Polypary Acalephæ--_Beroes_. 3. Typical Acalephæ--_Medusæ_.
3485. The _first_ have not yet attained to the unity or single character of the mouth, but imbibe their food through numerous tubes. They are bundles of ramified Vorticellæ, a thoracic duct full of glands and roots, which absorbs nutriment out of the sea, instead of an intestine.
As the first family they are the antetypes of the Mussels and Entozoa, and especially of the Hydatids and Tape-worms, and we shall not be far from hitting the mark, if we compare their air-bladder with the hindermost cystiform member of the body in the Hydatids.
3486. The _second_ family have a single mouth and ribs mostly full of lamellæ, which are probably the antetypes of gills; they are also frequently traversed by nutritive vessels.
They are antetypical of Snails, being in form an abdominal pouch, in substance a liver, and frequently possessing paired tentacula. Higher up, and they indicate the Crabs.
3487. The _third_ have, as a general rule, one central mouth surrounded by four arms and very numerous vessels, which run from the gastric cavity to its margins, and there mostly elongate into filaments or hairs, being thus true lymphatic vessels, which convey the nourishment directly from the stomach through the whole body, and become on its margin tentacula.
3488. As antetypes of the Kracken they particularly manifest the form of the Sepiæ, or Cephalopods, in the strong and frequently papillated arms that surround the mouth. Gland-like nodes are also developed in the margin of the pileus, but their meaning is as yet doubtful. Upon a higher stage, and they are antetypical of Insects.
Their vascular system forms an exceedingly regular, quaternary net, with branches and ramules ranging opposite to each other, which have been believed to be the vitelline network of the ovum when hatched. These vessels terminate likewise in a common marginal vessel, as in the vitellus.
3489. Viewed as a whole they resemble in form, appendages, and substance the foetal involucra or envelopes. The upper surface may be compared with the convex back of the envelopes, the lower to the concave funnel of the umbilical cord, and the succigerent filaments or hairs to the shags of the chorion. They are probably elongated by injection, like the so-called legs of the Star-fishes.
The secretion of the ova in four ovaries speaks also in favour of their higher grade of development. They lie in four cavities around the stomach, which also open near to the mouth. Their size also is significantly expressive of their higher position.
Finally, they are of separate sexes. In them is the milt for the first time distinctly displayed, and in the same situation indeed where in other animals the ovaries are.
3490. Hence, in these three classes there are found no more than three families, which together make up only _one_ order. The want of a fourth family and so on, proves that the development of these animals does not pass over into the succeeding circle. Their body therefore is only an homogeneous, transparent mass, variously excavated and perforated, but not separated into two cysts, namely, tegument and intestine, and without the other viscera, which are formed by the vascular system, such as the liver, kidneys, and salivary glands.
SECOND CIRCLE. VASCULAR, SEXUAL ANIMALS.
3491. These animals will both traverse the three classes of their circle, as also repeat the preceding three classes, and they consequently divide into two orders and six families.
_Fourth Class._
_Venous, Ovarial Animals--Mussels._
3492. The Mussels or bivalve Mollusca resolve themselves into two orders, according to the structure of their mantle or respiratory sac. It is either closed in a tubuliform manner, and opens posteriorly into two mostly tubular-shaped respiratory apertures, and in front has an orifice for the passage of the foot--_Camacea_ (_Lochmuscheln_); or it is slit along its whole length anteriorly; and the pedal aperture is confluent with the anterior and also indeed with the posterior respiratory opening--_Ostracea_ (_Spaltmuscheln_.)
The first order still represent the cystic form of the Oozoa, and partly by tubuliform calcareous shells placed around the two ordinary shell-valves; but the second, by the freer development of the organs, represent the animals of their own circle.
_Order_ 1. _Protozooid Mussels_--_Lochmuscheln_.
3493. These animals repeat the Mucus-animals; are tubular in form, and mostly provided with two long respiratory tubes.
_Fam. 1. Infusorial Mussels--Sackmuscheln or Pholades._
Cylindrical, with a tubuliform mantle almost entirely closed and frequently surrounded by a calcareous tube, in addition to the two ordinary shell-valves; the pedal aperture is situated at the oral extremity; the respiratory tubes retractile by means of muscles; the foot is cylindriform.
Of the same kind are the Pfahlmuscheln, Teredines, and Sandmuscheln, as also the Solenidæ. They typify the Nudibranchiate Snails and the Salpæ.
_Fam. 2. Polypary Mussels--Klaffmuscheln or Tellinidæ._
Body flat, mostly discoidal; there is a large fissure on the abdominal side of the mantle for the passage of the foot, while posteriorly there are two long respiratory tubes with retractor muscles. Here belong the Tellinidæ and Venusidæ.
I have shown, that they are already to be recognized by the uncinate or hook-shaped groove for the mantle in the shell, and which proceeds from the insertion of the retractor muscles. The foot is usually lancet-shaped. They typify the Patellæ and Ascidiæ.
_Fam. 3. Acalephan Mussels--Cardiacea._
Body globular in form, mantle having a pedal slit on the abdominal side; two respiratory apertures without retractor muscles; foot mostly uncinate or strap-shaped.
T have shown that these creatures are to be recognized by a discoid groove in the shell, which simply depends upon the mantle, the retractor muscles being wanting. Here belong the Cardiacea and gigantic Chamidæ, in which last a byssus occurs, and the union also of the two occlusor or sphincter muscles, as in the following order. They antetypify the air-breathing Snails and Cirripedia.
_Order 2. Conchozooid Mussels--Spaltmuscheln._
3494. These animals represent their own circle.
The pedal slit in the mantle occupies the whole length of the latter, and is confluent with the anterior or also with the posterior respiratory aperture; so that only one or even no aperture is left, and this when present is besides never elongated in a tubular form; the retractor muscles are therefore wanting also, and the shell simply exhibits a discoid groove for the mantle. The occlusor muscles approximate and unite at their terminations. Both usually coalesce into one.
_Fam. 4. Typical Mussels--Mytilaceæ._
Occlusor muscles separate, and mantle slit to such an extent, that only the posterior respiratory aperture remains; foot mostly tongue-shaped or coniform, with occasionally a byssus.
Here belong the fresh-water Mussels and the Mytili proper, of which last many bore into rocks. They typify the Capulidæ and Brachiopoda.
_Fam. 5. Snail-Mussels--Arcaceæ._
Have two separate occlusor muscles and a perfectly separate mantle without respiratory aperture; the foot small, mostly cartilaginous. Here belong the Arcæ, Aviculæ, and Pearl-mussels.
They typify the Trochidæ and Pteropoda.
_Fam. 6. Kracken-Mussels--Ostracea._
Only _one_ occlusor muscle, mantle entirely open or slit, so that both respiratory apertures are only oblique incisions therein; foot very small, frequently furnished with a byssus. They typify the Buccinidæ and Cuttle-fish.
_Fifth Class._
_Arteriose, Orchitic Animals--Snails._
3495. The Snails likewise divide into two orders, according to the two circles, preindicated by the Ovum-and Sexual system. Their branchiæ are either ramulate or pectiniform, the sexual parts combined or separated.
The first kind are still frequently gelatinous in texture, transparent and naked; their branchiæ usually stand out freely as filaments, lamellæ, or ramules upon the back, or lie simply as a vascular network within the mantle. All are androgynous. They therefore obviously repeat the Protozoa or Mucus-animals.
The second are invariably covered by the shell and by a mantle, within the cavity of which the branchiæ lie concealed as one or two comb-like bodies. Tentacula and eyes, which are occasionally wanting in the preceding order, are here universally present; the sexes separate.
As in the Snails, the male parts make their appearance for the first time distinct and individualized, and are also a characteristic organ of the class; so also do they serve as bases of division, and the Snails may be divided into those which are Androgynous or bisexual, and those with separate sexes, i. e. Dioecious.
_Order 1. Protozooid Snails--Androgyni._
3496. The male and female sexual parts united together in a single individual, branchiæ ramuliform, occurring either as filaments, leaflets or ramules, freely situate upon the naked body, or as a network lodged within the pallial cavity and surrounded by a shell.
The Nudibranchiate species live in the sea, those with hollow branchiæ or the Pulmonea in the air; the former feeding mostly upon animals, the latter upon plants.
_Fam. 1. Infusorial Snails--Notobranchiata, or Tritoniæ._
Body gelatinous and membranous, cylindrical and naked, devoid of shell, with branchial filaments or ramules disposed in two rows upon the back.
Here belong the Tritoniæ and Dorides. Their body is muscular; the tentacula not retractile; the male sexual parts open in company with the female upon the right side of the neck, as in the higher organized Snails. All inhabit the sea. They prefigure or typify the Salpæ.
_Fam. 2. Polypary Snails--Pleurobranchiata, or Patellæ._
Body and sexual parts as in the preceding family, the branchiæ, however, occurring as ramules or leaflets upon the sides of the body, are more or less covered.
Here belong the Phyllidiæ, Schüssel-and Schildschnecken. They are the antetypes of the Ascidiæ.
_Fam. 3. Acalephan Snails--Dictyobranchiata or Limacidæ._
The branchiæ form a rete or network within the pallial cavity, and respire the moisture of air; mantle and viscera are mostly surrounded by a shell; the body is therefore bipartite, being separated into a splanchnic or visceral body and a foot with head.
Here belong the Air-breathing Snails, both land as well as fresh-water species. They typify the Cirripedia.
The shells are mostly thin and horny, yet nevertheless contain a considerable quantity of calcareous earth, and are mostly devoid of opercula.
Those species, which dwell in fresh water, do not possess introvertible tentacula like the marine Snails, and the eyes are placed at their basis; the sexual orifices are separate.
In the Land-snails, the tentacula are introvertible, and support the eyes upon their apex; the sexual apertures are blended into one. The former, like the marine Snails, lay numerous small ova inclosed within a gelatinous mass in the water; the latter deposit their ova free, covered with a membranous, and occasionally calcareous shell, in the earth. Copulation is effected in all by a reciprocal interchange of the androgynous species.
I have observed that individuals of Limnæa auricularia can continue to propagate for several generations without being impregnated; they _must impregnate themselves_.
_Order 2. Conchozooid Snails--Dioecii._
3497. Branchiæ situated within the cavity of the mantle, and hanging down in the form of a comb; shell for the most part spirally contorted; sexes separate--_Pectinibranchiata_.
Here belong the Capulidæ, Turbinidæ, Neritæ, Conidæ, Volutidæ, Buccinidæ, Muricidæ and Strombusidæ.
The tentacula are not retractile, and have the eyes mostly seated at their basis; the penis is external, very large, and cannot be drawn within the body, but is only reflected into the pallial cavity; most of them have a protrusile perforating proboscis, and an operculum. They lay numerous small ova, contained within large membranous cases, which frequently hang in rows to each other like a necklace of pearls. The shells are in some instances horny, in others of a stony texture.
_Fam. 4. Mussel-Snails, Capulidæ._
Only one branchial comb within the mantle, and covered merely by a flat hood-shaped shell; no operculum.
Here belong the Capulidæ; all the species are marine. They prefigurate the Brachiopoda.
_Fam. 5. Typical Snails, Turbinidæ._
Two branchial combs, mantle devoid of a respiratory groove, shell turbinated; mostly furnished with an operculum, and that indeed of a stony texture. Here belong the Turbinidæ, and Trochidæ, such as the Cyclostomata, Paludinæ, Ampullariæ, Janthinæ, Neritæ. Live in the sea and in fresh-water.
_Fam. 6. Kracken-Snails, Buccinidæ._
Like the preceding family, but their mantle has a groove, and there is a horny operculum. Here belong the Conidæ and Volutidæ, Buccinidæ, Muricidæ, Strombusidæ. Almost all live in the sea and are of sanguisugal habits. They antetypify the Cuttle-fish.
_Sixth Class._
_Cardiac, Nephritic Animals--Kracken._
3498. Body cylindrical, without sole or foot; frequently venous hearts together with the arteriose hearts, and a kidney-like organ.
Such are the proper _Cylindrical Snails_, which are either sessile or fixed, or move themselves by fins and what have been called arms--Sessile and Natant Kracken. Here belong the Salpæ, Cirripedia, Brachiopoda, Pteropoda, Heteropoda, and Cephalopoda. They all live in the sea.
Branchiæ and sexual relations very varied.
They divide likewise into two orders, in accordance with the two circles of Mucus-animals and Conchozoa; the former are spathiform, mostly gelatinous, and firmly sessile or fixed; androgynous, without head and rudders or fins. The branchiæ trellis-like and filiform--Ascidiæ, Cirripedia and Brachiopoda.
The others have a kind of head with arms and eyes, or fin-shaped steerage-organs upon the body; branchiæ pectiniform, reticular, and leaf-shaped--Pteropoda, Heteropoda, and Cephalopoda.
_Order 1. Protozooid Kracken--Rumpfkracken._
3499. Body sacciform, mostly gelatinous, without head and eyes; species androgynous. They are destitute of independent locomotion, most of them being quite stationary or fixed, and surrounded frequently by a shell; some few float about passively in the sea.
_Fam. 1. Infusorial Kracken--Salpæ._
The _Salpæ_ are gelatinous, freely natant cylinders, perforated by an open tube, within which are situated the branchiæ, heart, mouth, intestine, and liver, without any tentacula. They continue to hang for a long time to each other, as though they were in the ovary.
They undergo a remarkable metamorphosis, which traverses two generations, the young meanwhile not resembling the mother but the grandmother.
Body within a sacciform mantle, with two opposite respiratory apertures.
_Fam. 2. Polypary Kracken--Ascidiæ._
The _Ascidiæ_ have a sacciform body with two co-approximate respiratory apertures; branchiæ internal and trellis-shaped.
These gelatinous or cartilaginous animals adhere firmly to rocks, with the respiratory apertures directed upwards as in the Mussels, but the branchiæ are not foliiform; the internal cavity is lined with a membranous sac, upon which the branchial vessels are dispersed in a trellis-like manner. At the bottom of and within the sac is the mouth, which is destitute of tentacula. Intestine, liver, and a heart that is simple or undivided. Mode of propagation unknown.
They are frequently connato-sessile, like the Polyps.
_Fam. 3. Acalephan Kracken--Cirripedia._
Body sacciform, without branchial openings, oral apertures situated at the inferior extremity of the longitudinal fissure in the mantle, and provided with a kind of maxillary apparatus; above or behind the mouth are annulate or horny filaments representing caudal fins or feet, upon the basal joints of which is a branchial filament. Here belong the Balani and Lepades--_Fuss-kracken_.
These animals are likewise sessile, at least when full grown, and are surrounded by shell-plates, which remind us of the Echini or Sea-Urchins, whose antetypes they are; this resemblance is particularly distinct in the Balani, which have similar odontoid valves upon the aperture or mouth of the shell. These animals also undergo a metamorphosis, since in the early periods of their existence they swim about like small Crustacea, and later on become fixed or sedentary, and surrounded by the plates of shell.
The Lepades have the ordinary shells of Mollusca, but each of the two valves consists of two pieces, between which an odd one is interposed down the back.
These five shell-pieces become in the Balani a regular set of teeth, which are adherent within the cylindrical shell, by which the posterior part of the body, or the peduncular and elongated mantle, is surrounded. (Vid. Oken's 'Allgemeine Naturgeschichte,' V, 1835, 509.)
They bear a resemblance to the Crustacea, not simply in the annulate horny feet, but also in the double and ganglionic nervous chord lying along the abdomen; only they are androgynous, which occurs in no Crustacean.
_Order 2. Conchozooid Kracken--Kopfkracken._
Tentacula upon the head or fins upon the body. They repeat the Conchozoa or Shell-animals.
_Fam. 4. Mussel-Kracken--Brachiopoda._
The body is surrounded by a mantle open above, and by two shells; on the sides of the mouth there are two arms.
These animals look exactly like Bivalve Mollusca; they are firmly adherent to rocks, being frequently supported upon a hollow pedicle, which is a prolongation of the mantle. Branchiæ and sexual parts but little known; species probably androgynous--_Armkracken_.
3500. In the following families the body is free and sacciform, provided with arms or fins, and mostly with eyes upon a kind of head.
These animals have the mouth situated above or in front of the body, and are distinctly separated into mantle and abdomen, while they are frequently surrounded by a shell. They have a kind of head with tentacles or arms, and frequently furnished with eyes like the Snails, or pretty nearly like the Fishes; the sexual parts united and separated. They all row, paddle, or steer themselves about in the sea.
_Fam. 5. Snail-Kracken--Pteropoda._
Body mostly gelatinous, sacciform, closed all round and free; fins upon the sides of the neck, with two tentacula frequently projecting from near the mouth; androgynous, naked, and inclosed within a shell.
These transparent animalcules swim about upright in the sea, and wave the fins in this position as rapidly as a butterfly does its wings. Most of them are covered with a spathiform and likewise transparent shell. The branchiæ are placed externally on the body, but are not very distinct. Several species have an appendage in front of the neck, which is obviously a rudiment of the Snail's pad or sole--_Flugelkracken_.
The Heteropoda appear to resemble the Plantar or Sole-Snails, but the body is mostly gelatinous, and the foot or sole is only compressed to form a fin, so that they can only swim but not creep with it. Many of them have a shell almost like that of the Argonauta. Pterotracheæ--_Ruderkracken_.
_Fam. 6. Typical Kracken, Cephalopoda or Sepiæ._
Muscular animals inclosed within a sacciform mantle, open in front; head furnished with eyes, and surrounded by more than four strong prehensile arms; laminated or leaf-like branchiæ within the mantle, sexes separate; in the body is a nephroidal or kidney-like gland, which secretes a dark-coloured or inky fluid, on which account they are called by us 'Dintenschnecken'--_Sternkracken_. They are obviously the highest organized Malacozoa, and already remind us strongly of Fishes, partly from their size, partly by their fleshy body, and in part by their perfect organs of vision.
The body is often as large as the trunk of a man; the head being separated from it by means of a neck, has in its interior a kind of cartilaginous brain-case or skull, with externally a pair of jaws resembling a Bird's beak, and with eyes tolerably similar to those of Fishes. Auditory organs are also present, which consist of a tympanic cavity inclosing an otolithe or ear-ossicle; nostrils are wanting. The so-called arms are perfect organs of locomotion, as also useful for seizing the prey, being provided for this purpose with sucking cups, which adhere by producing a vacuum, when applied. In other respects these arms are nothing else but the Snail's sole divided in front into lobes. The ova resemble berries, and hang attached to one another in branched clusters like a bunch of grapes. The Sepiæ possess a remarkable gland, which is complicated with the liver, and secretes a dark brown juice, which has been called ink or sepia; it probably ranks in the significance of the kidneys.
Many are covered with a shell externally, as the Nautili and Argonautæ; but in the common Sepiæ this shell adheres within the mantle upon the back, and forms a straight lamina or plate, which is sometimes horny, sometimes calcareous in its texture. It has been called the "white fish-bone." What is regarded as the dorsal side of the animal is its ventral side, because upon the former lies the pallial aperture and the anus, while the vitelline sac is also inserted there.
In the form of the body, as well as in the circular position of the cephalic arms, the Sepiæ strikingly resemble their antetypes, the Acalephæ and Cirripedia, and their metatypes, the Asteriadæ and Crustacea.
With these animals the second stage or circle through which the Dermatozoa pass, or that of the Sexual animals, is closed. They require only a slight additional grade of perfection, and they would pass over into another class of animals. Thus were the arms to become horny and articulate, they would be Crustacea; had the head a nose, and the body consequently a myelon or spinal chord, they would then be Fishes.
SECOND CIRCLE. RESPIRATORY, CUTANEOUS ANIMALS--ANCYLIOZOA.
3501. Body annulate.
Here range the Worms, Crustacea, and the proper or volant Insects.
