Discourses: Biological & Geological Essays
Chapter 8
I cannot say that the theory put forward tentatively, and with much reservation by Professor Thomson, that the calcareous matter is dissolved out by the relatively fresh water of the deep currents from the Antarctic regions, appears satisfactory to me. Nor do I see my way to the acceptance of the suggestion of Dr. Carpenter, that the red clay is the result of the decomposition of previously-formed greensand. At present there is no evidence that greensand casts are ever formed at great depths; nor has it been proved that _Glauconite_ is decomposable by the agency of water and carbonic acid.
I think it probable that we shall have to wait some time for a sufficient explanation of the origin of the abyssal red clay, no less than for that of the sublittoral greensand in the intermediate zone. But the importance of the establishment of the fact that these various deposits are being formed in the ocean, at the present day, remains the same; whether its _rationale_ be understood or not.
For, suppose the globe to be evenly covered with sea, to a depth say of a thousand fathoms--then, whatever might be the mineral matter composing the sea-bottom, little or no deposit would be formed upon it, the abrading and denuding action of water, at such a depth, being exceedingly slight.
Next, imagine sponges, _Radiolaria, Foraminifera_, and diatomaceous plants, such as those which now exist in the deep-sea, to be introduced: they would be distributed according to the same laws as at present, the sponges (and possibly some of the _Foraminifera_), covering the bottom, while other _Foraminifera_, with the _Radiolaria_ and _Diatomacea_, would increase and multiply in the surface waters. In accordance with the existing state of things, the _Radiolaria_ and Diatoms would have a universal distribution, the latter gathering most thickly in the polar regions, while the _Foraminifera_ would be largely, if not exclusively, confined to the intermediate zone; and, as a consequence of this distribution, a bed of "chalk" would begin to form in the intermediate zone, while caps of silicious rock would accumulate on the circumpolar regions.
Suppose, further, that a part of the intermediate area were raised to within two or three hundred fathoms of the surface--for anything that we know to the contrary, the change of level might determine the substitution of greensand for the "chalk"; while, on the other hand, if part of the same area were depressed to three thousand fathoms, that change might determine the substitution of a different silicate of alumina and iron--namely, clay--for the "chalk" that would otherwise be formed.
If the _Challenger_ hypothesis, that the red clay is the residue left by dissolved _Foraminiferous_ skeletons, is correct, then all these deposits alike would be directly, or indirectly, the product of living organisms. But just as a silicious deposit may be metamorphosed into opal or quartzite, and chalk into marble, so known metamorphic agencies may metamorphose clay into schist, clay-slate, slate, gneiss, or even granite. And thus, by the agency of the lowest and simplest of organisms, our imaginary globe might be covered with strata, of all the chief kinds of rock of which the known crust of the earth is composed, of indefinite thickness and extent.
The bearing of the conclusions which are now either established, or highly probable, respecting the origin of silicious, calcareous, and clayey rocks, and their metamorphic derivatives, upon the archaeology of the earth, the elucidation of which is the ultimate object of the geologist, is of no small importance.
A hundred years ago the singular insight of Linnaeus enabled him to say that "fossils are not the children but the parents of rocks,"[9] and the whole effect of the discoveries made since his time has been to compile a larger and larger commentary upon this text. It is, at present, a perfectly tenable hypothesis that all siliceous and calcareous rocks are either directly, or indirectly, derived from material which has, at one time or other, formed part of the organized framework of living organisms. Whether the same generalization may be extended to aluminous rocks, depends upon the conclusion to be drawn from the facts respecting the red clay areas brought to light by the _Challenger_. If we accept the view taken by Wyville Thomson and his colleagues--that the red clay is the residuum left after the calcareous matter of the _Globigerinoe_ ooze has been dissolved away--then clay is as much a product of life as limestone, and all known derivatives of clay may have formed part of animal bodies.
[Footnote 9: "Petrificata montium calcariorum non filii sed parentes sunt, cum omnis calx oriatur ab animalibus."--_Systema Naturae_, Ed. xii., t. iii., p. 154. It must be recollected that Linnaeus included silex, as well as limestone, under the name of "calx," and that he would probably have arranged Diatoms among animals, as part of "chaos." Ehrenberg quotes another even more pithy passage, which I have not been able to find in any edition of the _Systema_ accessible to me: "Sic lapides ab animalibus, nec vice versa. Sic runes saxei non primaevi, sed temporis filiae."]
So long as the _Globigerinoe_;, actually collected at the surface, have not been demonstrated to contain the elements of clay, the _Challenger_ hypothesis, as I may term it, must be accepted with reserve and provisionally, but, at present, I cannot but think that it is more probable than any other suggestion which has been made.
Accepting it provisionally, we arrive at the remarkable result that all the chief known constituents of the crust of the earth may have formed part of living bodies; that they may be the "ash" of protoplasm; that the "_rupes saxei_" are not only _"temporis,"_ but "_vitae filiae_"; and, consequently, that the time during which life has been active on the globe may be indefinitely greater than the period, the commencement of which is marked by the oldest known rocks, whether fossiliferous or unfossiliferous.