3502. In this circle a remarkable relation is revealed. If, namely, its classes, orders, and families be compared with those of the two preceding circles, it is then distinctly shown that the present one again commences quite from below, ascends parallel with both these, and transcends them in its highest inmates or members.
As an example of the first case, the imperfect condition of some Entozoa, and their great resemblance to Infusoria, is sufficiently expressive.
The Worms evidently pass parallel to the Mucus-animals, and the Entozoa indeed to the Infusoria, the Red-blooded Worms to the Polyps, but the Holothuriæ chiefly to the Acalephæ, near which they are still arranged up to the present day, although they are articulated, and have an intestine and vessels.
Thus this class traverses the three inferior classes, and consequently imparts to them only the value of orders, or must itself be raised only to the rank of an order, a step, however, which, considering the great number of Worms, would not be suitable or just. The names may therefore remain for the sake of uniformity; only it must be borne in mind that they are of unequal value.
The same relation is exhibited in the Crustacea or Crabs. They obviously admit of being parallelized with no other animals than the Malaco-or Conchozoa. The Entomostraca and Crabs evidently repeat the Mussels, the Aselli the Snails; but the Spiders and Scorpions the Kracken or Cephalopoda. Consequently here also the orders correspond to the preceding classes.
The true or proper Insects transcend the Sexual animals, and therefore conform to the classification of their own circle, namely, they repeat the Worms and Crabs, and finally mount forth upon their own stage.
The inferior or lower organized animals form consequently two series, viz. the Smooth and Annulate, which in their lower members range parallel, and that indeed after the following manner:
A. _Smooth._ B. _Annulated or Ringed._
I. _Protozoa._ I. _Worms._ a. _Stomach_, 1. Infusoria. 1. White-blooded Worms. b. _Intestine_, 2. Polypi. 2. Red-blooded Worms. c. _Absorbents_, 3. Acalephæ. 3. Holothuriæ.
II. _Conchozoa._ II. _Crustacea._ d. _Veins_, 4. Mussels. 4. Crabs. e. _Arteries_, 5. Snails. 5. Aselli. f. _Hearts_, 6. Kracken. 6. Spiders.
III. _Ancyliozoa._ III. _Flies._ g. _Retia_, 7. Worms. 7. Tracheoptera. h. _Branchiæ_, 8. Crabs. 8. Dictyoptera. i. _Tracheæ_, 9. Flies. 9. Ceratoptera.
The Worms have simply an annulate body with reticular or filamentary branchiæ, but no feet; the branchiæ are supported by the body.
The Crabs have an annulate body with feet; the branchiæ are supported by limbs.
The Insects have an annulate body with feet and wings; the body is supported by the branchiæ.
_Seventh Class._
_Reticular, Papillose Animals--Worms._
_Protozooid Ancyliozoa._
3503. There are Worms with a soft body and white blood, without proper tentacula; others with red blood, with and without tentacula, having also bristles along the sides of the body; lastly, the rings of the body become hard or muscular, and a circle or ring of osseous maxillæ forms around the mouth, which is usually surrounded by numerous tentacula, as in the Star-fishes and Holothuriæ, in whom the blood is nevertheless colourless.
They divide accordingly into three orders, and each of these again into three families, in accordance with the classes, namely, those of the Protozoa or Mucus-animals, through which they pass.
_Order 1. Infusorial Worms--Parasites._
3504. The White-blooded Worms or Entozoa are very imperfect animals, a fact which proves that this circle recommences quite inferiorly or from below. They live for the most part in the interior of other animals, being thus located in darkness and in places where they obtain but little oxygen. Their blood therefore is not simply colourless, but even its vessels are only imperfectly developed. They respire without doubt through the tegument. In many, a distinct or separate intestine being wanting, it is the tegument also which digests; in others the intestine is simply a sac without an anus. The sexual parts too are in many species of a doubtful character, while meantime there are both androgynous and dioecious individuals. In the latter the male parts open invariably at the posterior extremity of the body, as in Insects; the female parts in front of its caudal end, as in the Crustacea. Both are in other respects constructed as in Insects, namely, there are two oviducts or seminal ducts, which unite before reaching their external orifice. They divide into three groups.
In one of these the body is tolerably smooth, and the pharynx commencing from a simple suctorial mouth elongates into a ramified intestine without an anus; they are androgynous--_Saugwürmer_.
In others the body itself supplies the place of the stomach; it is corrugated, and receives the food through one or several orifices, without separating into a special intestine. They have nearly all a claviform proboscis or rostrum, with which they perforate substantial cavities for themselves; it would seem that they are both androgynous and of separate sexes--_Hydatids_ and _Tæniæ_.
Others are separated into tegument and intestine like the Mussels and Snails, but without having a distinct vascular system, heart, and liver, though provided with a nervous chord and separate sexual parts--_Ascarides_.
_Fam. 1. Monad-like Worms, Saugwürmer._
Body tolerably smooth, suctorial mouth and sucker, the intestine losing itself in the tissue of the body, and without an anus; androgynous.
They remind us through their small size and structure of the Monades, especially the Cercariæ, and among the Worms typify or prefigurate in particular the Hirudines or Leeches, both by their form and power of suction, as also in the ramification of the intestine.
Many Cercariæ might be metamorphosed into Distomata, and so be or constitute their young.
The Planariæ are slightly different from the Liver-flukes or Fasciolæ hepaticæ, since they are white, bloodless, have a ramified intestine without anus, and increase also by fissiparous generation. They belong to the present order, although they live not as parasites, but in a state of freedom.
_Fam. 2. Rhizopodoid Worms, Hydatids._
Body annulate or articulate, with a claviform proboscis without an intestine; they appear to have several mouths.
The _Cystica_ develop ova simply upon the internal wall of the tegument; the body's last ring is expanded into a cystic form, and is mostly much larger than the whole body. The Coenurus cerebralis, or Brain-hydatid, ramifies like the Polyps.
The Cestoidea have in each joint or member of the body an ovarium, and, as it would appear, male parts also, so that they are androgynous. The sexual orifices are at the border or upon the surface of the posterior members. The member can be viewed as an ovarium liberating itself; as in the Lerneæ and Arguli.
The Filariæ or Thread-worms, which also belong here, are in the form of a cylindrical tube, with separate sexual parts, which open posteriorly.
_Fam. 3. Rotiferal Worms, Ascarides._
Body cylindrical, intestine free, with mouth and anus; sexual parts separate.
Upon the mouth these Entozoa have some papillæ or points, which may be regarded as tentacula. The larger species have a distinct nervous cord. The Gordius cannot well take any other position than the present, although it does live in a state of freedom.
_Order 2. Polypary Worms--Hirudines._
3505. Body cylindrical with branchiæ in or upon the tegument, blood red; two ganglionic nervous threads run along the ventral surface of the body, as in Insects; all are indeed androgynous.
Here belong the _Red-blooded Worms_, as Hirudines, Lumbrici, Nereides, and Serpulæ.
_Fam. 4. Tubularial Worms--Dermatobranchiata._
Body cylindrical. The branchiæ are only a vascular network in the tegument.
Here belong the Hirudines and Lumbrici.
The Hirudines have a perfectly naked body, without filaments and bristles, a perfect network of vessels containing red blood within the tegument, an intestine with an anus, both sexual parts androgynoid; posteriorly they have a pad or sucker; in the mouth there are mostly maxillæ, and upon the head simple eyes.
The Naiades and Lumbrici have bristles in longitudinal rows upon the sides of the body. The latter are androgynous. The Naiades increase by division.
The Thalassemæ have a protuberant white body; red blood-vessels only upon the intestine; mouth rostriform.
These Worms stick in the mud, and are nourished by it. They have here and there bristles, but which do not form any longitudinal rows. They appear to absorb water through the skin, and respire by the intestine. They cannot, by reason of their red blood, range with the Holothuriæ.
_Fam. 5. Alcyonioid Worms--Notobranchiata._
Body depressed, with pedal filaments, tentacula and branchiæ in two longitudinal rows upon the back.
These Worms all live in the sea, adhering mostly upright in the earth, and have frequently annulate, rigid tentacula, ocelli, and pairs of maxillæ, like Insects. The gills are not unfrequently covered with pergamentaceous scales, as in many Nereides, and especially in Aphrodite.
_Fam. 6. Actinioid Worms--Auchenobranchiata._
Body inclosed within a pergamentaceous or calcareous tube with lateral bristles, branchiæ and tentacula upon the neck or head.
The neck is surrounded by a kind of mantle, almost as in the Snails, so that several animals have been arranged here, which are now known to be veritable Snails. Upon the head many have a horny operculum, whereby they can close the shell.
Here belong the Amphitritæ, Terebellæ, Serpulæ, and Sabellæ.
_Order 3. Acalephan Worms--Sternwürmer._
3506. These creatures attain the most perfect structure of the Worms. The blood is white, the form cylindrical, globular and stellate; the mouth surrounded by a ring or wreath of maxillæ. The nerves form a ring around the pharynx, and upon the latter are placed membranous cysts, which spirt water into the tentacula or feet as they have been called, and thereby expand them.
It is impossible for these animals to continue to range with the Acalephæ, although they resemble them in outward form; for they consist of two cysts, the intestine having become freed from the outer one as a special sac; moreover, they have a perfect vascular system, distinct muscles, a mouth with a dental apparatus, which prefigurates a complete skeleton, a peculiar vascular system for the injection of the tentacula or feet, a nervous ring surrounding the pharynx, an ovarium entirely separated, and lastly, a perfectly annulate body.
_Fam 7. Physalial Worms, Holothuriæ._
Body cylindrical, very plentifully supplied with muscles. Mouth and anus present, the former of these being surrounded by a dental wreath and ramified branchimorphous tentacula; branchiæ upon the intestine, and branchial aperture at the anal extremity. They correspond to the Physaliæ.
Their tegument is a perfect muscular tunic, consisting of several longitudinal bands, which pursue a downward course upon its internal surface; in other respects being transversely corrugated and full of papillæ, partly replete with hollow podoidal filaments, which by the injection of water into them are elongated; thus, they present a resemblance to the Acalephæ.
_Fam. 8. Beroeal Worms, Echini._
Are at bottom Holothuriæ with ossified tegument, or Asteriadæ with shortened rays.
The Echini have an anus, pretty like that of the Holothuriæ, and also similar feet, which are protruded in a similar manner through apertures in the shell. The dental wreath surrounding the mouth is a complete bony framework, which has been called, on account of its form, "Laterna Aristotelis," and bears a considerable resemblance to the valves of the Balani, the antetypes in general of the Echini, just as these are of the Opossum Shrimps.
_Fam. 9. Medusal Worms, Asteriadæ._
Body stelliform, consisting of bony rings; contains a free multilobular intestine with blood-vessels and branchiæ; several ovaria; it would appear that they are without male parts.
Here belong the Encrinites, Pentacrinites, and Asteriadæ proper. Around the mouth of the Encrinites and Pentacrinites stand long, ramified, and likewise articulated tentacula, which remind us of the arms of the Acalephæ and Cephalopoda.
Fundamentally too, in the Star-fishes, the disk only is the body proper, and the rays are the tentacula, which, being monstrously developed, surround the mouth; what have been called pedicles upon these, represent the suckers of the Sepiæ.
_Eighth Class._
_Branchial, Pedal Animals--Crustacea._
_Conchozooid Ancyliozoa._
3507. The Crustacea are Worms with horny rings to the body, jointed feet and tentacula, which mostly breathe by branchiæ.
They correspond to the Conchozoa, or Shell-animals, and divide therefore into three orders, each of these including six families.
Here belong the Crabs, Wood-lice, and Spiders.
In the lowest forms the head, thorax, and abdomen are blended together, and the back mostly covered with a great horny scute--_Mussel-like Insecta_.
In the next place the thorax and abdomen admit of being clearly distinguished, both by their form as well as by their appendages, which are much larger on the thorax, as in the Cray-fish.
Then follow cylindrical forms with numerous feet, appended to uniform rings, but having a head free and distinguished by tentacula, maxillæ, and eyes--_Wood-lice_.
The Wood-lice bear a resemblance to the air-breathing Insects, although all parts of the body still pass over or blend gradually into each other.
In the _Arachnida_ or _Spiders_ aerial respiration takes place, and their body enters into relations with the more highly organized Insects, since the head becomes small, the abdomen large, thick, and short.
As the Crustacea pass parallel to the Conchozoa or Shell-animals, so do they divide into three cohorts or groups: the _Mussel-_, _Snail-_, and _Kracken-like_ Crustacea, or _Crabs_, _Wood-lice_, and _Spiders_.
COHORT I. MUSSEL-LIKE CRUSTACEA--CRABS.
3508. Head and thorax connate, being mostly covered with a shell or shield, abdomen stunted or caudiform; maxillæ and branchiæ present.
_Order 1. Lochmuschelartige, Pfriemenkrebse._
3509. The feet mostly simple and pointed, without large claws or forcipes, and furnished with setæ or lamelliform branchiæ; eyes sessile.
_Fam. 1. Pholadoid, Mussel-Insects._
Small, almost microscopical, and slightly annulated animals with an uni-or bi-valvular dorsal shield or testa, stunted maxillæ, and few legs, having attached to them setiform branchiæ.
These animals are found in all stagnant waters, in which they are incessantly paddling or rowing themselves about. They remind us of those Infusoria which, like Brachionus, are covered by a scute or shield. They are what have been called Monoculi (Entomostraca or Lophyropoda), e. g. Daphniæ.
Number of feet small, and beyond their appended setæ there are no branchiæ; two eyes frequently blended together; antennæ mostly furcate like the feet; sexes separate; a perfect circulation within a true Mussel-like ventral cavity. They are microscopic Mussels with eyes and feet.
_Fam. 2. Tellinoid, Branchiopoda._
Similar animalcules to the above, but with a body strongly annulated, naked or covered with a double shell, and provided with numerous feet, unto which are attached leaflets that serve as branchiæ.
Here belong the Branchiopoda properly so called.
_Fam. 3. Cardiaceoid, Shield-Crabs._
Large animals, with strongly annulated body and numerous feet; head and back covered by a shield, whereupon are the eyes--Phyllopoda.
Here range the Trilobites, Apus, and the Molucca or King-Crabs.
_Order 2. Spaltmuschelartige, Scheerenkrebse._
3510. Five pairs of thoracic feet, of which the first pair is mostly large and forcipiform; the eyes upon moveable pedicles or foot-stalks.
These animals usually attain a striking size, and are generally the largest among the horny Ancyliozoa.
The sexual parts open mostly upon the posterior legs.
The abdomen or tail usually supports five pairs of stunted feet, to which the ova hang.
The number of maxillæ agrees tolerably well with that of the thoracic feet.
_Fam. 4. Mytiloid, Locust-Crabs._
All the feet of pretty equal length, and the forceps stunted in size; the branchial plates free upon the abdominal feet--Squilla.
_Fam. 5. Arcaceous, Macroura._
Abdominal feet arrested, branchial comb upon the femora of the thoracic feet and beneath the dorsal shield, tail extended, as in the common Cray-fish.
_Fam. 6. Oyster-like Crustacea, Brachyura._
Characters similar, but tail bent under the body--_Taschenkrebse_.
COHORT II. SNAIL-LIKE CRUSTACEA--ASSELN.
3511. Body mostly cylindrical, annulate, without a true carapax or scute; head free; feet short and simple; branchiæ vesici-or folii-form; mostly placed under the tail; eyes non-pediculated, or even wanting.
_Order 3. Androgynoid--Saugasseln._
3512. Body soft and slightly annulate, maxillæ, feet and branchiæ arrested; live by suction as parasites upon other animals, mostly fishes.
_Fam. 1. Tritonia-like, Lernæaceæ._
Body soft, without shield, eyes and branchiæ; feet and maxillæ stunted; ova carried in two tubes at the hinder part of the body--_Kiemenwürmer_ or _Lerneæ_.
_Fam. 2. Patella-like, Argulaceæ._
Body distinguished into head, thorax, and abdomen, with few natatory feet, maxillæ moulded to form suctorial organs, mostly eyes; lay also strings of ova; head elongated in many in a scutiform manner. The so-called Fish-lice or Argulaceæ.
_Fam. 3. Luftschneckenartige--Pycnogonidæ._
Body short, with four pairs of long feet; eyes, but no branchiæ and maxillæ; abdomen puny in size. The _Whale-lice_.
_Order 4. Kammschneckenartige--Nagasseln._
3513. Body cylindrical, horny, and distinctly annulate, maxillæ, eyes, and mostly seven simple pairs of feet, branchiæ as cysts or leaves.
_Fam. 4. Capulidenartige, Walzenasseln._
Body cylindrical, with five or seven pairs of feet, and some branchial vesicles; body and abdomen very puny. The _Læmodipoda_; Caprella, Cyamus.
_Fam. 5. Turbinidenartige, Seitenasseln._
Body horny and distinctly ringed, mostly compressed, with perfect maxillæ; seven pairs of thoracic feet and branchial vesicles; abdominal feet rudder-shaped. They swim usually lying upon the side; many leap--_Amphipoda_; _Flohkrebse_ or _Gammarinæ_.
_Fam. 6. Buccinidenartige, Sohlenasseln._
Similar to the preceding family, but the body is depressed, and the abdominal feet furnished with branchial plates. The _Isopoda_; _Oniscidæ_, to which belongs the genus _Armadillo_.
COHORT III. KRACKEN-CRUSTACEA--KOBE.
3514. Body not tripartite; spiracula or air-openings; more than three pairs of feet, no wings. Here belong the air-breathing Crustacea; Scolopendræ, Acari, Scorpions, and Spiders.
These animals are abruptly distinguished from the preceding by a conversion of the branchiæ into spiral-shaped tracheæ, which ramify and traverse the whole body. They all therefore live in the air, and if they do also dwell in the water, they still come to the surface to inhale that element. The eyes are only simple points or ocelli, which are accumulated frequently upon the sides of the head.
The most inferior of them are distinguished from the preceding cohort, or the Asseln, by almost nothing save the essential character of their own cohort, the tracheæ. They have mostly a number of feet, and only simple eyes, as the Scolopendræ.
The following have a short body, in which the abdomen predominates; thorax and head connate; never more than four pairs of feet--_Acari_, _Scorpions_, and _Spiders_.
They likewise divide into two orders, like the Kracken.
_Order 1. Rumpfkrackenartige--Langkobe._
3515. Body horny, tolerably cylindrical and uniform; feet mostly very numerous.
_Fam. 1. Salpoid, Spindelkobe._
Only three pairs of thoracic feet, but still podoidal appendages to the abdomen--Podura, Lepisma.
_Fam. 2. Ascidioid, Schnurkobe._
Body cylindrical, with very many feet; sexual parts on the thorax--_Juli_.
_Fam. 3. Cirrhopodoid, Bandkobe._
Body band-shaped and depressed, maxillæ perforate, sexual parts placed posteriorly--_Scolopendræ_.
_Order 2. Kopfkrackenartige--Kurzkobe_.
3516. Body thick, mostly globiform; head and thorax blended together; only four pairs of feet.
_Fam. 4. Brachiopodoid, Acarides._
Body rounded; all three parts confluent; usually only two simple eyes. The Acari are mostly so small, that their parts can only be distinctly seen through the microscope. Their mouth is always very much arrested, and has maxillæ, which are in some cases adapted for manducation, in others for suction.
_Fam. 5. Pteropodoid, Scorpions._
Body tolerably cylindrical, and all three parts connate; palpi very large and forcipiform.
_Fam. 6. Sepioid, Spiders._
Body rounded, head and thorax connate, abdomen separate, mostly eight simple eyes.
Their most remarkable organs are the four spinnerets in front of the anus, which probably stand in the signification of the renal organs, just as the material of the Spider-threads does in that of the urine.
The tracheæ are but few in number, and expand into lung-like vesicles.
The sexual parts do not lie posteriorly by the anus, but at the root or base of the abdomen.
It is moreover remarkable, that their mandibles are perforated and pour a poisonous juice into the wounds they inflict. They must be therefore regarded, like the venom-teeth of Serpents, as elongated salivary ducts.
The Crustacean families can also be named according to the orders of their circle, namely, the Worms and Crustacea, e. g. thus:
ORDER I. _Worm-Crabs_--Pfriemenkrebse.
Fam. 1. _Entozooid_--Mussel-Insects. 2. _Hirudinous_--Branchiopoda. 3. _Holothurioid_--Shield-Crabs.
ORDER II. _Crustacean Crabs_--Scheerenkrebse. Fam. 4. _Typical Crustacea_--Sea-Mantes. 5. _Asselkrebse_ " --Macrura. 6. _Arachnoid_ " --Brachyura.
_Ninth Class._
_Tracheal, Alary Animals--Flies._
_Typical Ancyliozoa._
3517. Body tripartite, only three pairs of thoracic legs, spiracula and wings. They divide, according to the classes of their circle, into three cohorts, of Worm-, Crab-like, and perfect Flies.
As the wings are their characteristic organ, so also must they be divided according to them, and not after the cibarial instruments. Those wings which are developed to the least degree are the homogeneous, transparent wings traversed by few tracheæ or ducts--_Tracheoptera_, as Flies, Bees, and Butterflies.
Their chrysalis or pupa condition is perfect.
Then follow wings having very numerous veins united by transverse veins so as to form a network--_Dictyoptera_, as Neuropterous, Orthopterous, and Hemipterous Insects.
The pupa is moveable.
Finally, the anterior and posterior pair of wings become wholly dissimilar to each other, the former being horny, and the latter membranous in texture with reticulated veins, provided likewise with joints like the legs, so that they can be folded up under the anterior pair or elytra--_Ceratoptera_, as the Beetles.
Pupa state perfect.
Strictly speaking, it is a matter of indifference whether the names of orders and families be adopted from the first or second parallel series; whether e. g. we speak in the first cohort of Protozooid or Worm-flies. The nearest series will, however, bear the greater amount of resemblance to them. Meanwhile I will in the sequel vary in the choice of names, in order to exhibit different samples of this double parallelism.
COHORT I. WORM-FLIES--TRACHEOPTERA.
3518. Wings membranous with few longitudinal ducts, and almost devoid of transverse ducts, eyes larger than head. Here belong the Flies, Bees, and Butterflies. The abdomen is indeed annulate, but soft; its first ring is frequently set free and unites with the thorax, but supports neither feet nor wings.
The sexual parts always lie at the anal extremity.
The head is almost nothing but eye, and the Insects of this order may very well be termed Megalopidæ, or large-eyed, out of contrast to the succeeding ones. Between the two large compound eyes there are usually found three simple ocular puncta or ocelli, which they have adopted from the preceding class.
The larvæ are either entirely apodal, white and soft like Entozoa, or they have, in addition to the thoracic feet, numerous abdominal feet like the higher Worms.
The Flies with their soft and imperfect body, and the apodal larvæ, repeat the Infusoria and Entozoa; the Bees therefore the Polyps and Red-blooded Worms; the Butterflies with their large farinose or dusty wings and polypodal caterpillars, the Acalephæ and Holothuriæ.
_Order 1. Entozooid Flies--Diptera._
3519. All the thoracic rings coalesced, and along with them the first or basi-abdominal ring, unto which are attached the halteres or balancers; only the two posterior wings present; the labium prolonged into a proboscis, which incloses the setiform maxillæ. Larvæ apodal and white.
The Diptera resemble the Entozoa in a striking manner through their apodal, soft and white larvæ, and even through their habitation, which is mostly in fetid and moist animal matters. Added to this, they respire usually through two tubes which open upon the anus.
Many during their metamorphosis do not shed their skin; but it becomes, during the pupa condition, only horny, representing a small case, whose bottom springs up in front like an operculum, and gives exit to the perfect Fly which has been therein developed.
Upon the first abdominal ring of the imago two nodose pedicles stand out, which are called halteres; they are probably the old respiratory tubes.
The maxillæ have changed into bristles, which act like pestels within the groove-shaped labium, puncture and pump in the fluid.
They divide into three families like the White-blooded Worms, or according to the orders of their cohort.
_Fam. 1. Typical Diptera, Schmeissen._
Antennæ triarticulate, the last joint being mostly spatulate and furnished with a lateral awn or bristle; only two suctorial setæ inclosed between the terminal valves of a fleshy proboscis, which admits of being retracted within a large cephalic or oral cavity. The Muscidæ, unto whom the Common House-flies belong, and also the Hippoboscidæ or Louse-flies. It is my opinion too, that the Fleas also are to be ranged among the Diptera.
Alliance 1. _Klappenmucken._ The two punctuating setæ placed between two valves without a proboscis--Flea (Pulex) and Louse-fly (Hippobosca.)
Alliance 2. _Acalyptera._ Proboscis having thick lips, and retractile into a large cephalic cavity; alulæ or halteral opercula arrested--Hypocera to Dolichocera.
Alliance 3. Dung-flies. Characters similar to the above, but the alulets are of considerable size--Muscidæ.
_Fam. 2. Hymenopteroid Diptera, Dasseln._
Antennæ as in preceding family, but the bisetaceous proboscis is thin and horny, with small labia, or else large labia with four setæ--OEstridæ, Conopidæ, and Syrphidæ.
Alliance 4. _Parasitic Diptera._ Two setæ with or without a lipless proboscis--OEstridæ, Myopariæ, and Conopidæ.
Alliance 5. _Syrphidæ._ Four setæ within a similar kind of proboscis; the third joint of the antennæ spatulate; the palpi thickened.
Alliance 6. _Leptidæ._ Four setæ within a short thick-lipped proboscis, the third antennal joint mostly coniform--Therexidæ, Leptidæ, and Dolichopidæ.
_Fam. 3. Lepidopteroid Diptera, Gölsen._
Antennæ multiarticulate and stipiform; proboscis with and without labia, mostly four and six setæ for punctuation--Tipulidæ, Tanystomidæ, Stratiomydæ, Tabanidæ.
Alliance 7. _Gnats._ Antennæ filamentary and multi-articular. Proboscis varied.
Alliance 8. _Spiessmucken._ Four setæ within a hastate horny proboscis with or without arrested labia; third antennal joint not annulated, with the bristle or awn at the extremity--Tanystomidæ, as Asilidæ, Empidæ, Bombylidæ.
Alliance 9. _Stielmucken._ Four or six setæ within a thick-lipped proboscis; third antennal joint stipiform and annulated--Stratiomydæ and Tabanidæ.
_Order 2. Leech-flies--Hymenoptera._
3520. Four naked veined wings, labium mostly elongated, the two pairs of maxillæ acting above it like spears. Larvæ mostly apodal, or with more abdominal feet or prolegs than in the true Caterpillars.
Most of the larvæ still bear a great resemblance to the Entozoa, though they do not respire like them through the anus, but by lateral apertures or spiracula; the larvæ with abdominal feet repeat the Nereidæ and Aselli. In other respects they live no longer in putrid moisture, fungi, roots, and such like bodies, but in living animals or in cavities specially prepared for their reception by the parents, and even in a state of freedom upon leaves.
And here is particularly worthy of our notice the structure of the cells, which are by many species fabricated quite substantially of wax or wood-shavings, and are to be compared with the webs of Spiders, since both serve as nests for the young. They repeat the Polypidoms.
Others make cases of leaves, and carry into them honey upon which to deposit their ova.
Others again simply bore holes in wood or in the earth, in order that they may in a similar manner provide their young with honey or larvæ. Finally, others simply stick, by means of their ovipositor, the ova into animals or leaves.
The dwellings are fabricated by the maxillæ, which but seldom serve as cibarial instruments, seeing that the labium undertakes this office as a lick-organ.
Another remarkable feature of this order is the arrested development of the sexual parts of the females in certain generations, an occurrence which depends upon the time of the year or the size of the cells, and whereby they are constrained to lead a social life as workers.
They divide, according to the Red-blooded Worms or the orders of their cohort, into three families.
_Fam. 1. Dipteroid Hymenoptera, Bees._
Abdomen aculeate; labium elongated in a rostriform manner. They dig or build cells, and carry thither honey to the larvæ, which are apodal.
Alliance 1. _Fossorial Bees_--Andrenidæ. 2. _Carpenter-Bees_--Anthophoridæ. 3. _Cell-Bees_--Apidæ.
_Fam. 2. Typical Hymenoptera, Wasps._
Aculeate; labium not elongated--rapacious Hymenoptera, such as Ants, Fossorial, and Cell-wasps, larvæ apodal.
Alliance 1. _Hohlenwespen_--Formicidæ. 2. _Fossorial Wasps_--Sphegidæ. 3. _Cell-Wasps_--Vespidæ.
_Fam. 3. Lepidopteroid Hymenoptera, Terebrantia._
Instead of the sting there is an ovipositor, with which the females stick their eggs into other insects, mostly into caterpillars, or else into leaves and wood--Ichneumonidæ, Tenthredinidæ, and Uroceridæ. The larvæ of the first are apodal, those of the second provided with thoracic and abdominal feet like caterpillars, those of the third with thoracic feet only like the larvæ of Coleoptera.
Alliance 1. _Stutzwespen._ Ovipositor short; wings nearly veinless. Deposit their eggs in small insects; larvæ apodal--Chalcididæ, Oxyuri, Chrysididæ.
Alliance 2. _Schlupfwespen._ Ovipositor very long, divided into three hairs; wings veined. Deposit their eggs in caterpillars; larvæ apodal--Ichneumonidæ, Evaniidæ.
Alliance 3. _Pflanzenwespen._ Ovipositor spiral or saw-shaped. Deposit their ova in plants; larvæ mostly furnished with feet, and frequently resembling caterpillars in form--Cynipidæ, Tenthredinidæ, and Uroceridæ.
_Order 3. Holothurioid Flies--Lepidoptera._
3521. Four veined wings, covered with small dust-like scales; maxillæ have coalesced into a proboscis; larvæ with thoracic and abdominal legs.
The larvæ or Caterpillars remind us of the Nereides, especially the setaceous Aphrodites, such as the Holothuriæ, and further still, the Aselli and Scolopendræ. There are Caterpillars, which are scarcely to be distinguished from a Wood-louse. They subsist almost throughout upon leaves, and being exposed to the light are therefore variously coloured. They have maxillæ, and in the labium is the orifice of the salivary glands, from which they spin the threads used in weaving their cocoons, or social tents. Where they crawl they leave, like the Spiders, their threads beneath them.
The pupa state is perfect; the perfect insect generally creeps out of the chrysalis by a slit taking place down the back.
The abdomen is almost throughout covered with hair, which is indicative likewise of a strong mucous secretion.
They take in their fluid nourishment by suction, but the mechanism by which this is effected is not yet known, as from the jaws themselves forming the proboscis, there can be no suckers lodged within the latter that might act. It is probably effected by expansion of the abdomen taking place during respiration. Their deglutition would be therefore a respiratory act.
Besides their very non-artistic webs, no artistic instincts are to be observed in this order. The ova are just layed without more ado upon plants, and rarely upon other objects. The Butterflies are generally related to the plants, and especially to their corollæ, whose colours and forms they carry in themselves.
They divide according to their proximal orders into three families.
_Fam. 1. Dipteroid Lepidoptera, Moths._
Antennæ filiform, wings mostly thrown like a mantle round the body; proboscis short.
Small and nocturnal in their habits, proceeding from tolerably apodal Caterpillars, which reside mostly concealed in plants, or make themselves cases of hairs and leaves.
Alliance 1. Typical Moths, _Tineidæ_.
a. Typical Tineidæ: Alucitidæ. b. Pyraloid: Tineæ. c. Tortrix-like: Crambidæ.
Alliance 2. Bombycoid Moths, _Pyralidæ_.
a. Tinea-like: Aglossæ. b. Typical: Hydrocampæ. c. Tortrix-like: Deltoides or Herminiæ.
Alliance 3. Homalopteroid Moths, _Tortricidæ_.
a. Tinea-like: Fruit-Tortrices. b. Pyraloid: Heterogeneæ. c. Typical: Leaf-rolling Tortrices.
_Fam. 2. Hymenopteroid Lepidoptera, Silk-Spinners._
Antennæ filiform, wings tile-shaped.
Lepidoptera of considerable size, proceeding out of
Caterpillars, with many feet, and frequently covered with hair, which live mostly in a state of freedom upon plants, and in some cases make large cocoons freely exposed, in others under the earth--_Noctuidæ_ and _Bombycidæ_.
Alliance 1. Moth-like Spinners, _Noctuidæ_.
a. Typical: Hadenæ. b. Pseudobombycoid: Catocalæ. c. Bombycoid: Erebidæ.
Alliance 2. Typical Spinners, _Pseudo-Bombyces_.
a. Noctua-like: Psyches, Limacodes. b. Typical: Notodontæ. c. Bombycoid: Callimorphæ.
Alliance 3. Homalopteroid Spinners, _Bombycidæ_.
a. Noctua-like: Bombyces. b. Pseudo-Bombycoid: Saturniæ. c. Typical: Hepialidæ.
_Fam. 3. Typical Lepidoptera, Homaloptera._
Antennæ varied; wings flat and expanded; proboscis long--_Geometridæ_, _Vespertina_, _Diurna_.
Large Lepidoptera proceeding from free-living, strongly-coloured, and mostly naked Caterpillars.
The Geometridæ have level, tolerably triangular wings and filiform antennæ; they proceed from naked Caterpillars with few abdominal feet.
The Vespertina have level, long, and narrow wings, and spindle-shaped antennæ; they emerge from perfect Caterpillars.
The Diurna have the wings folded upwards in a reverse position, claviform antennæ, and proceed from perfect Caterpillars.
Alliance 1. _Moth-like Homaloptera, Geometridæ._
a. Typical: Geometræ. b. Sphingoid: Aposuræ. c. Papilionaceous: Uraniidæ.
Alliance 2. _Bombycoid Homaloptera, Sphingidæ._
a. Geometroid: Zygænidæ and Sesiadæ. b. Typical: Sphingidæ. c. Papilionaceous: Castniæ.
Alliance 3. _Typical Homaloptera, Papilionidæ._
a. Geometroid: Hesperiidæ, Lycænidæ and Erycinidæ. b. Sphingoid: Nymphalidæ and Heliconiidæ. c. Typical: Pieridæ and Papilionidæ.
COHORT II. CRUSTACEOUS FLIES--DICTYOPTERA.
3522. Four wings, with longitudinal and numerous transverse veins, the anterior pair being mostly leathery in texture; eyes mostly smaller than head.
Here belong the Neuroptera, Orthoptera, and Hemiptera.
The metamorphosis is tolerably imperfect, and the larvæ have never more than three pairs of thoracic feet, and therefore correspond no longer to the Worms, but to the Crabs, where the number of the feet has been already more determined. Many also strikingly simulate the Crustacea.
In all three families also the pupa runs about, eats, and has rudiments of wings.
The abdominal rings are mostly horny and hairless. The spiracula lie upon the upper margin.
The eyes are generally much smaller than the head, and may therefore, in comparison with those of the preceding and subsequent groups, be said to be of _middling size_; the simple eyes have for the greatest part disappeared. Meanwhile the first family displays _large eyes_, the second eyes of a _middling size_, the third _small eyes_.
They divide like their antetypes, the Crabs, into three orders.
_Order 4. Crab-Flies--Neuroptera._
3523. All the wings alike and membranous; maxillæ with mostly large eyes; abdomen soft.
These Insects which, on account of their delicate wings, are known with us by the name of Flohrfliegen, live mostly upon flesh, and many of them by capturing living prey. Many undergo a short pupa stage. Many live in the water and have, too, branchiæ, whereby they strikingly remind us of the Crustacea, especially the Branchiopoda.
Like the Crustacea, or in accordance with the two first cohorts of their class, they divide into two families.
_Fam. 1. Tracheopteroid Neuroptera, Nagbolden._
Do not subsist by rapine, but gnaw slowly animal and vegetable matters; many also as flies eat no longer.
Alliance 1. _Dipteroid Neuroptera, Kieferläuse._ No wings; gnaw skin, feathers and hair, mostly Bird-lice--Philopterus, Liotheum.
Alliance 2. _Hymenopteroid Neuroptera_, Blumen-and Mulmlaüse, Thrips, Psocus, Termes.
Alliance 3. _Lepidopteroid Neuroptera_, _Phryganeidæ_ (_Wassermotten._)
_Fam. 2. Dictyopteroid Neuroptera, Raubbolden._
Alliance 4. _Typical Neuroptera, Flohrfliegen_, Panorpidæ, Hemerobiidæ, Myrmeleonidæ.
Alliance 5. _Orthopteroid Neuroptera_, Raphidiidæ, Mantispidæ.
Alliance 6. _Hemipteroid Neuroptera_, Ephemeridæ, Libellulidæ.
_Order 5. Oniscal Flies--Orthoptera._
3524. Maxillæ and dissimilar reticular wings; the anterior, pergamentaceous; abdominal rings, horny; eyes of moderate size; pupa moveable.
Live for the greatest part on plants.
_Fam. 1. Tracheopteroid Orthoptera, Saltatoria._
Alliance 1. _Dipteroid Orthoptera_, Grylli.
Alliance 2. _Hymenopteroid_ " Locustæ.
Have a hard ovipositor, which reminds us of the sting in the Hymenoptera.
Alliance 3. _Lepidopteroid Orthoptera_, Acrydia.
_Fam. 2. Dictyopteroid Orthoptera, Cursoria._
Alliance 4. _Neuropteroid Orthoptera_, Mantidæ.
Alliance 5. _Typical_ " Blattidæ.
Alliance 6. _Hemipteroid_ " Forficulidæ.
_Order 6. Arachnoid Flies--Hemiptera or Bugs._
3525. Horny suctorial proboscis with puncturating setæ; dissimilar reticular wings, the anterior mostly pergamentaceous, the posterior membranous, and mostly capable of being folded up, as in the Chafers; eyes small, pupæ moveable.
The Bugs have much resemblance to the Mites, and Spiders are as it were winged Mites. One party of them suck blood, another vegetable sap. They manifest no artistic instincts.
The Aphides or Leaf-lice require only a single pairing, to enable them to propagate through several generations. During the summer they produce only females, and in autumn the males first appear.
_Fam. 1. Tracheopteroid Hemiptera, Leptoptera._
Alliance 1. Dipteroid Hemiptera, _Coccidæ_, _Aphidæ_.
Alliance 2. Hymenopteroid " _Cicadæ_.
Alliance 3. Lepidopteroid " _Fulgoridæ_.
_Fam. 2. Dictyopteroid Hemiptera, Pachyptera._
Alliance 4. Neuropteroid Hemiptera, _Water-bugs_.
Alliance 5. Orthopteroid " _Narrow bugs_, _Hydrometræ_, _Reduvii_, _Cimicidæ_.
Alliance 6. Typical Hemiptera, _Broad bugs_, _Capsidæ_, _Lygæidæ_, _Coreidæ_, _Scutelleridæ_.
COHORT III. PERFECT FLIES--CERATOPTERA, BEETLES.
3526. Maxillæ and dissimilar reticular wings, the anterior being horny, the posterior membranous and susceptible of being folded upon themselves; eyes small; pupæ motionless.
The Chafers or Beetles are the highest Insects, because they possess the greatest variety of organs, namely, two kinds of wings; not to mention the perfect condition of their manducatory apparatus, antennæ, and the first thoracic ring which is free.
The Beetle represents the Flies in their entire perfection; everything is rendered horny in the former even to the lower wings, which are furnished with several joints, almost like the feet. The upper wings are horny and meet together so closely by their inner margins upon the back, that they form a closed suture, like the shells of the bivalve Mollusc. The antennæ are also more perfect than in other families, and mostly jointed like the feet, while at their extremity they are frequently thickened into laminated moveable clubs, which open when the animal would fly, as if to listen.
Their habitation, mode of living and subsistence is exceedingly complex, and therein also they combine again all the families of this class, and it might be said, all the preceding classes.
The Beetles are also much more numerous than any other order, and could in this point of view even hold good for an entire class, especially, if they represented a special organic system, but which is not the case.
They live upon vegetable saps, and matters, blossoms, leaves, and wood, living animals, putrid flesh, dung, and such like.
They dwell mostly indeed in concealed situations, but also in those that are freely exposed, while many live in water, and are so subjected to its influence that their larvæ actually respire this element through branchiæ.
The larvæ are white and have three pairs of horny thoracic feet. They live concealed.
The pupæ are invested by a transparent tegument, which tears in an irregular manner.
The Beetles divide distinctly into three divisions, which correspond to the three cohorts of this class, or repeat the three cohorts of their circle.
The Beetles have been pretty generally divided according to the number of tarsal joints into those having three, four, five, or an unequal set of these "articuli," whereby, however, the greatest disorder has originated. I have therefore divided them in my 'Naturgeschichte' according to their mode of living into _Phytophaga_, _Sarcophaga_, and _Rypo-_ or _Coprophaga_, which gradually seems to meet with approval. At least a much more natural arrangement comes to light by using these means.
I have also declared the Rhyncophora to be the lowest and the Lamellicornes the uppermost in rank, a view, which at present appears to be generally adopted.
By these means I obtained two firm points, whereby the division or classification of the intermediate members is uncommonly facilitated, although many may still stand in the wrong place. But who can point out a single system in which Insects range correctly, or, what is more, wherein it would not be easy to prove that no single order and family occupies the right place.
In such trumpery systems of classification, one would think, that some respect might be had for principles, or at least a feeling of shame for the blockhead's weakness of intellect. But in vain! Where the sense for philosophy or for principles is wanting, it is not to be inculcated or driven in. We still see Natural Histories shoot forth, with whose shrub-like ramification we must have compassion. Every thicket of briars is rooted deep, and admits only of being extirpated by wearisome and patient culture.
The Rhyncophora indicate that the Phytophagous beetles are the lowest; the Lamellicornes that the Coprophagous are the highest. The Sarcophagous chafers accordingly take their place between the two.
Now, however, the Rhyncophora are tetrameral. In like manner are the Phytophagous beetles, such as the Borkenkäfer and Holzschröter. They consequently form the main stem, unto which all the true Phytophaga must be annexed. But among them also belong the Pentameral, namely, the Holzböhrer, Schnell-and Prachtkäfer; I have therefore disposed them in this order, and I believe quite correctly.
The Lamellicorn or Dung-beetles are pentameral, and on that account to be co-arranged with the Necrophaga or Carrion-feeders and Sexton-beetles, as well as with the parasitic Lampyridæ, and the predaceous Raub-and Laufkäfer, despite their exceedingly great difference of living and even anomalous structure. On the contrary, most of the heteromeral kinds agree in their mode of living, and tolerably too in structure, with the Dung-beetles, at least the Mulmfressenden, such as the Meal-beetles or Tenebrionidæ, and indeed the Stenelytra also, while the Cantharides are parasitic at least as larvæ.
It seems also to me, that the fungi must be viewed as flour or dung, and that therefore those very Fungivora, whose structure does not point the way otherwise, can be placed with the Mulm-and Mistkäfern, although they are only tri-and tetrameral, like the Lycoperdinæ and Erotyli. Besides this the antennæ of the Fungivora are usually thickened, and even club shaped, whereby they thus stand nearer to the Necrophaga, such as the Speckkäfern and Sexton-beetles, as being those which likewise live upon putrid substances. It is difficult to separate the Coccinellidæ from them; their external resemblance to the Erotyli is also striking.
The pentameral Raub-Lauf-and Sandkäfern form, as it were, the trunk or main stem of the Zoophagous beetles, to which are annexed likewise the pentameral Malacopterous chafers, namely, the Lampyridæ, as parasites.
But the heteromeral Cantharides have also the same soft wings and mode of life, so that they are not in a natural system, even if devoid of principles, to be separated from the Malacopterous beetles.
The nourishment of the Lamellicorn beetles appears to me to be most perfect in kind. Mould or dung can be regarded as a fully prepared aliment, or, as it were, a minced and cooked meat with greens prepared by Nature, like as Man restores it by art. Thus the lower Thricozoa, e. g. the Mice, eat the crudest vegetable substances, such as roots and seeds; those that stand higher, grass and leaves; then snails, worms, and insects; finally flesh, and last of all fruits, as the Bears and Apes. But Man lets the crude matters ferment or reduces them to rapid decomposition by cooking, whereby a mixed kind of food results, which bears obviously the greatest resemblance to dung which, as just observed, is a food cooked by nature.
According to these considerations I now arrange what have been called Beetles in the following manner into divisions drawn from philosophical principles.
The Beetles again commence, like the whole class of Flies, from below, and the inferior kinds pass therefore parallel to the Tracheo-and Dictyoptera, while the superior project above them, as was the case also in the preceding classes.
In a more remote manner also they repeat the lower classes, namely, the Protozoa, Conchozoa, Worms and Crustacea, a fact which is, properly speaking, self-evident, and which is rendered clear by the following table. We have thus:
_Order_ I. Tracheopteroid Coleoptera--_Phytophaga_. _Order_ II. Dictyopteroid " --_Zoophaga_. _Order_ III. Ceratopteroid " --_Rypophaga_.
It can also be said; the first correspond to the Worms, the second to the Crustacea, the third to the Flies.
Lastly, it may still be said; the first correspond to the Protozoa, the second to the Conchozoa, the third to the Ancyliozoa.
_Order 7. Tracheopteroid Beetles--Phytophaga._
3527. Body cylindrical, head mostly long, antennæ setiform, maxillary teeth obtuse, tarsi mostly tetrameral.
They gnaw hard seeds, leaves, and wood, and mostly live concealed. The larvæ almost or entirely apodal.
_Fam. 1. Dipteroid Beetles--Rhyncophora._
Head rhynchiform or snout-shaped, tarsi tetrameral.
_Fam. 2. Hymenopteroid Beetles, Blattkäfer._
Head tolerably short as well as the antennæ, tarsi tetrameral.
_Fam. 3. Lepidopteroid Beetles, Holzkäfer._
Head pretty short, antennæ very long, tarsi tetra-and pentameral--Borkenkäfer, Holzbohrer and Schröter.
_Order 8. Dictyopteroid Beetles--Zoophaga._
3528. Body long and depressed, antennæ short, maxillæ large with pointed teeth, feet mostly pentameral and without ungues.
They subsist upon living or dead animals, dwell usually in water or upon dry land, and run very swiftly. The one kind seize upon living beasts, and on that account constantly swarm about them, as the Raub-and Laufkäfer; the others only subsist as larvæ by suction upon other beetles or snails externally, but live, as Flies, upon leaves and flowers, as the Lampyridæ and Cantharidæ.
_Fam. 4. Neuropteroid Beetles--Water-beetles._
Like the following, but all three parts of the body closely annexed, and rudder-like fringes of hairs upon the feet--Parnidæ, Hydrophilidæ, Dyticidæ.
_Fam. 5. Orthopteroid Beetles--Prædatoria._
Maxillæ very large and pointed, antennæ setiform, wings hard and tolerably flat like the body, tarsi mostly pentameral.--Raub-Lauf-and Sandkäfer.
_Fam. 6. Hemipteroid Beetles--Parasitica._
Body pretty cylindrical, head rounded with moderate sized maxillæ, elytra narrow and soft, tarsi penta-and heteromeral.
They hang as larvæ like Mites to other beetles, and suck them; but after the metamorphosis they live upon leaves and flowers--Lampyridæ and Cantharidæ.
_Order 9. Ceratopteroid Beetles--Rypophaga._
3529. Body quadripeltate and mostly short, with hard elytra, thorax large, head small, with short claviform antennæ and blunt maxillary teeth; feet mostly furnished with ungues, tarsal joints varied. Their habitation is very varied, being both free and concealed, and they live for the greatest part on decaying excrementitious matters, such as fungi, dry rot, and even animal ordure.
_Fam. 7. Phytophagoid--Necrophaga._
Body mostly rounded, with hard and large elytra; antennæ claviform; tarsi tri-tetra-and pentameral--Lycoperdinæ, Erotyli, Coccinellidæ, Byrrhidæ, Dermestidæ, Sylphidæ. They correspond to the Worms and Tracheopterous insects.
_Fam. 8. Zoophagoid--Mulmkäfer._
Body elongated and also nearly spherical, mostly of a dark spotted colour, with hard elytra, thorax rounded, head retractile, palpi filiform, tarsal joints heteromeral or unequal in number--Helopidæ, Diaperidæ, Tenebrionidæ, and Pimeliariæ.
They live usually in dark situations, eating meal and dry-rot; have an unpleasant smell. They correspond to the Crabs, Wood-lice, and Spiders, as likewise to the Dictyoptera.
_Fam. 9. Perfect Beetles_--_Erdkäfer_ or _Lamellicornes_.
Mostly large, convex and short bodied Beetles with large thorax, broad head, lamellated antennal clubs, spinose tibiæ and pentameral tarsi. Their habitation is in dung or under the earth, where they live in the larval state frequently for years before they change; some Chafers eat also leaves of trees and flowers. They are distinguished by their striking size and singular forms, being mostly furnished with horns on the thorax or head. I do veritably believe that they may be generally interpreted as being the highest organized beetles and members of their class, although they are generally placed or arranged midway between the lower forms, while the Sand-and Laufkäfer are esteemed the most perfect.
_Tabular Co-arrangement._
The Dermatozoa now admit of being co-arranged in the following manner, from which their three Cardinal relations of Proximity, Repetition, and Serial analogy or Parallelism, admit of being distinctly recognized.
_PARALLELISM OF THE LOWER ANIMALS._
| | IX CL. PTILOTA. | A. PROTOZOA. | VII CL. WORMS. | Co. I. TRACHEOPTERA. | | | | I CL. INFUSORIA. | O. 1. _Weisswürmer._ | O. 1. _Diptera._ | | | | F. 1. Monades | F. 1. Saugwürmer | F. 1. Schmeissen | 2. Rhizopoda | 2. Tæniæ | 2. Dasseln | 3. Rotifera. | 3. Ascarides. | 3. Gölsen. | | | | II CL. POLYPI. | O. 2. _Rothwürmer._ | O. 2. _Hymenoptera._ | | | | F. 1. Hydræ | 4. Earth-worms | F. 1. Bees | 2. Alcyonia | 5. Nereides | 2. Wasps | 3. Actiniæ. | 6. Serpulæ. | 3. Terebrantia. | | | | III CL. ACALEPHÆ. | O. 3. _Sternwürmer._ | O. 3. _Lepidoptera._ | | | | F. 1. Physaliæ | 7. Holothuriæ | F. 1. Moths | 2. Beroeæ | 8. Echinidæ | 2. Spinner | 3. Acalephæ. | 9. Asteridæ. | 3. Wanner. | | | | B. CONCHOZOA. | VIII CL. CRUSTACEA. | Co. II. DICTYOPTERA. | | | | IV CL. MUSSELS. | Co. 1. KREBSE. | O. 4. _Neuroptera._ | | | | O. 1. _Röhrenmuscheln._ |O. 1. _Pfriemenkrebse._| F. 1. _Nagbolden._ | F. 1. Pholades | F. 1. Entomostraca | A. 1. Kieferläuse | 2. Tellinidæ | 2. Branchiopoda | 2. Termitidæ | 3. Cardiacea. | 3. Phyllopoda. | 3. Phryganeidæ. | O. 2. _Spaltmuscheln._ |O. 2. _Scheerenkrebse._| F. 2. _Raubbolden._ | 4. Mytilaceæ | 4. Squillæ | 4. Hemerobiidæ | 5. Arcaceæ | 5. Macroura | 5. Raphidiidæ | 6. Ostraceæ. | 6. Brachyura | 6. Libellulidæ. | | | | V CL. SNAILS. | Co. II. ASSELN. | O. 5. _Orthoptera._ | | | | O. 1. _Androgyni._ | O. 3. _Saugasseln._ | F. 1. _Saltatoria._ | F. 1. Tritoniæ | F. 1. Lernoeæ | A. 1. Achetidæ | 2. Patellæ | 2. Arguli | 2. Gryllidæ | 3. Pulmonea. | 3. Pyncnogonides. | 3 Locustidæ. | O. 2. _Dioecii._ | O. 4. _Nagasseln_ | F. 2. _Cursoria._ | 4. Capulidæ | 4. Læmodipoda | 4. Mantides | 5. Turbinidæ | 5. Amphipoda | 5. Blattidæ | 6. Buccinidæ. | 6. Isopoda. | 6. Forficulidæ. | | | | VI CL. KRACKEN. | Co. III. KOBE. | O. 6. _Hemiptera._ | | | | O. 1. _Rumpfkracken._ | O. 1. _Langkobe._ | F. 1. _Leptoptera._ | F. 1. Salpæ | F. 1. Poduræ | A. 1. Aphidii | 2. Ascidiæ | 2. Juli | 2. Cicadæ | 3. Cirripedia. | 3. Scolopendræ. | 3. Fulgoridæ. | O. 2. _Kopfkracken._ | O. 2. _Kurzkobe._ | F. 2. _Pachyptera._ | 4. Brachiopoda | 4. Acaridæ | 4. Water-bugs | 5. Pteropoda | 5. Scorpions | 5. Narrow bugs | 6. Cephalopoda. | 6. Spiders. | 6. Broad bugs. | | | | C. ANCYLIOZOA. | IX. CL. PTILOTA. | Co. III. CERATOPTERA. | | | | VII CL. WORMS | Co. I. TRACHEOPTERA | O. 1. _Phytophaga_ | VIII CL. CRUSTACEA | II. DICTYOPTERA | 2. _Zoophaga_ | IX CL. PTILOTA or FLIES. | III. CERATOPTERA. | 3. _Moderkäfer._ |
| Co. III. CERATOPTERA. | O. 3. _Rypophaga._ | | O. 1. _Phytophaga._ | F. 7. _Necrophaga._ | F. 7. _Erdkäfer._ | | | | F. 1. _Rhyncophora_ | A. 1. _Fungivora._ | A. 1. _Phyllophaga._ | | | | A. 1. Langrüssler | Scaphidiidæ | a. Melolonthidæ | 2. Kurzrüssler | Erotyli | b. Anoplognathidæ | 3. Breitrüssler. | Encidæ. | c. Rutelidæ. | | | | F. 2. _Blattkäfer._ | A. 2. _Speckfresser._ | 2. _Anthobii._ | | | | A. 1. Galerucidæ | 1. Dermestidæ | a. Lepitrichidæ | 2. Chrysomelidæ | 2. Nitidulidæ | b. Dichelidæ | 3. Crioceridæ. | 3. Silphidæ. | c. Glaphyridæ. | | | | F. 3. _Holzkäfer._ | A. 1. _Dung-feeders._ | 3. _Melitophaga._ | | | | 7. Bostrychidæ | 1. Byrrhidæ | a. Cetoniæ | 8. Elateridæ | 2. Sphæridiidæ | b. Trichii | 9. Cerambycidæ. | 3. Histeridæ. | c. Goliathi. | | | | O. 2. _Zoophaga._ | | | | | | F. 4. _Water-beetles._| F. 8. _Mulmkäfer._ | A. 2. _Coprophaga._ | | | | A. 1. _Sumpkäfer._ | A. 1. _Trachelidæ._ | a. Aphodii | a. Heteroceridæ | a. Lagriæ | | b. Parnidæ | b. Pyrochroæ | | c. Hydrophilidæ. | c. Mordellæ. | | A. 2. _Flusskäfer._ | _Serropalpidæ._ | b. Copridæ. | d. Girinidæ | d. Mycteridæ | | e. Haliplidæ | e. OEdemeræ | | f. Dytiscidæ. | f. Melandryæ. | | | | | F. 5. _Mordkäfer._ | A. 2. _Stenelytra._ | _Arenicolæ._ | | | | A. 1. _Raubkäfer_ | | | a. Pselaphidæ | a. Cistelidæ | a. Æglaliæ. | b. Scydmænidæ | b. Helopidæ | | c. Staphylinidæ. | c. Cnodalidæ. | | A. 2. _Laufkäfer._ | _Taxicornes._ | b. Trogidæ. | d. Carabidæ | d. Niliondæ | | e. Brachinidæ | e. Cossyhidæ | | f. Cicindelidæ. | f. Diaperidæ. | | | | |F. 6. _Schmarotzkäfer._| A. 3. _Tenebrionidæ._ | _Geotrupidæ._ | | | | A. 1. _Leuchtkäfer._ | | | a. Lampyridæ | a. Tenebriones | a. Geotrupes. | b Melyridæ | b. Toxicidæ | | c. Cleridæ. | c. Opatridæ. | | A. 2. _Ziehkäfer._ | _Melanosomata._ | b. Lethri. | d. Notoxidæ | d. Blapsidæ | | e. Horiidæ | e. Pimeladiidæ | | f. Cantharides. | f. Sepidiæ. | | | | | O. 3. _Moderkäfer._ | F. 9. _Erdkäfer._ | A. 3. _Lohfresser._ | | | | F. 7. Aaskäfer | A. 1. Phyllophaga | a. Lucanidæ | 8. Mulmkäfer | 2. Coprophaga | b. Passalidæ | 9. Erdkäfer. | 3. Lohfresser. | c. Oryctidæ.
_Second Province._
--FOURTH CIRCLE--SARCOZOA.
_Pisces, Reptilia, Aves and Thricozoa._
3530. These animals necessarily pass through all 4 circles, and each class therefore divides into 4 orders, or into 5, if the senses be allowed to hold good as constituting a distinct order in themselves.
Two points of departure for classification here admit of being thought of, either wholly from below, or first of all from the commencement of the present circle. In order to be convinced of this, we need only make a slight attempt. The animal series is as follows:
A. SPLANCHNOZOA.
_Circle I. Intestinal animals_ _Protozoa._
Class 1. Gastric animals Infusoria. 2. Intestinal " Polypi. 3. Absorbent " Acalephæ.
_Circle II. Vascular animals_ _Conchozoa._ 4. Venous animals Mussels. 5. Arteriose " Snails. 6. Cardiac " Kracken.
_Circle III. Respiratory animals_ _Ancyliozoa._ 7. Reticular animals Worms. 8. Branchial " Crabs. 9. Tracheal " Flies.
B. SOMATOZOA.
_Circle IV. Sarcozoa._ 10. Osseous animals Fishes. 11. Muscular " Reptiles. 12. Nervose " Birds.
_Circle V. Aistheseozoa._ 13. Sense-animals Thricozoa.
3531. Now it is here conceivable, either that the families of Fishes, Reptiles, Birds, and Thricozoa, pass parallel to all the preceding classes from the Infusoria upwards, or that they first commence with the class of Fishes.
In regard to this question the Birds and Thricozoa give the most clear and decisive answer. If we adopt the last proposition, then the Birds must follow each other thus:
1. Fish-like Birds Palmipedes. 2. Reptilian " Grallæ. 3. Typical " Gallinæ. 4. Thricozooid Birds Struthionidæ.
According to this, the Aquatic Birds would occupy the lowest rank, just as they have actually done hitherto in all other systems. The three other families too do not admit of any other position. But what is to be done meanwhile with the large majority of other Birds? They would extend or pass beyond the Thricozoa, and have thus no anatomical system whatever for their basis. Besides, Humming-birds, Sparrows and Linnets would in this way rank higher than the Gallinaceous and Struthious Birds, which some indeed believe to be the case, but without any proof, so that it is scarcely necessary to abide by such an opinion.
The Thricozoa must stand thus:
1. Icthyoid Cetacea. 2. Herpetoid Pachydermata. 3. Ornithoid Ruminantia. 4. Typical All the remaining Thricozoa.
In the systems now in vogue the Thricozoa certainly follow in this manner, and no scruple whatever is made of placing the Mice, Ant-eaters, Shrews, and Bats above Elephants, Horses, and Deer. Yet apart from all this, the former must pass beyond the Thricozoa, and would thus likewise have no other organ as a foundation than perhaps the organs of sense, of which there are but five, while the families of Thricozoa not yet provided for are much more numerous, namely, besides those named, the Seals, Dogs, Bears, and Apes.
The above methods of arrangement must be consequently cast aside without limitation or reserve, although they may have held good for hundreds of years. But what does or will not hold good in an age that is devoid of principles?
It is thus evident that the smaller Birds stand below the Aquatics, just as the smaller Thricozoa do beneath the Whales, quite apart from organic structure, which in every respect is imperfect. They must correspond, consequently, to the Asarcous animals, wherewith also their great number agrees.
If this holds good of the two highest classes, so also must it hold good of the two lower, namely, Fishes and Reptiles, and we shall thus have:
I. Protozooid Fishes, Reptiles, Birds, and Thricozoa. II. Conchozooid Fishes, &c. III. Ancyliozooid " &c. IV. Icthyoid " &c. V. Herpetoid " &c. VI. Thricozooid " &c. VII. Ornithoid " &c.
3532. But this parallelism depends essentially upon the organs, and they are called, in the language of Physio-philosophy, more correctly:
I. Intestinal Fishes, Reptiles, Birds, and Thricozoa. II. Vascular Fishes, &c. III. Pulmonary " &c. IV. Osseous " &c. V. Muscular " &c. VI. Nervose " &c. VII. Sensorial " &c.
But as these names are unused, and give or convey only the anatomical idea, but no external image, the first appellations are to be preferred.
_Tenth Class._
_Osteozoa, Glossozoa--Pisces._
3533. No one of the upper classes is in such an extensive state of confusion as that of Fishes. Nor does this result simply from the great deviations in their structure from the normal type, these being rather useful than otherwise for the purposes of classification; but mainly from the utter want of any principle which might serve systematists as a guide. Thus at one time we find them having recourse to the nature of the osseous tissue, at another to the fins, now to the teeth or to the scales, ay, even to the fin-rays, and all this because the characteristic or typical organ has never been sought after, nor the presence of such an organ as indispensable been so much as known.
3534. The characteristic organ of Fishes is the Osseous system, which is consequently the principle also of their division.
The physical nature, form, position, and number of the osseous parts must therefore be principally considered, and hence, above all, the substance or texture of the bones; the limbs also, and the maxillæ with their teeth, as well as the teeth upon the palate, upon the tongue and the branchial arches.
With regard to the component substance or texture of the bones we encounter a great difficulty. The Cartilaginous Fishes appear to belong to each other, and are also usually arranged together. Yet amongst them we find those species, such as the Lampreys, which obviously occupy the lowest grade of all Fishes, while the Sharks and Rays remind us of the Reptilia and Thricozoa, as well by their external structure as the development of their sexual parts, since they possess perfect testes, and ovaria separate or distinct from the oviducts, while they no longer deposit roe, but large ova inclosed in leathery shells, like the higher Reptilia. Now, if we separate these Fishes from the Lampreys, with whom in the scaleless tegument, the branchial foramina, and even the external form they have many points of resemblance, nothing else remains to be done than assign them the uppermost place, and so parallelize them with the Thricozoa. But one is next constrained to unite into one family the Pikes and Herrings, which perhaps admits of being done.
There belong namely to the upper Fishes without doubt the Abdominales, which are divisible into five families: the Carps, Pikes, Shads, Salmons, and Herrings. Now, if the Sheat-fishes be placed inferiorly on account of their scaleless body and amorphous maxillæ, four families will still be left, which should correspond to the Fishes, Reptiles, Birds, and Thricozoa, so that no place is left remaining for the Sharks. Now, however, the Salmons correspond decidedly to the Reptilia; and the Flying Fishes, which are ranged below the Pikes, probably to the Birds. If we unite them with the Herrings, then the Sharks may occupy the place of the Thricozoa.
This being preassumed, we can now attempt the classification. The substance or texture of the bones is of such importance, that notwithstanding the separation of the Sharks and their congeners, the other Cartilaginous Fishes must be left along with them, and range upon the lowest stage, so that they thus correspond to the Intestinal animals.
The next great distinction in the osseous system is the regular and irregular form which it imparts to the body, so that the Regular-shaped can be separated in a tolerably "tranchant" manner from the Irregular Fishes.
The regular form of a Fish is obviously the ellipse, as we find to be the case in our fresh-water Fishes, namely, the Perches, Salmons and Carps. They are collectively covered with large scales, which is therefore also a sign of their regularity.
The Irregular Fishes are cylindrical, fusi-clavi-spheri-and tubuliform, usually destitute of scales or covered with plates, scutes, and spines. Thus, since the Cartilaginous Fishes are collectively irregular in form, the Irregular Osseous Fishes must be allowed to follow them.
The greatest variety of the osseous system is shown in the limbs, especially the posterior pair, which in the other classes also are generally imperfect and make their appearance the last. In Fishes they are not divided into digits, but only into rays, which probably correspond only to the digital ungues or to feathers. An animal which has fin-rays is assuredly a Fish, for fin-rays do not occur in any Reptile.
The posterior fins change even their situation. Those Fishes in whom they are placed near to the anus, are obviously the more perfect, as the Abdominales.
In others they advance to the rear of the thoracic fins, and are even attached to what has been called the "girdle" or humerus--Thoracici.
In others they even get in front of the thoracic fins on the throat--Jugulares.
Lastly, they are actually wanting--Apodales.
In the Lampreys there are neither thoracic nor ventral fins.
The skeleton of Fishes is not simply divided to a greater extent than in other animals, but has actually a greater number of bones, such as the rays in the perpendicular fins, which are wanting in all animals, even in the Reptilia. An animal with dorsal rays must surely be placed among the Fishes, and consequently the Lepidosiren also.
The misshapen Fishes will therefore occupy the lower stages, the regular the upper, not directly by reason of their form alone; but because the other organs are also more imperfectly developed, the bones being cartilaginous, the tegument asquamous, mucous, or covered with spines, scutes, and plates; the fins wanting or abortive, or displaced from their proper situation; the head disproportionate in size to the body, the mouth wide or narrow, the eyes placed superiorly or upon the forehead.
A lower character is afforded also by very long dorsal fins, such namely as extend from the head to the tail. In the Abdominales, Sturgeons, Sharks, and Rays, the perpendicular fins are small; in the Thoracici, Jugulares, and Apodes they are, on the contrary, mostly very long.
Moreover, a lower character is a very long coccygeal or caudal fin, which denotes that the anus lies far forwards, and therefore that the tail has a great preponderance over the trunk. In the Abdominales and the Sharks the tail is short; in the Rays thin and terminated abruptly, as in Reptiles and Thricozoa. Thus the higher the animals ascend the more does the tail diminish in length.
Regard being had to all these relations, the Irregular Fishes must be viewed as those which correspond to the lower classes of animals, and the Cartilaginous Fishes will indeed make the commencement; to these are annexed the Irregular Osseous Fishes, and nearest to them indeed those with arrested ventral fins, whether wanting or placed on the throat; then come the Regular Fishes, and of these first of all the Thoracici, and next the Abdominales. In this manner we obtain four divisions.
A. Body irregular.
I. Pisces Cartilaginei. II. Stummelflosser--Apodales and Jugulares.
B. Body regular.
III. Thoracici--Tunnies, Breams, and Perch. IV. Abdominales--Carps, Pikes, Salmons and Herrings, Sharks.
Now, these divisions, having been discovered by a simple analysis or testing of facts, are to be arranged according to philosophical principles, and further subdivided, whereby the ground and legality of their existence will be recognized.
A. IRREGULAR FISHES.
3535. Body deviating from the elliptical form, devoid of scales, or covered with spines, scutes, and plates; head and tail disproportionate; fins mostly arrested.
They correspond to the unarticulate Proto-and Conchozoa; their irregular-shaped mucous or mailed body agreeing perfectly with these animals.
_Order 1. Intestinal, Protozooid Fishes._
3536. Mouth round and without maxillæ, or disproportionately narrow and wide.
There can be no doubt that the Lampreys are the lowest Fishes, since they remind us in every respect of the Worms by their naked, mucous, and lineiform body, with indistinct head, almost devoid of bones and true teeth, having a circular mouth, obliterated nostrils, puny eyes, and finally branchial cysts, which occupy a higher rank only from their opening into the oesophagus. They pass therefore parallel to the Infusoria, or rather to the commencements of the second animal series, namely, the Etozoa Ancyliozoa.
Now, although much dispute prevails concerning the division of Fishes which should be associated with the Lampreys, yet it appears to me that no others but the narrow-mouthed Fistularidæ and Pipe-fishes can follow, since they resemble them not only in their cartilaginous bones, but in the structure also of their branchiæ. They will thus truly occupy the place of the second family. Whether the narrow-mouthed Globe-fishes are likewise to be united with them or to be set up as a third family, may seem to be matter of doubt. I adopt the first course, and arrange at present the wide-mouthed or Frog-fishes and Shads in the third family.
_Fam. 1. Infusorial Fishes, Lampreys._
Body vermiform, naked, and slimy, without membral fins. Ex. Branchiostoma or Amphioxys, Myxine, Petromyzon.
In these Fishes the mouth is quite in front and round, being without maxillæ and adapted for the purposes of suction; only one nostril, but mostly several branchial foramina, which lead to cysts provided with reticular branchiæ, but without opercula.
The Branchiostomata are the smallest Fishes, not much above 1" in length, almost devoid of head, yet with traces of eyes and a nostril. The Myxinæ crawl even into the rectum of other Fishes, and live therein like Entozoa. The river or lesser Lampreys stick in the mud; the Lampreys cling fast by suction to stones, and do not draw the water in through the mouth, but through the branchial foramina themselves, like the lower animals.
_Fam. 2. Polypary Fishes, Narrow-mouthed._
Body cartilaginous, mouth having maxillæ, but unusually narrow, only one branchial foramen with immoveable operculum--Fistularidæ, Pipe-fishes and Globe-fishes.
In this family we still meet with species entirely naked, but covered also with plates, scutes, nails, and spines. The corymbiform or tufted branchiæ of the Syngnathi or the Lophobranchii still remind us strongly of the cystiform reticular branchiæ of the first family.
_Fam. 3. Acalephoid Fishes, Wide-mouthed._
Body naked or covered with plates; mouth in front and mostly unusually wide.
I here arrange in doubt the Frog-fishes and the Shads, although the last are Abdominales. But they deviate from the Carps and Pikes by their asymmetrical, naked or mailed body, the large transverse mouth, the eyes staring upwards, and the arrested branchial opercula.
_Order 2. Vascular, Conchozooid Fishes--Stummelflosser._
3537. Asymmetrical Osseous Fishes, Apodales, and Jugulares.
Among the true Osseous Fishes the Eels must undoubtedly range the lowest on account of their vermiform and asquamous body, and the want of ventral fins. To them are allied the Jugulares, namely, the Blennii and Gadidæ, as well as the Plaice; lastly, the asymmetrical Thoracici without scales, being quite naked or covered with scutes, as the Gobii and Triglæ.
_Fam. 4. Mussel-Fishes, Eels._
Body naked and serpentiform, without ventral fins.
The Eels, from their naked, cylindrical or riband-shaped body, the long dorsal and coccygeal fin, the small branchial foramina occasionally confluent beneath the neck, and by their dwelling in the mud, rank among the imperfect Fishes. The one set have soft, the other hard, fin-rays, and on that account they have not simply been separated, but even far removed from each other. This difference alone is not so great as to justify their being arranged, when the structure of the body agrees in other respects, into distinct orders. The influence of the dorsal spines upon the life and natatory or waving movement of Fishes is so slight, and its value generally, in comparison with other parts, to be taken so little into account, that a natural arrangement can never result from these appendages, which do not deserve to be called organs, and viewed as principal characters.
_Fam. 5. Snail-Fishes, Haddocks._
Jugulares without distinct scales and spines on the branchial operculum, and hard fin-rays.
These Fishes are allied to the Eels by the aborted ventral fins, the tolerably cylindrical, naked, or small-scaled body and soft dorsal fins. The viviparous Blennius bears with us its name of Aalmutter not in vain, for it resembles a shortened, slimy Eel. The Gadidæ are indeed less slimy and have in part scales, but, by reason of their lengthened form and their fins, cannot be removed far from the Eels; the same holds good of the Plaice.
_Fam. 6. Kracken-Fishes, Grundeln._
Asymmetrical Jugulares and Thoracici, with naked and mailed body, in addition to hard fin-rays.
Here belong the Gobiidæ and Triglidæ, the first whereof are usually naked, the second mailed, with roughnesses upon the head and spines on the opercula; in all the eyes placed high up.
B. REGULAR FISHES.
3538. Body elliptical, mostly covered with scales, Thoracici and Abdominales; eyes placed sideways.
_Order 3. Pulmonary, Ancyliozooid Fishes._
3539. Regular Thoracici.
The position of the abdominal fins immediately behind the thoracic is obviously a step further in the perfection of structure, and these Fishes must be therefore placed above the preceding kinds, in whom, apart from the asymmetrical form of body, the position of the fins is mostly upon the neck or advanced very close to it. The anus is still situated far forwards, and the tail is therefore mostly larger than the trunk. The dorsal fins are still very predominant. Among them we still meet with naked or microlepidal species, which in their abnormal form also remind us of the irregular Fishes, as the Tunnies and Haberdines or Stock-fish. They are therefore to be regarded as the lowest.
As is the case here in Fishes, so also in the Ancyliozoa the truly regular or bilateral body originates in them for the first time, or at least constitutes a persistent character.
_Fam. 7. Worm-Fishes, Tunnies._
Tolerably naked or microlepidal, cylindriform or very much compressed, with small head and mouth; the teeth very feeble, only like a brush; the branchial opercula without spines. Here belong the proper Tunnies and Haberdines: all of them marine Fishes.
_Fam. 8. Crustacean Fishes, Brassen._
Body perfectly regular, covered with great scales; mouth small with strong teeth; branchial opercula unarmed. Here belong the Labridæ or Lipped Fishes, Seabream and Osphromanus.
_Fam. 9. Ptilotoid Fishes, Perch._
Body quite regular with large scales, mouth of moderate size with scythe-shaped teeth, branchial opercula armed. Here belong the Scianoidæ and Percoidæ. Dwell in the sea and rivers.
_Order 4. Sarcose Fishes._
3540. Abdominales, dorsal fins small, with soft ramified rays, mostly placed far behind. Plainly regular Fishes with large scales. The head is regular, with the eyes upon its sides; the set of teeth varied; the trunk large, tail small, as are also the dorsal fins, which proceed more and more backwards to the sacrum or even the tail, a fact indicating their gradual disappearance, and therefore a sign also of greater perfection. Added to this, they are generally distributed over the whole earth in rivers and seas. Finally, it is they that yield the most nutriment to Man, which is also a constant sign of greater perfection; as is evidenced in the vegetable kingdom by the Fruit-trees or plants; in the animal kingdom by the Oysters, Snails, Sepiæ, Holothuriæ, Geese, Fowls, Cattle, &c.
_Fam. 10. Typical Fishes, Carps._
Body of the Carps covered with large scales, mostly but one dorsal fin placed pretty far back, mouth nearly edentate, supra-maxillary bones arrested, large teeth on the posterior branchial arches, or what have been called pharyngeal bones. They are for the greatest part fresh-water Fishes, and those which are most used as articles of food.
_Fam. 11. Reptilian Fishes, Salmons._
Mouth with strong teeth in the superior and intermaxillary bone; behind the radial dorsal fin there is still a fatty fin and no large scales. Here belongs the Lizard-like fish (Saurus), so called on account of its resemblance to the Lizards. Live in sea and fresh-water.
_Fam. 12. Ornithic Fishes, Herrings and Pikes._
The Herrings have teeth in the superior and intermaxillary bone; only one dorsal fin; mostly large scales.
Body of the Pike slightly scaled, furnished mostly with a small dorsal fin situated very far back, mouth full of teeth, but none of these in the rudimental intermaxillary bone. Dwellers in the sea and in fresh water.
Among the Pikes is placed the Exocoetus or Flying fish.
_Order 5. Sensorial Fishes._
3541. Bones cartilaginous, mouth opening transversely under the snout.
_Fam. 13. Thricozooid Fishes, Sharks._
Abdominales; bones cartilaginous, mouth opening transversely under the projecting snout; mostly several pairs of separate branchial apertures.
Here belong the Chimæræ, Sturgeons, Sharks, and Rays. The last ought to be held as higher in rank, partly on account of their slender tail, partly because the huge Rays, which are called Cephalopterus, have the anterior thoracic rays free and so moveable that they can seize their prey with them as with hands. All lay, with the exception of the Sturgeons, large and leathery ova, and in this approximate pretty closely to the Reptilia.
It has been already remarked, that the large Cartilaginei would not pass correctly into the others, and obviously seem to claim the highest post. At some later period the principle may probably be discovered whereupon their union with the higher Osseous fishes depends. Both are at all events Abdominales.
Eleventh Class.
_Myozoa, Rhinozoa--Reptilia._
3542. Body entirely naked or covered with scales, with distinctly separate and red-coloured muscles; two nostrils permeable throughout.
As the Muscular system is here the characteristic organ, it must be regarded principally in the division of the present class; the limbs also exhibit nowhere so great a variety as in this class, since they are in some cases wanting, while in others two only, in some four, are present. In place of fin-rays true toes have, however, made their appearance, these again indicating the greatest variety in the number of the joints; but meanwhile there are in no instance more than five toes.
The osseous system is constructed after the pattern of that of the Thricozoa, and is never furnished with dorsal rays as in Fishes.
The dental formula begins also to be regular. The teeth stand usually in the superior intermaxillary bone, and are sometimes pointed, at others obtuse; in the Ichthyosauri and Crocodiles they are even inserted by gomphosis, as in the Thricozoa. In many Serpents additional kinds of teeth occur, namely, the curved poison-teeth or fangs, which have an involuted groove traversing their concavity. In most Reptiles teeth also occur upon the palatal, but there are no longer any upon the lingual bone. In the Chelonia or Tortoises the teeth are entirely wanting, and they are also scarcely indicated in the Asquamous Salamanders and Frogs.
The os quadratum is found as in Birds; in Serpents, however, the mastoid bone has been freed, and hence the capability possessed by these creatures of expanding the mouth.
Viewed in accordance with the perfection of the limbs, the Salamanders and Frogs ought to be regarded as the highest in rank; but their scaleless tegument, their development out of spawn in the water, as well as branchiæ brings them near to the Fishes. In other respects their position is determined by the dental formula, which, as I have already shown, belongs to the limbs and thus to the motor system.
The position of the nostrils is now, throughout the class, in front of and upon the snout, being no longer situated almost on the forehead or vertex, as in Fishes. Their relation to the scales is likewise of importance for the purposes of division.
The naked Salamanders and Frogs will occupy the lowest place. By their form, absence of teeth, and mode of life, the Chelonia are allied to them.
Then follow the Serpents and Lizards with a perfect set of teeth. The Serpents are distinguished from the Lizards by their want of feet and the long bifid tongue inclosed within a sheath.
Among the Lizards apodal species occur, as the Blind-worms; but they have under the skin some pedal bones, and are, in addition to this, sufficiently distinguished from the Serpents by the short and sheathless tongue.
Among the Lizards with perfect feet there are some with small visual organs, like as in the Serpents; others with unusually large eyes, e. g. the Chameleon, Gecko, Ichthyosauri, and Crocodiles. All other Lizards, the Serpents, Chelonia, Salamanders and Frogs, have small eyes and consequently follow each other.
The Crocodile has teeth articulated by gomphosis, and consequently resembles the Thricozoa.
I divide the present class therefore into the following groups.
A. DERMAL REPTILES--SMALL-EYED.
_Order 1. Protozooid Reptilia-Kröten._
3543. Body slimy and scaleless, or maxillæ devoid of teeth.
_Fam. 1._ Infusorial Reptiles, Petromyzoid--_Caudate Batrachia, Salamanders_.
Body naked and furnished with a tail.
These cylindrical animals bear the greatest resemblance to the Lampreys, or are Petromyzoid, being provided with setaceous and scarcely discernible teeth, frequently only one pair of feet; they are developed also out of spawn in the water, and many retain the branchiæ during the whole of life.
_Fam. 2._ Polypary Reptilia, Kugelfischartige--_Stutzkröten_, _Frogs_.
Body thick and naked and without a tail, but with four feet.
The Frogs and Toads proper are likewise developed out of spawn, but soon lose their branchiæ. In their form and even the structure of their mouth they are Kugelfischartige, i. e. remind us strikingly of the Globe-fishes, among the Engmäulern or Plectognathi.
_Fam. 3._ Acalephan Reptilia, Welsartige--_Chelonia_.
Body thick and coated with scales, maxillæ quite edentulous.
The Chelonia appear certainly to occupy a pretty high rank; they lay large ova covered with a calcareous shell, but frequently live or dwell in the water, and have, in their mode of living as well as form, a striking resemblance to the naked Batrachia. Through their want of teeth they range among all the succeeding families.
_Order 2. Conchozooid Reptilia--Ophidia._
3544. Body cylindriform and scaly, teeth acuminate, tongue longitudinally bifid and inclosed within a sheath; no feet and eyelids.
The Serpents stand without doubt below the Lizards; yet it is difficult to arrange them properly into families. The poison-teeth appear to indicate a lower character, because the cranium is thereby removed or recedes from the usual type of structure, while the superior maxilla becomes very much arrested. But as there are moveable and immoveable poison-teeth, and the latter gradually pass over into the ordinary kinds of teeth, while externally also, no character has been found to distinguish the venomous from the non-venomous Serpents, it is best at present to discontinue this separation.
Then the Serpents can be brought according to the structure of their scales into 3 families. They are either of equal and small size, around the whole body; or there are plates upon the belly and tail; or finally, the plates under the latter are divided into two tablets.
_Fam. 4._ Mussel-Reptiles, Eel-like--_Schuppenschlangen_.
All the scales around the body small and of equal size; upon the belly only being somewhat larger. Here belong the venomous Hydridæ or Sea-snakes, and huge Boas or Pythons. If once the venomous Serpents could be distinguished as a distinct family, they will indeed be placed higher, and the non-venomous be separated from them. Then probably the Boas may be raised, together with their allies, as the highest family.
_Fam. 5._ Snail-Reptiles, Haddock-like--_Tafelschlangen_.
Plates under the belly, those beneath the tail being halved or divided into two.
Here belong our Colubri, but also the venomous Adders.
_Fam. 6._ Kracken-Reptilia, Grundelartige--_Schienenschlangen_.
Entire plates upon belly and tail, as in the Rattle-snakes.
_Order 3. Ancyliozooid Reptilia--Lizards._
3545. Scales, the usual kind of teeth and tongue; inferior maxillæ anchylosed in front, mastoid bone not freed, mostly feet and eyelids.
They divide likewise into three families.
_Fam. 7._ Worm-Reptiles, Tunny-like--_Schleichen_.
Body round and serpentiform, with small scales, feet rudimental, or even wanting.
Here belong the Cæciliæ, Blind-worms, and Scinci. I have united these animals, which elsewhere have stood dispersed among the Serpents and Lizards as well as the Salamanders, and placed them as a distinct family between the two former. By reason of their rudimental feet they keep simply upon the earth and bore themselves passages therein. Their motion is serpent-like, because they lie upon the belly and cannot assist themselves with the feet.
_Fam. 8._ Crustacean Reptiles, Bream-like--_Schuppenechsen_.
Four perfect feet, body covered all round with small granular scales, tongue short and hardly slit.
Here belong the Flying Lizards, the Iguanæ and Basilisks. They usually climb about on trees and seek for beetles and berries.
_Fam. 9._ Ptilotoid Reptilia, Perch-like--_Schienenechsen_.
Four perfect feet, but plates upon the belly and tail, tongue thin and bifid. Here belong the common Lizards and the "Sauvegards" or Monitors. Their body is usually depressed. They cannot climb, but run about briskly upon the ground and eat beetles, and even the higher animals. Many are a fathom in length.
B. SARCOSE REPTILIA--LARGE-EYED.
3546. These animals have tubercles, spines and plates, with four perfect feet, as in the higher Lizards, but the toes are of pretty equal length; there are no palatal teeth, and the tongue is not fissured. They are of varied size, lead a sluggish and mostly nocturnal mode of life, and have a slow pace, occurring too only in warm climates.
_Order 4. Sarcose Reptiles._
3547. Feet abnormal, being fin-and wing-shaped, adapted for climbing or clinging. They correspond quite closely to the three next animal classes.
_Fam. 10._ Ichthyoid Reptilia, Herring-like--_Ichthyosauri_.
All four feet converted into fins.
Here belong clearly the monstrous extinct animals, which formerly lived in the sea. They had gomphotic teeth almost like the Crocodile.--Ichthyosaurus, Plesiosaurus.
_Fam. 11._ Typical Reptiles, Salmon-like--_Geckos_.
Climbing feet, or retractile claws as in the Cats, and perpendicular laminæ or plates under the soles, by means of which they cling to walls. They usually dwell in houses and catch beetles--Chamæleo, Gecko.
_Fam. 12._ Ornithoid Reptilia, Pike-like--_Dragons_.
Body short and bird-like, with very long neck and a long digit, unto which an alary membrane has probably been attached.
Here belong plainly the fossilized species of a single genus known by the name of Pterodactylus. They have been hitherto discovered in different parts of Europe, and probably flew about, like the Bats, by night.
_Order 5. Sensorial Reptiles._
3548. Teeth wedged into the jaws, toes regular.
_Fam. 13._ Thricozooid Reptilia, Shark-like--_Crocodiles_.
Feet and toes of equal length, with swimming webs between them.
The Crocodiles must be without doubt regarded as the most perfect Reptilia, on account of their notched teeth and regular feet. In many respects they remind us, like the Sharks do among Fishes, of the Thricozoa.
_Twelfth Class._
_Neurozoa, Otozoa_--_Aves_.
3549. In these animals it is again shown in a striking manner, that the characteristic organ is the principle of classification. Without recourse being had to the varied structure of the head, especially of the beak, the division of Birds would not be possible, although the feet frequently yield good characters.
3550. The development of the young makes an essential distinction upon a large scale, since one great body of Birds must be fed as nestlings, whilst the other, when scarcely emerged from the egg, runs about and seeks its own nourishment. Although in the lower animals the young do not require the assistance of the parents, and, on the other hand, those of the Thricozoa are suckled for a longer time by the mother; yet still those Birds which can, upon issuing from the egg, at once nourish themselves, such as the Fowls, Geese, &c., are probably the most perfect, for they pass parallel to the upper classes of animals, and for the Ostrich to be a Thricozoon, there is indeed as little wanting as to the Crocodile.
3551. There is no doubt whatever that the Natatores in every respect, both in the structure of the feet and head as also in their mode of living and feeding, repeat the Fishes. This opinion the empirical zoologists have already adopted from the Physio-philosophy.
The same may be said of the Grallæ or Wading Birds in reference to the Reptiles. Their feet, neck, and beak are serpentiform, and associated with a frog-like body. Their mode of living and feeding is likewise amphibious. But these birds pass over so directly into the Fowls, that the latter could not be arranged elsewhere, apart even from their displaying by their domestication to Man the higher grade of understanding, which is manifested for the first time in the class of Birds.
The Bustards and Ostriches are, finally, the highest stage of Birds, and form the closest alliance with the Thricozoa.
3552. This point being once settled, it is self-evident that the Birds, which do not stand in need of being fed, occupy the uppermost place, and consequently the nestling or parent-fed Birds the lowest, i. e. the former correspond to the Sarcozoa, the latter to the Dermatozoa.
A. SPLANCHNIC BIRDS.--NESTHÖCKER.
3553. Remain, after exclusion from the egg, in the nest and are fed; neck and feet short, toes four in number and ununited, beak pointed.
By their short neck, which rarely appears longer than the head, these Birds approximate the Reptiles and Fishes, in whom also the head has scarcely separated itself from the thorax.
The uniformity of the feet and toes is likewise an inferior character, as it is found in the Dermal Reptiles and Fishes; while in the Sarcose Reptilia and Fishes the feet and fins make their appearance in the greatest variety, both as regards the form and length of the feet themselves, as the structure of the toes; and such is the case too in the higher organized Birds.
The same holds good also of the form of the head, and especially of the maxillæ and teeth, which are very uniform in the lower Fishes and Reptiles, but occur under very varied conditions in the upper kinds, exactly as in the Natatores and Grallæ, in the Gallinæ and Struthionidæ.
_Order 1. Protozooid Birds--Tenuirostres._
3554. Bill awl-shaped.
These Birds present a resemblance to each other, not simply in their mode of feeding, since they are collectively insectivorous, but also in the slenderness of their form, and in the dazzling, varied, sharply defined, and very striking colours of their plumage, as well as in their habits, for they employ their feet and tail usually for the purposes of support, and so climb about the upright stems and branches of trees.
Among them also occur the smallest sized Birds, a fact which, compared with a similar one in the Thricozoa, indicates likewise their lower grade or rank.
_Fam. 1._ Infusorial Birds, _Tree-runners or Creepers_.
Bill awl-shaped, three toes in front--Humming-birds, Tree-creepers.
The small size of the Humming-birds seems to render them the lowest in rank of the class, and by this means the system obtains a point of departure, unto which similar forms may be annexed. Their manner of feeding is rather a process of lapping, than an actual snapping with the bill; their food also, which consists of small Beetles and their larvæ, requires scarcely any operation of the bill, so that here the cibarial instruments obviously rank upon the lowest stage, and remind us of the proboscis in Flies, Butterflies, and Bugs.
_Fam. 2._ Polypary Birds, _Woodpeckers_.
Bill straight and chisel-shaped, two of the toes directed forwards and two backwards.
The Woodpeckers stand obviously a step higher, because their bill is specially active in seeking out larvæ, and their body is held securely by the toes and stiff tail.
_Fam. 3._ Acalephan Birds, _Cuckoos_.
Bill rounded, slightly arched and obtuse; feet scansorial.
These Birds are less animated or lively than the preceding ones, keep themselves more concealed, and live mostly upon caterpillars in warmer countries. Some of them lay their eggs in the nests of other Birds.
_Order 2. Conchozooid Birds--Conirostres._
3555. Bill short, thick, and straight, without a notch, three toes directed forwards.
These Birds usually perch upon branches, without, however, being able to walk upon them, since they usually hop, and must therefore make an auxiliary use of their wings. They are all of them granivorous, and in a condition both to crush the seeds with their strong bill, as well as pound them with their fleshy stomach or gizzard.
Their instruments of manducation and digestion are therefore perfect, added to which they are so allied to the subsequent order of Rapaces, that they could not be arranged lower; probably they ought even to rank higher.
_Fam. 4._ Mussel-Birds, _Sparrows_.
Bill short and coniform, without bristles; eat simply grains.
_Fam. 5._ Snail-Birds, _Crows_.
Bill long and coniform; eat grains, beetles and flesh, and pass gradually over into the Rapaces.
_Fam. 6._ Kracken-Vögel, _Parrots_.
Bill very thick and curved. The food consists of grains and fruits. Their spiritual energies or capacities are also more strongly developed.
_Order 3. Ancyliozooid Birds--Dentirostres._
3556. Bill with a notch; food consists of worms and flesh.
_Fam. 7._ Worm-Birds, _Cantores or Songsters_.
Bill tolerably long and straight; eat worms and berries.
_Fam. 8._ Crustacean Birds, _Fly-catchers_.
Bill straight, with sharp edges superiorly, point de-curved or bent downwards. Their aliment consists of insects, which they mostly snap at during flight--Fly-catchers, Shrikes, Swallows.
_Fam. 9._ Ptilotoid Birds--_Rapaces_.
Bill unciform; seize upon the Sarcozoa with their claws.
B. SARCOSE BIRDS--NESTFLÜCHTER.
3557. Run about soon after being hatched, and nourish themselves. Bill and feet very varied, the former being mostly obtuse.
These Birds do not fly much nor hop, but walk, run, or swim. In them are found all the diversities of bill and feet; the latter are mostly placed far back, so that the body is usually directed upwards.
Their food also is very varied, consisting of seeds, grass, fruits, worms, and flesh.
_Order 4. Sarcose Birds._
3558. Neck long, i. e. much longer than the head and bill, frequently longer than the body.
_Fam. 10._ Fish-Birds, _Natatores_.
Natatory feet short.
Mostly large Birds, which live upon fishes, worms, and many of them even upon herbs. Their bills are so varied, that they could represent several families, a fact which likewise speaks in favour of their higher position. It may be regarded as an instructive hint, that animals occupy a higher rank, if in them a richness or variety of forms is to be perceived. This is the case throughout the Thricozoa. The Natatores, through the structure and posterad insertion of their feet, through their closely-set plumage, which frequently presents short and scale-like feathers, by their swimming and diving, and lastly, through their fish-catching, approximate as closely to the structure and mode of living of Fishes as it is possible for a Bird to do, while still retaining the characters of its class. Many row even with the wings, and consequently use them as veritable fins.
_Fam. 11._ Reptilian Birds, _Grallæ or Waders_.
Legs, neck, and bill very long.
These Birds are a lively image of, or a composition from, the Reptilia, having a Frog's body with its long feet, and a Serpent's neck with a Tortoise's head.
They wade about in marshes to catch worms and fish; have also bills of very varied structure, yet generally very long and slender, being in some cases naked, in others covered with a skin.
_Fam. 12._ Typical Birds, _Gallinæ_.
Bill and legs shorter, the former being arched, the latter armed with strong toes for scraping.
Feed usually upon grains and worms, and live in dry situations. The Marsh-hens pass at once into the true Fowls, and these are attached, like as is no other family, through their capacity for domestication, to Man.
_Order 5. Sensorial Birds._
3559. Run only, cannot fly.
_Fam. 13._ Thricozooid Birds, _Bustards_.
Fowls with long legs and mostly diminished toes--Bustard, Cassowary, and Ostrich.
These Birds have mostly such shortened wings, that they cannot elevate themselves from the earth. In the Cassowary we find, in place of the primary feathers upon the wings, five barbless quills like so many claws. The Ostrich has a pelvis closed in front like the Mammalia. It has not incorrectly been compared with the Camel, since the Ruminant animals in general have many resemblances in common with Birds, especially in the development of the horny substance, which obviously agrees with the feathers.
_Thirteenth Class._
_Aistheseozoa, Opthalmozoa--Thricozoa._
3560. All the senses perfect, five digits; the face covered with skin and flesh, the body with hairs; mammæ present.
3561. The Thricozoa combine in themselves all the animal classes, and indeed, so far as the development by grades makes no distinction, in equal proportion.
The differences are in no class therefore so numerous and so strongly pronounced as in this; and yet one is in more doubt almost concerning the rank of the families than in the former classes. It turns out, therefore, that they are not treated according to sound or solid principles, but that these, after the manner in which Natural History is still prosecuted, are despised.
Upon viewing superficially the groups of Thricozoa, it is certainly difficult to determine, which are the lowest families, although the uppermost ones are very well known; so that here matters are the reverse of what they were in Plants. In the case of Birds the empirical inquirer into nature knows neither which are the upper, nor which the lower groups; while in the Reptiles and Fishes he fares still worse.
It is pretty generally conceded that the _Whales_ are the lowest in point of rank, because their posterior feet are wanting, and in this they certainly do depart the most from the usual or normal form of the Thricozoa; yet still it is just these animals which form the proper starting-point for the present class. Even the empirical naturalists here begin to employ physio-philosophical language, and designate these animals by the title of Fish-like Thricozoa. But, if they repeat the Fishes, then there must be Thricozoa, which stand or rank below them, and pass parallel to the Dermatozoa. Which these latter are, is not indeed to be declared until the other groups have been separated and brought into their proper station or place.
If once there are Ichthyoidal there must also be Herpetoidal or Reptile-like Thricozoa. Now, if the Whales correspond to the former, then the serial arrangement of the latter kinds is readily enough decided. To the Whales no other animals admit of being annexed but the _Swine-like_ genera, such as the Hippopotamus, Pig, and Elephant, which, through their muscular mass, almost naked skin, and residence in marshy situations, agree very closely with the Reptilia.
In the next place come the _Ruminantia_, to take their site opposite the class of Birds, with whom they correspond in their susceptibility to domestication, large ears, fine sense of hearing, and timid disposition. The horns must be regarded as the obscure metatype (nachregung) of the feathers. These three families may be aptly termed _Ungulata_ or Hoofed-animals.
Were matters to fare simply thus, then the center-building of this class would stand firm, and it would consequently not be difficult to say which animals stood below and which above it. For to the Apes are allied the Bears, to these the Dogs, Cats, and Martens, and to the latter again the Seals; all would rank above the Ungulata or Hoofed animals, and represent the proper or typical class of Thricozoa.
But the Thricozoa are Æsthetic or Sense-animals, and consequently the upper families must pass parallel to the organs of sense, if the lower correspond to the anatomical systems.
Now, if the eye has attained its maximum development in Man, the same must be said of the ear in the Apes, of the nose in the Bears, of the tongue in the Carnivora, and of the skin in the Seals.
Having so disposed of these, the only remaining Thricozoa are the Bats, Shrews, Moles, Marsupials, Sloths, and Rodentia.
In common parlance we compare the Bats with the Birds on account of their wings, unto which may be further added the large size of their ears. Their close relation, however, with the Shrews and Rodents assigns them a lower rank. They must be placed parallel with the Ptilota or Flies. The dental formula of the Bats resembles too so strikingly that of the Shrews that a rusty-grown prejudice can alone place the former in the neighbourhood of the Apes. They have obtained this post, without doubt, simply because they have but two mammæ, and these placed upon the chest.
The Moles cannot be removed from the Shrews.
The choice of position now remains between the Marsupials and Rodents. And to determine this point cannot prove difficult, for every one will place the former above the latter on account of their size, more perfect dental formula, and the hands, upon whose model the hind-feet have been in many species formed. To the Marsupials are annexed the Sloths, because several of them still possess marsupial or pouch-bones.
Viewed in this and in every other respect, the Rodentia stand or rank the lowest; and since the members of this family are much more numerous than those of any other, we may conclude that they fill up several families of the asarcose animals.
3562. The series would accordingly be as follows; first of all Rodents, then Sloths and Marsupials; with Moles, Shrews, and Bats, all as the repetition of the Dermatozoa. They are all Myoidal or Mouse-like in character. Next come the Whales, Pigs, and Ruminants as the repetitions of the Fishes, Reptiles, and Birds.
The Carnivora with the Bears, Apes, and Man, as being the proper representatives of the senses, conclude or wind up the list. They alone have a regular dental formula. We have thus--
I. Splanchno-Thricozoa; Mice, Edentata, Marsupialia, Shrews and Bats. II. Sarco-Thricozoa; Whales, Pachyderms, Ruminants. III. Æsthesio-Thricozoa; Carnivora, Seals, Bears, Apes and Man.
3563. It is here shown, just as distinctly as in the series of the classes, that no simple scale exists in the history of development, and consequently in the arrangement of animals. The Mouse-like species stand off from the rest, and then follow the entirely different Ungulata with the Pigs and Ruminants, which once again diverge in like manner, and make room for the development of the Seals, which then proceed through the Dogs, &c., in a less interrupted series up to Man.
He who marvels at this, let him take and set the table of the class-series before his eyes, and he must give utterance with us to the following words; namely, that the lower animals diverge or turn aside, and the entirely different Fishes, Reptiles, and Birds follow, which, once again diverging, make room for the development of the Thricozoa, or, in other words, the "Compendium Animalium." A perfect parallelism is thus found to exist between the classes of animals generally and the families of Thricozoa; but no linear or continuously progressive connexion is discoverable between one set and the other, but an appearance by fits or starts of new forms, just as the systems and organs also are not gradually evolved metamorphoses of one system, but sudden productions "en avant" with new tissues, forms, and functions. The animal system is a multifariously-constructed temple, with its nave, choir, chapels, and towers, while these again are present with the whole diversity of forms, which belongs to them in their several characters or bearings.
A. SPLANCHNO-THRICOZOA--PFOTENTHIERE, MAUSARTIGE.
3564. Small animals with irregular set of teeth; four feet with claws. The regular set of teeth has included all kinds of teeth, and with them four or six incisors.
A set of teeth is irregular, which has more or less than the ordinary number of incisors, in which moreover one or the other kind of tooth is wanting, or if it is separated by a breach or interspace.
The small Thricozoa divide into three orders.
The one have blunt uniform molars, two rodent or gnawing-teeth, and no canines--Rodentia.
The others have, one might say, a wholly aberrant and confused dental formula, there being at one time too few, at another too many teeth; molars uniform, with perfectly irregular incisors and canines--Sloths, Marsupials.
Others, lastly, have a tolerably regular set of teeth, presenting quadriacuminate molars, mostly small canines, and rodent-like incisors--Shrews and Bats.
_Order 1. Gedärm-, Eyer, Schleimthier-Haarthiere--Rodentia._
3565. Rodent teeth, without canines.
The dental formula of the gnawing Rats and Mice is so varied, and so devoid of influence upon their bodily structure and mode of life, that all attempts to arrange these animals in accordance with it have failed.
3566. The lower position of the Rodentia admits of being easily proved by taking to our aid the meaning of the dental system. The splanchnic or visceral teeth obviously rank lower than the membral teeth. Now, it so happens that the anterior teeth are in the Rodentia the principal organ, not only of the dental formula, but of the entire animal. Without gnawing-teeth the Mice could not maintain their existence, much less their character. They thus depend wholly and solely upon the visceral teeth, and are consequently the lowest Thricozoa.
In addition to this comes their small size, which is by no means a character to be despised, and one to which all naturalists pay regard, even while they keep it, upon the other hand, in the background; or else they must place the Elephant below the Field-mice.
3567. I divide them, as well according to structure as habit, into three groups. The one has the front and hind feet of equal length, with blunt claws for digging.
The other has similar feet, with sharp claws for climbing.
Lastly, the third has the hind feet longer than the fore for leaping.
_Fam. 1._ Infusorial Thricozoa--_Wühlmäuse._
Eyes and auditory conchæ feebly developed, feet for scraping, tail lax; live always under the ground, and eat roots and grains--Spalax, Rat, Common and Field-mouse, Beaver; mostly only three molars.
_Fam. 2._ Polypary Thricozoa--_Klettermäuse_.
Eyes, ears, and tail large, the latter being stiff and hairy, claws pointed; live mostly upon trees, eating nut-kernels and fruit; usually four molars--Squirrels.
_Fam. 3._ Acalephan Thricozoa--_Laufmäuse_.
Eyes and ears large, tail hairy, hind feet longest, claws obtuse, lamellar or plicated teeth; live upon the surface of the earth and eat grass--Hares and Guinea-pigs; mostly more than four molar teeth.
_Order 2. Ader-, Geschlechts, Schalthier-Haarthiere--Kaumäuse._
3568. Teeth and toes deviating completely from those of every other order; nor in a less degree the method of propagation.
In some instances all the teeth are wanting, in some they exceed the ordinary number, and are quite uniform; sometimes the lateral teeth only are similar, but in this case there are no canines and incisors; or else there are canines with more than six incisors, or also with rodent teeth.
In like manner the toes are irregular; the one kind being disproportionately large and the other absurdly small; some are for the most part wholly connate; in many cases there are hands posteriorly. The claws also are in some instances obtuse, in others sharp for climbing, or else asymmetrically large and unciform. Hands adapted for swimming or flight also occur.
Finally, the sexual parts are quite aberrant, being mostly very large and singularly formed; the mammæ frequently lodged in a pouch, or at least furnished with marsupial bones. The young are born as naked and immoveable embryos, and suckled for a very long time.
All this reminds us of the Conchozoa or Shell-animals as doth also their unusual covering of belts, scales, spines and long hairs.
_Fam. 4._ Mussel-like Thricozoa, _Sloths_.
Lateral and canine teeth equal in size and obtuse, incisors mostly wanting, and occasionally all the teeth; claws very large and curved--Ornithorynchi, Ant-eaters, Armadillos and Sloths.
_Fam. 5._ Snail-like Thricozoa, _Herbivorous Marsupials_.
Rodent teeth, usually with stunted proximal and canine teeth, lateral teeth level; toes mostly connate and very unequal; they live in the Old World upon roots, grass, and fruit--Wombat, Dasyure, Kangaroo, Opossum.
_Fam. 6._ Kracken-Haarthiere, _Carnivorous Marsupials_.
Mostly more than six incisors, triangular molars and large canines; live in the New World and in Australia, eating worms, insects, eggs and flesh--Vulpine Phalanger, Phascogale, Beutelratze.
The abnormal structure of the sexual parts reminds us of the same relation in the Snails and Kracken.
_Order 3. Lungen-, Fell-, Ringelthier-Haarthiere--Raubmäuse._
3569. Molar teeth mostly quadriacuminate, with a break in the series, canines and pointed incisors, or rodent teeth with lateral incisors, five toes; live upon worms and insects.
_Fam. 7._ Worm-Thricozoa, _Moles_.
Claws, sharp incisors or rodent teeth, with lateral incisors or false molars; live exclusively under the earth, and cast up the mould.
_Fam. 8._ Crustaceous Thricozoa, _Shrews_.
Paws, rodent teeth, with small lateral incisors and canine teeth. Many burrow passages without throwing up the soil.
_Fam. 9._ Ptilotal Thricozoa, _Bats_.
Alary membrane between the feet and anterior digits; pointed canine and incisor teeth.
_Order 4. Sarco-Thricozoa--Ungulata._
3570. Body large and heavy; teeth stunted, molars uniform, tolerably obtuse; feet fin-or hoof-like; mostly udders, rarely mammæ.
_Fam. 10. Ichthyoid Thricozoa, Whales._
Skin naked; no hind feet; two horizontal caudal fins; toes of the anterior feet surrounded by a common skin; no auditory conchs; posteriorly two udders. All live in the sea.
It is hardly necessary, in speaking of the Whales, to direct attention to their monstrously-developed osseous system and large fleshy tongue, as also to their correspondence with Fishes in the entire form, manner of living, and imperfect nose. Their head is still confluent with the neck, the teeth are horny plates of coalesced hairs, or uniform simple points, as in Fishes. Most of them have even dorsal fins; the two udders are hardly separated from the sexual parts.
_Fam. 11. Reptilian Thricozoa, Pigs._
Four feet with hoofs; canines and mostly also incisors; stomach simple, do not ruminate--Nylghau, Pig, Elephant, Rhinoceros, Horse. These animals love the marshes, and are through their mode of living, as also their form, similar to the larger Reptiles; or, in other words, through their colossal skeleton, with preponderating muscular mass, they are Myozoa, through the proboscis or snout, Rhinozoa.
_Fam. 12. Ornithic Thricozoa, Ruminants._
Toes bifid, surrounded by a hoof; above there are rarely incisor and canine teeth; udders behind; stomach fourfold; they ruminate. The horn-formation indicates a relationship with feathers; the want of incisor teeth, large ears, and timidity of disposition that of the family with the class of Birds.
Both families enter into connexion with the Whales through the size of their body, the structure of their feet, form of the head and disposition towards water and mud. They are quadrupedal Whales, which have come out of the water, and adopted a manner of living like the Amphibious Reptiles and Grallatorial Birds.
_Order 5. Æsthesio-Thricozoa--Unguiculata._
3571. Here for the first time an equiponderance of the sensorial organs makes its appearance and along with it therefore a resemblance between the animals, which is no more interrupted by such strange or odd forms, as in the preceding orders.
All of them have divided toes with claws or nails, and all kinds of teeth with multiacuminate enamelled molars.
3572. No doubt can exist about the animals belonging to the present order; they are the Apes, Bears, and Carnivora generally, as Seals, Cats, Dogs, Martens, and such like beasts.
I also cherish no doubt concerning the rank of the two first families, namely, the Apes and Bears, although they have been separated in a strange manner by the interposition of the Bats and likewise the Shrews. At some future time one will not believe that the Bats and Shrews were once placed next to the Apes.
3573. Doubt, however, may exist concerning the rank of the Carnivorous or Rapacious animals; so that here the principles of our philosophy must be brought to bear in our behalf. I regard then these animals as the highest representatives of the sensorial organs. By this step three families at once take their proper positions, viz.:
Man upon the station or rank of the Eye.
The Apes on that of the Ear.
The Bears upon the post of the Nose.
Difficulties are consequently presented in regard only to the Rapacious animals, but which are removed, so soon as the three families just named are parallelized with the others thus:
Bone, Tongue, Whales.
Muscles, Nose, Pigs--Bears.
Nerves, Ear, Cattle--Apes.
Senses, Eye--Man.
Now the Seals range of themselves next the Whales.
There are thus left the Rapacious animals proper, which as Dermatozoa or Sentient animals must consequently correspond to the Splanchnozoa. In favour of this, evidence is afforded not only by the particular use made of their feet, but their great number also, which can admit of comparison with no individual family or order.
Now, however, there are three orders of Murine animals.
1. Rodent Mice--the Rodentia proper. 2. Chewing Mice--the Sloths and Marsupialia. 3. Rapacious Mice--the Moles, Shrews, and Flitter-mice, or Bats.
In like manner do the rapacious Carnivora divide into three groups.
1. The mostly sneaking and scansorial Martens and Viverræ.
2. The sneaking and fossorial Skunks, Gluttons, and Badgers, with soles and blunt claws.
3. The high-legged Digitigrades, as Dogs, Hyænæ, and Cats.
As I have given up the dental formula in the Rodentia as a means of division, so now it seems to me that it must be abandoned also in the Beasts of Prey. The whole appearance of them and their mode of living, which is still the main point in view, obviously directs us more towards consideration of the feet, than of the dental formula. The Marten or Weasel kind were formerly compared with the Mice, and called on that account Mustela. They cannot be regarded otherwise than as the lowest in rank.
Unto them are obviously annexed the short-legged Civets, despite their cunoidal set of teeth. Many have half soles or pads under the feet. With these again the Fox-like animals, notwithstanding their viverrine dental formula, enter into alliance. I believe that I have rightly parted the Badger from the Bears, and rightly done it too in this place.
The highest are without doubt the Dogs, Hyænas, and Cats, with their long and upright legs, not to speak of their mental faculties. We accordingly obtain the following arrangement.
_Fam. 13._ Dermal Thricozoa--_Carnaria_.
All kinds of teeth included; six broad incisors, a longer canine, two to three small false molars, a large carnivorous tooth and large Querzahn, and frequently also the Kornzahn. Mostly five separated toes, with nails resting upwards, and either sharp or obtuse feet, occasionally soled. They eat flesh, mostly that of warm-blooded animals, and kill their prey themselves.
_Alliance 1. Rodent-like Unguiculate Animals--Schleicher._
Feet short and oblique, mostly sharp claws without entire soles.
_1. Mole-like Animals--Martens._
Sharp claws without soles, and no Kornzahn--Martens and Otters.
_2. Squirrel-like--Civets._
Sharp, curved, and mostly retractile claws, superiorly the Kornzahn--Civet, Paradoxurus.
_3. Leporine or Hare-like--Ichneumons._
Straight claws with half soles--Ichneumon, Ryzæna.
_Alliance 2. Kaumausartige--Fossores._
Large and straight claws with soles, no Kornzahn.
_1. Sloth-like Animals--Mephitic Animals._
Dental formula, like that of the Marten--Skunk, Mydaus.
_2. Wombat-like--Gluttons._
Dental formula, like that of Marten, but behind the carnivorous is a small tubercular tooth which is wider, being broader than long.
_3. Opossum-like--Badgers._
The tuberculous tooth larger than the carnivorous tooth, and nearly quadrangular.
_Alliance 3. Raubmausartige--Digitigrada._
Legs high and upright; no soles.
_1. Mole-like--Dogs._
Claws blunt, the Kornzahn both in upper and lower jaw.
_2. Shrew-like--Hyænas._
Claws blunt; no Kornzahn but a small Querzahn.
_3. Bat-like--Cats._
Claws sharp, curved, and retractile; no lower tubercular tooth, but a small upper one.
_Fam. 14._ Lingual Thricozoa, _Seals_.
Feet fin-shaped, the hinder pair stretched out, dental formula complete, but the lateral teeth tolerably even, and six or four incisors; tongue mostly somewhat slit. They correspond to the Whales.
_Fam. 15._ Nasal Thricozoa, _Bears_.
Nose elongated into a snout, walk upon soles; all kinds of teeth, of which, however, the carnivorous or tearing tooth is similar to the grinders, six incisors.
The slower gait, originating from their walking on the soles of the feet, with the less pointed molars, assign to these animals a less rapacious habit of living. They therefore kill no large animals, and are satisfied with worms and even roots, fruit, and honey. They repeat the Pigs.
_Fam. 16._ Nasal Thricozoa, _Apes_.
The ears begin to acquire the human form, as do even the teeth; never more than four incisors, but a longer canine; hands both fore and aft.
These animals live upon fruits and beetles, and are by their scansorial feet destined to live upon trees. Their varied, piping, and sonorous cry is a property of the sense of hearing, and along with it the larynx also usually obtains a stronger amount of development.
3574. It seems that every family of Thricozoa contains five genera, and that these accord with the organs of sense. In many families this relation is at least striking, e. g.
Among the Pigs the Elephant is obviously characterized by the nose, the Hippopotamus by the skin, the Pig by the dental system; and thus by the sense of taste, the Rhinoceros by large ears, the Horse by the eyes, thus
1. Skin Hippopotamus. 2. Tongue Pig. 3. Nose Elephant. 4. Ear Rhinoceros. 5. Eye Horse.
In like manner do the Ruminantia or Cattle become distinctly marshalled according to the five senses. The Camel recedes from the others by its simply uplying hoofs, the Musk-deer by its monstrous canine; the Goat is distinguished by its fine sense of smell, and the Giraffe by its large ears, the Ox by the large and beautiful eye, thus
1. Hautrind Camel. 2. Zungen- Musk-deer. 3. Nasen- Goat. 4. Ohren- Giraffe. 5. Augen- Ox.
In the Bears also we are confronted by this principle of classification.
The Skin-bear is the Common Bear. The Tongue- " is the Raccoon. The Nose- " is the Nasua. The Ear- " is the Arctitis. The Eye- " is the Cercoleptes.
After this principle had been so strikingly verified in several families, I proceeded with confidence also to those which were more difficult, and it resulted that each consists only of five genera severally distinguished by the predominance of some one organ of sense. This method has been carried out throughout my "Allgemeine Naturgeschichte."
_Fam. 17._ Ophthalmozoa, _Man_.
Superiorly or in front hands, inferiorly or behind soles.
3575. Here all the senses enter for the first time into a state of perfect equiponderance or proportion. Skin naked, and therefore a perfect organ of feeling; feet and hands differently constructed for progression and manipulation; tongue and lips fleshy, while the latter have hitherto been only tegumental; all the kinds of teeth different, but still very similar, being of equal height and nearly equal size; nose elevated by its whole length from the face, and fleshy; ears oval, laid close against the head and having regular windings or convolutions; eyes directed forwards, with perfect eyelids, and moveable in all directions.
3576. Man by the upright walk obtains his character, namely, that of bodily freedom, for his hind feet take the place of all the four feet of other animals, by which means the hands become free and can achieve all other offices, the feet alone serving to support the body.
He is the only animal that surveys with the axes of the eyes borne parallel the most extensive horizon. All animals whose eyes look higher up or above the ground, as the Horse, Elephant, Ostrich, and such like creatures, have eyes directed sideways.
3577. With the freedom of the body has been granted also the freedom of the mind. Man sees everything, the whole universe, while the animals can only view individual parts thereof, two of these even invariably appearing different, so that the images seen by them are never reduced to unity.
3578. There is only one human family, only one human genus, and only one species; and this just because Man is the whole Animal Kingdom.
3579. But yet there are five kinds or varieties of Men, according with the development of the sensorial organs.
1. The Skin-Man is the _Black_, African. 2. The Tongue- " is the _Brown_, Australian--Malayan. 3. The Nose- " is the _Red_, American. 4. The Ear- " is the _Yellow_, Asiatic--Mongolian. 5. The Eye- " is the _White_, European.
_Co-arrangement._
3580. The classes of Sarcozoa pass or rank parallel to each other in the following manner:
| CLASS X. | CLASS XI. | CLASS XII. | | | | | Anatomical | Osteozoa, | Myozoa, | Neurozoa, | _Systems_. | _Fishes_. | _Reptiles_. | _Birds_. | | | | | I. _Intestinal._ | _Ord._ I. | _Ord._ I. | _Ord._ I. | | | | | |F. |F. |F. | Stomach--Infusoria |1. Lampreys |1. Salamanders |1. Tree Creepers| Intestine--Polypi |2. Plectognathi|2. Frogs |2. Woodpeckers | Absorbents--Acalephæ.|3. Shads. |3. Tortoises. |3. Cuckoos. | | | | | II. _Vascular._ | _Ord._ II. | _Ord._ II. | _Ord._ II. | | | | | Veins--Mussels |4. Eels |4. Schuppen- |4. Sparrows | Arteries--Snails |5. Haddocks |5. Tafel- |5. Crows | Heart--Kracken. |6. Grundeln. |6. Schienens- |6. Parrots. | | | chlangen. | | | | | | III. _Pulmonary_. | _Ord._ III. | _Ord._ III. | _Ord._ III. | | | | | Skin--Worms. |7. Tunnies |7. Schleichen- |7. Singing-birds| Branchiæ--Crabs |8. Breams |8. Schuppen- |8. Fly-catchers | Tracheæ--Flies. |9. Perch. |9. Schienen- |9. Hawks. | | | Echsen.| | | | | | IV. _Flesh._ | _Ord._ IV. | _Ord._ IV. | _Ord._ IV. | | | | | Bones--Fishes |10. Herrings |10. Icthyosauri |10. Geese | Muscles--Reptiles |11. Salmons |11. Geckos |11. Herons | Nerves--Birds. |12. Pike. |12. Pterodactyli.|12. Fowls. | | | | | V. _Senses._ | _Ord._ V. | _Ord._ V. |_Ord._ V. | | | | | Thricozoa. |13. Sharks. |13. Crocodiles. |13. Bustards. |
| CLASS XIII. | | | Anatomical | Sense-animals,| Aisthesio- | Sense-Organs. _Systems_. | _Thricozoa_. | _Thricozoa_. | 1. _Tegument._ | | 13. Mausartige | I. _Intestinal._ | _Ord._ I. | 1. Wühlmäuse | I. _Ovum._ | | | |F. |F. | Stomach--Infusoria |1. Wühlmäuse |1. Martens |Vitellus--Infusoria Intestine--Polypi |2. Kletter- " |2. Civets |Albumen--Polypi Absorbents--Acalephæ.|3. Lauf- " |3. Ichneumons. |Envelopes--Acalephæ. | | | II. _Vascular._ | _Ord._ II. | 2. Kaumäuse | II. _Sex._ | | | Veins--Mussels |4. Sloths |4. Skunks |Spawn--Mussels Arteries--Snails |5. Beutelhasen |5. Gluttons |Milt--Snails Heart--Kracken. |6. Beutelmarder.|6. Badgers. |Kidneys--Kracken. | | | III. _Pulmonary._ | _Ord._ III. | 3. Raubmäuse | III. _Limbs._ | | | Skin--Worms. |7. Scheermäuse |7. Dogs |Papillæ--Worms Branchiæ--Crabs |8. Shrews |8. Hyænas |Feet--Crabs Tracheæ--Flies. |9. Bats. |9. Cats. |Wings--Flies. | | | | | | IV. _Cephalic IV. _Flesh._ | _Ord._ IV. | | Senses._ | | | Bones--Fishes |10. Whales |14. Seals |2. _Tongue_--Fishes Muscles--Reptiles |11. Pigs |15. Bears |3. _Nose_--Reptiles Nerves--Birds. |12. Ruminants. |16. Apes. |4. _Ears_--Birds | | | V. _Senses._ | _Ord._ V. | | | | | Thricozoa. |13. Aisthesio- |17. Man. |5. _Eyes_ | | | --Thricozoa. | Thricozoa.| |
3581. By the accompanying table the parallelism of the different families is recognized, as well as their remaining relationships, both among themselves as also with the Asarcous animals, if their table at p. 614 be compared with it, and which could not for want of room be inserted here.
3582. It is, moreover, proved from this table that the classes stand one above the other, but yet that each recommences from below, so that the lower animals of a higher class are more stunted or rudimental than the upper ones of a lower class. Thus the Salamanders are more rudimental, that is, they have organs more imperfect than the Sharks; the Tree-creepers are more rudimental than the Crocodiles, the Mice than the Fowls and Bustards.
Nevertheless these stunted animals stand higher than those of the lower classes, because they are characterized by a higher organ.
What holds good of the classes holds good again of the orders and families also. The lower animal of a consecutive family is again more rudimental than the upper one of the antecedent family. Thus the Ornithorynchus is more rudimental than the Beaver, the Shrew-mouse more rudimental than the Opossum and such like creatures.
In the highest families the equiponderance is first restored, and the lowest Man is still higher than the uppermost Ape.
IV. PSYCHOLOGY.
FUNCTIONS OF THE ANIMALS.
3583. The present section treats concerning the functions of the Whole animal, just as the preceding one did of those belonging to individual organs. It is at bottom the psychological part of Physio-philosophy. The functions are so numerous and difficult also to arrange, that I place this section here, to point out rather its situation than to follow out or trace its development.
3584. All the functions of an entire animal are spiritual or sensorial functions; at least they are conditionated by the senses, and I will also speak of them only in this respect. The mechanical and chemical functions have been already comprehended in the physiological part. The senses only make their appearance gradually in animals, and with them also the spiritual functions.
A. FUNCTIONS OF THE DERMATOZOA.
1. _Enterozoa or Oozoa._
3585. These animals are governed chiefly by a passive sense of feeling, from their consisting for the most part of a naked, homogeneous, and gelatinous integument, and living in water, an element wherein the other senses can be but slightly active.
3586. Their sense of feeling stands upon the lowest grade, since it is only the sensibility of the tegument, there being no articulated organs of touch, so that it consists only in the discrimination of an opposing object.
The ability or power to discriminate is not yet consciousness; for unto this a reflexion upon the object discriminated is necessary.
The Infusoria, Polypi, and Acalephæ, simply feel that something else is there, but they are so completely imprisoned in this feeling that they are unable to submit the same to an internal process of comparison.
3587. By reason of this inability to compare their own feelings, not a trace is left unto them of internal change; so that these creatures are truly devoid of memory or recollection.
The Infusoria have only sensation, nothing else; they are therefore in ceaseless motion. They are actually capable of nothing but moving and eating. Of all other spiritual functions they are utterly devoid.
3588. Their spiritual life is in some degree a mesmeric condition. Destitute of the senses of seeing, hearing, smelling and tasting, they feel every thing, or, properly speaking, perform all these functions at _one and the same time_, and by _one_ organ, the visceral mass. By mesmerism they find their food, perceive the light, and become transparent unto themselves, just as they are really in a physical point of view. For they are only viscera or visceral nerves.
_Development of the Mind._
3589. The mind, just as the body, must be developed out of these animals. The human body has been formed by an extreme separation of the neuro-protoplasmic or mucous mass. So must the human mind be a separation, a memberment of infusorial sensation.
3590. The highest mind is an anatomized or dismembered mesmerism, each member whereof has been constituted independent in itself.
The skeleton of this dissected mind, when scientifically represented, would be the science of the mind, i. e. Philosophy, properly so called.
Pneumato-philosophy is the likeness of Physio-philosophy. For spirit is only the tension of nature, and nature only the spirit set in motion.
The philosophy of spirit must develop itself out of the philosophy of nature, as doth the flower out of the stem. For nature is the spirit analyzed and at rest, which we can handle at our pleasure. It does not appear only for an instant; but as stone, air and such like entities, abideth always, as if to solicit and preserve us for its investigation.
A Philosophy or Ethicks apart from Physio-philosophy is a nonentity, a bare contradiction, just as a flower without a stem is a non-existent thing.
3591. As many essential members as Physio-philosophy hath, into so many must Pneumato-philosophy also divide, and this too so exactly that the two shall cover each other.
The reason why one has hitherto rambled about in Pneumato-philosophy without ballast and without compass, depends solely upon the disregard which has been paid to the science or knowledge of Nature. It is in fact not difficult to understand how impossible it must be, from observations made upon the rapidly evanescent phenomena of the spirit, to thence abstract a system of the laws in conformity wherewith this spirit manifests itself or acts. Spirit is nothing different from Nature, but simply her purest outbirth or offspring, and therefore her symbol, her language. With such a basis as this, we shall no longer pursue the _ignes fatui_ of the mind, but first of all endeavour to banish them into the provinces or realms of Nature, and there co-ordinate them in conformity with her laws; then for the first time shall we recognize the flaming lights of the mind and the divine voices, which all matter proclaimeth through the speech of Man.
He, who were once in a condition to reveal or disclose this conformity of Nature's phenomena with those of Spirit, will have learnt the philosophy of the latter.
2. _Functions of the Vascular or Sexual Animals._
3592. These animals are no longer merely sentient, 'clairvoyant' Acalephæ, because, in addition to the nervous mass, they are provided with other systems, such as the sexual and vascular, with the special organs of digestion and taste.
These three or four organs must also resolve themselves into three spiritual functions; the vascular system furnishes special organs of feeling and therewith a voluntary sense of feeling; the intestine and chiefly the liver is now the cardinal organ, and will therefore execute the mesmerically percipient functions.
3593. In the liver the faculty of anticipation and foresight, with melancholy, choleric passion, and anger, appear to reside. Encephalic thought is reflected in it.
The liver is the soul in a state of sleep, the brain is the soul active and awakening. In it the spirit broods unconsciously for years, and then breaks forth fearfully, as capriciousness, tyranny and sorrow, but also as earnestness and strength.
Circumspection and foresight appear to be the thoughts of the Bivalve Mollusca and Snails.
Gazing upon a Snail, one believes that he finds the prophesying goddess sitting upon the tripod. What majesty is in a creeping Snail, what reflection, what earnestness, what timidity and yet at the same time what firm confidence! Surely a Snail is an exalted symbol of mind slumbering deeply within itself.
The old artists must have felt this signification, as in many of their representations they have introduced a Snail. One can hardly think, that in so doing they wished to express such common and lascivious ideas, as are at present manifested openly or secretly by our daily enjoyments.
3594. The intestine must moreover be concerned with the sense of taste. Taste, however, leads to voracity, gluttony, daintiness, sluggishness and drowsiness.
3595. Taste in union with the sexual function is the expression of venery or lust.
This is indicated by the secretion of slime, by the monstrous size too of the sexual organs, and by their androgynism, which enables either individual during copulation to enjoy the delicious feeling, belonging to the male and female, either at once or alternately. Their food also appears to be selected from a feeling of desire.
Circumspection in feeling, dainty voracity, and immoderate lust appear to constitute the spiritual character of the Malacozoa, especially of the Snails.
_3. Functions of the Respiratory or Arthric Animals._
3596. The Insect is mainly an aerial and motor organ, and therefore its spirit is also of an aerial and motor kind.
The respiratory process produces strength, and this again courage, both which are the distinguishing properties of Insects. The Insect is the strongest and boldest animal upon the earth.
Health, plenitude of life, generosity, nobleness, and heroism dwell in the thorax.
3597. But besides these virtues the spirit of the thorax is also that of smell. Insects have an excellent sense of smell, the spirit of which is _cunningness_ and treachery, wherein no animal will easily surpass them.
3598. The Insect has moreover a spirit of motion or versatility of the tactile sense, which is displayed in the representation of symmetrical figures. This faculty proceeds especially from the creative sexual functions--as mechanical or _artistic instinct_.
All spirit of motion launches out into mechanical instinct. It disappears in all classes of animals, which chiefly correspond to the sex and belly, as, e. g. in Fishes and Reptiles. On the other hand, in the moveable thoracic animals or Birds, the mechanical instincts at once re-appear.
Mechanical instinct and dexterity of limbs run parallel together.
The dexterity of the limbs taken up into the spirit is an _art-sense_.
B. FUNCTIONS OF THE CEPHALOZOA.
3599. Here the head is for the first time placed in a perfect condition, and hence an antagonism arises for the first time between head and trunk.
The Cephalozoon no longer distinguishes nature and self only like the acephalous and amnemonic animals; but it distinguishes even its body from its head, because the Fish has begun to be a double animal.
3600. The Cephalozoon hath _consciousness_; consciousness of its condition, of its body, but not of its head and the operations therein. It has _no self-consciousness_.
3601. As soon as an animal contemplates a part of its body, of its world, and hath consciousness in a general sense, it has also _memory_. For memory is a repetition of _its own_ condition, not the reiterated feeling of a foreign object.
The Acephalous or anencephalic animals have therefore no memory, because they live only in opposition to the world, but never in antagonism to themselves. Every perception is therefore a new one for them, because it is always an actual object which excites them. Whether Insects have memory, has not yet been made out.
3602. The brainless animals have no ideas, and naturally so, because they have no consciousness.
It would appear likewise that they do not feel pain.
The Cephalozoa have ideas, and quite certainly pains, because they become partly an object unto themselves.
4. _Functions of the Osteo-or Glossozoa._
3603. The Fish's head is the lowest, and therefore its mind also will manifest only the first function, that ranks above the mind of the Acephalozoa, the _memory_.
With this memory, however, all the spiritual functions, exhibited by the preceding classes, but chiefly mesmerism, have been bestowed.
3604. Fishes are again provident, zealous animals, that, drawn together by mysterious bands, make the longest voyages, wherein they ascend and descend rivers, knowing how to find their prey over miles in extent.
All the mechanical instincts are, on the contrary, obliterated in them, as being fingerless, finned animals. Their principal business is propagation--Pelvic animals.
3605. Gluttony is the principal character of the Glossozoa, in so far as taste stands upon the lowest stage. Touch and taste are only motion and deglutition.
Smell becomes evidently stronger.
The ear still ranks upon the lowest stage, but yet they hear closely. In other respects they are mute, and exhibit all the consequences of aphony.
3606. Fishes are of the Phlegmatic temperament.
5. _Functions of the Myo-or Rhinozoa._
3607. Reptiles are Cephalozoa with well-developed lungs and nose, and are therefore gifted with voice.
3608. To memory comes the art of _ambuscade_ or lying in wait, a property of the olfactory sense, the application of memory. The circumspection of the Snails passes over into ambuscade and surprise.
With this character the higher perfection of the belly or of the digestive system, along with the poisoning saliva, is in parallel accordance. To take by surprise and to poison are acts of _one_ series.
3609. Reptiles appear enabled to _reflect_, i. e. several reminiscences are at their bidding. Now, the comparison of these constitutes reflection. They are therefore more sagacious than Fishes, docile and in some measure susceptible of instruction.
3610. The courage, which they have probably obtained through their aerial respiration, passes over more into impudence and sauciness.
3611. Reptiles are of the Melancholic crasis.
6. _Functions of the Neuro-or Otozoa._
3612. The spirit of the thorax and limbs is here predominant, whence comes the restlessness and mechanical instinct of the Bird.
The motor sense, the ear, is the prominent one. The ear, however, is partly the sense for the Indefinite, partly for the demolition of matter. The auditory spirit is _fear_.
But it passes over into joy, passion, levity, if it perfectly perceives tones.
3613. With the ear and the moveable organs of voice originates a kind of language, which is in a condition to express a multitude of sentiments. The language of Birds hath not a few tones, and expresses not a few passions.
3614. The Bird knits or associates for the first time with some completeness a sense, or definite sensation, to a simple tone. The Bird hath for the first time _signs_, or symbols, which are not the things themselves, but only _signify_ or mean them.
The Bird understands the relation of the spiritual expression to the organ or matter. It comprehends a connexion where none is materially, but which is only imparted by the idea.
The capacity to understand the thing in the image or idea, I call _imagination_, and of this Birds furnish us with very definite proofs.
Birds can therefore dream.
3615. Reptiles and Fishes appear to have no ideas because they have no _signs_, or tones, indicating the resolution of the organ into mind. For the tone is none other than the ghost of the organ or animal. Fishes and Reptiles do not indeed dream.
3616. The Bird, however, appears to get no further than to mere images or ideas. The conception is wanting unto them.
It has therefore no sense of shame. But possesses in a full degree circumspection, desire of imitation, comparison.
3617. To the Bird it is not simply the sensation of its body that becomes, like a foreign product, objective; but its own product, its voice, as something already distinct from its own mind.
3618. It is clear, that if all the sensorial functions were to become objective to the Bird, it would be self-apparent and resolve itself into self-consciousness. Thus does self-consciousness sprout forth gradually with the sensorial functions.
3619. Birds are of the Sanguineous temperament.
7. _Functions of the Æsthesio-or Ophthalmozoa._
3620. All the faculties hitherto mentioned occur here. The soul of the eye is still associated with them, and therewith the faculties of perception, understanding and conception appear to be bestowed.
One cannot refuse understanding to the Thricozoa. The actions of the Dog, Horse and Elephant, do not admit otherwise of being conceived; nor also the shame and pride, fidelity, animosity, desire of revenge, and yet many other properties exhibited by these animals.
But it is an understanding without self-consciousness, if we may so venture to express ourselves; an understanding of many signs, but devoid of any combination and separation of these signs; in a word, there is no faculty of _judgment_.
The Thricozoa are Choleric animals.
8. _Functions of the Panæsthetic Animal._
3621. If finally all its organs become objective to the animal, through contemplation of the universe, through hearkening of the animal restored into symbols; it thus contemplates itself, is apparent unto itself, and is quite conscious of itself.
Then is the animal equivalent to the whole animal kingdom and to the universe.
3622. The universal spirit is _Man_.
In the human race the world has become individual. Man is the entire image or likeness of the world. His language is the spirit of the world. All the functions of animals have attained unto unity, unto self-consciousness, in Man.
3623. The thorough penetration of all the animal's symbols, the comparison of all the world's symbols, and thus free comparison is _Reason_.
The understanding compares only the symbols of sounds, of men; but the reason compares also the symbols of light, of the world.
Reason is world-understanding; the understanding is animal reason.
All the mental functions of animals have become ratiocinative in Man.
Feeling is in him consciousness, consciousness is self-consciousness, the understanding is reason, the passion, freedom, the mechanical instinct artistic sense, the comparison science.
3624. The spirits of the senses are Art-spirits, the spirit of reason is the spirit of Science.
_Art._
3625. Art is the representation of the senses in Nature.
3626. The sense is, however, the last or ultimate design of Nature.
Art is consequently the representation of _Nature's design_. (Ed. 1st, 1811. § 3517.)
3627. That is beautiful which represents the will of Nature.
But non-beautiful is that which represents real Nature by means of art.
3628. Art is one universal business. Beautiful is that, which represents the world within a fragment or portion of the world.
3629. There is also one natural beauty--unconscious formation of the world's laws.
3630. The highest natural beauty is the universal portion of nature, i. e. Man.
Man expresses the ultimate goal or purpose of Nature's design.
3631. The terminus or goal of Nature is, in Man to revert again into herself. The human countenance most perfectly repeats the trunk, and again reverts wholly and actually into the trunk. That human countenance is beautiful, in which the vertebral column runs back again parallel with the vertebral column of the trunk. The facial vertebral column is the nose.
3632. The face is _beautiful_, whose nose runs parallel to the spine.
No human face has grown unto this estate, but every nose makes an acute angle with the spine. The facial angle is, as is well known, 80°.
What as yet no Man has remarked, and what is not to be remarked either without our view of the cranial signification, the old artists have felt through inspiration. They have not only made the facial angle a right angle, but have even stepped beyond this, the Romans going up to 96°, the Greeks even to 100°.
Whence comes it, that this unnatural face of the Grecian works of art, is still more beautiful than that of the Roman, when the latter comes nearer unto Nature? The reason thereof resides in the fact of the Grecian's artistic face representing Nature's design more than that of the Roman; for in the former the nose is placed quite perpendicular, or parallel to the spinal cord, and thus returns whither it has been derived.
3633. He who paints or otherwise copies Nature in a purely mechanical manner, is consequently a bungler; he is devoid of ideas, and imitates her no better than a bird does song, or an ape the postures of the human body. The province of Art is alas! not yet understood.
3634. In Man all the beauties of nature are associated or combined.
3635. Thus, Nature can be beautiful, in so far as she represents the individual ideas of Man.
3636. There are only two art-senses, the eye and the ear; and thus but two departments of art, the Plastic and Sonant, or that of _form_ and of _motion_.
3637. The province of form represents the material universe in its ideas, its design, and thus in its freedom.
3638. The representation of the bodily universe in the ideas is the _Architectural art_. (Ed. 1st 1811, § 3533.)
3639. The representation of heaven in the Plastic, is the church architecture.
The temple or church is the art of heaven.
3640. The representation of the planet in the Plastic is the house.
The house is the art of the planet.
The architectural art is the cosmical art.
3641. The representation of the Individual is the _Sculptor's art_.
The sculptural art represents the Terrestrial, and in its highest estate Man. It is the _Heroic art_.
3642. This art rendered manifest in matter, or repeated in light, is Painting.
Painting represents the symbol, the naught of the world, the heavenly, and also in its lowest estate a Spiritual.
Painting is the art of religion, the _Sacred art_.
Painting is the art of heathens, whose deities are men; Painting is the art of Christians, whose Men, being sacred or holy, are gods.
God can be painted, but not formed or sculptured.
3643. The art of motion represents the material and the spiritual motion.
3644. The representation of the world's material laws of motion is the _Dance_.
3645. The representation of the motion of individuals is the _Histrionic art_.
3646. The representation of the world's spiritual laws of motion, of the laws of the dance, is _Music_.
3647. The spiritual representation of the Histrionic is the _Poetic art_.
_Science._
3648. The representation of the rational world is Science.
3649. The first science is the _Science of Language_, the architecture of science, the earth.
3650. The second science is the _Art of Rhetoric_, the sculpture of science, the river.
3651. The third science is _Philosophy_, the painting of science, the respiration or breath.
This, like painting, divides into a number of branches, whereof the art of government is the highest.
3652. The fourth science is the _Art of War_, the art of motion, histrionism, music, poetic art of science, the light.
As in the art of poetry all arts have been blended, so in the art of war have all sciences and all arts.
The art of War is the highest, most exalted art; the art of freedom and of right, of the blessed condition of Man and of humanity--the Principle of Peace.
THE END.
End of Project Gutenberg's Elements of Physiophilosophy, by Lorenz Oken