And thus we are led to see where the solution of a great problem and apparent paradox of geology may lie. Satisfactory evidence now exists that some animals in the existing world have been derived by a process of gradual modification from pre-existing forms. It is undeniable, for example, that the evidence in favour of the derivation of the horse from the later tertiary _Hipparion_, and that of the _Hipparion_ from _Anchitherium_, is as complete and cogent as such evidence can reasonably be expected to be; and the further investigations into the history of the tertiary mammalia are pushed, the greater is the accumulation of evidence having the same tendency. So far from palaeontology lending no support to the doctrine of evolution--as one sees constantly asserted--that doctrine, if it had no other support, would have been irresistibly forced upon us by the palaeontological discoveries of the last twenty years.
If, however, the diverse forms of life which now exist have been produced by the modification of previously-existing less divergent forms, the recent and extinct species, taken as a whole, must fall into series which must converge as we go back in time. Hence, if the period represented by the rocks is greater than, or co-extensive with, that during which life has existed, we ought, somewhere among the ancient formations, to arrive at the point to which all these series converge, or from which, in other words, they have diverged--the primitive undifferentiated protoplasmic living things, whence the two great series of plants and animals have taken their departure.
But, as a matter of fact, the amount of convergence of series, in relation to the time occupied by the deposition of geological formations, is extraordinarily small. Of all animals the higher _Vertebrata_ are the most complex; and among these the carnivores and hoofed animals (_Ungulata_) are highly differentiated. Nevertheless, although the different lines of modification of the _Carnivora_ and those of the _Ungulata_, respectively, approach one another, and, although each group is represented by less differentiated forms in the older tertiary rocks than at the present day, the oldest tertiary rocks do not bring us near the primitive form of either. If, in the same way, the convergence of the varied forms of reptiles is measured against the time during which their remains are preserved--which is represented by the whole of the tertiary and mesozoic formations--the amount of that convergence is far smaller than that of the lines of mammals between the present time and the beginning of the tertiary epoch. And it is a broad fact that, the lower we go in the scale of organization, the fewer signs are there of convergence towards the primitive form from whence all must have diverged, if evolution be a fact. Nevertheless, that it is a fact in some cases, is proved, and I, for one, have not the courage to suppose that the mode in which some species have taken their origin is different from that in which the rest have originated.
What, then, has become of all the marine animals which, on the hypothesis of evolution, must have existed in myriads in those seas, wherein the many thousand feet of Cambrian and Laurentian rocks now devoid, or almost devoid, of any trace of life were deposited?
Sir Charles Lyell long ago suggested that the azoic character of these ancient formations might be due to the fact that they had undergone extensive metamorphosis; and readers of the "Principles of Geology" will be familiar with the ingenious manner in which he contrasts the theory of the Gnome, who is acquainted only with the interior of the earth, with those of ordinary philosophers, who know only its exterior.
The metamorphism contemplated by the great modern champion of rational geology is, mainly, that brought about by the exposure of rocks to subterranean heat; and where no such heat could be shown to have operated, his opponents assumed that no metamorphosis could have taken place. But the formation of greensand, and still more that of the "red clay" (if the _Challenger_ hypothesis be correct) affords an insight into a new kind of metamorphosis--not igneous, but aqueous--by which the primitive nature of a deposit may be masked as completely as it can be by the agency of heat. And, as Wyville Thomson suggests, in the passage I have quoted above (p. 17), it further enables us to assign a new cause for the occurrence, so puzzling hitherto, of thousands of feet of unfossiliferous fine-grained schists and slates, in the midst of formations deposited in seas which certainly abounded in life. If the great deposit of "red clay" now forming in the eastern valley of the Atlantic were metamorphosed into slate and then upheaved, it would constitute an "azoic" rock of enormous extent. And yet that rock is now forming in the midst of a sea which swarms with living beings, the great majority of which are provided with calcareous or silicious shells and skeletons; and, therefore, are such as, up to this time, we should have termed eminently preservable.
Thus the discoveries made by the _Challenger_ expedition, like all recent advances in our knowledge of the phenomena of biology, or of the changes now being effected in the structure of the surface of the earth, are in accordance with and lend strong support to, that doctrine of Uniformitarianism, which, fifty years ago, was held only by a small minority of English geologists--Lyell, Scrope, and De la Beche--but now, thanks to the long-continued labours of the first two, and mainly to those of Sir Charles Lyell, has gradually passed from the position of a heresy to that of catholic doctrine.
Applied within the limits of the time registered by the known fraction of the crust of the earth, I believe that uniformitarianism is unassailable. The evidence that, in the enormous lapse of time between the deposition of the lowest Laurentian strata and the present day, the forces which have modified the surface of the crust of the earth were different in kind, or greater in the intensity of their action, than those which are now occupied in the same work, has yet to be produced. Such evidence as we possess all tends in the contrary direction, and is in favour of the same slow and gradual changes occurring then as now.
But this conclusion in nowise conflicts with the deductions of the physicist from his no less clear and certain data. It may be certain that this globe has cooled down from a condition in which life could not have existed; it may be certain that, in so cooling, its contracting crust must have undergone sudden convulsions, which were to our earthquakes as an earthquake is to the vibration caused by the periodical eruption of a Geyser; but in that case, the earth must, like other respectable parents, have sowed her wild oats, and got through her turbulent youth, before we, her children, have any knowledge of her.
So far as the evidence afforded by the superficial crust of the earth goes, the modern geologist can, _ex animo_, repeat the saying of Hutton, "We find no vestige of a beginning--no prospect of an end." However, he will add, with Hutton, "But in thus tracing back the natural operations which have succeeded each other, and mark to us the course of time past, we come to a period in which we cannot see any further." And if he seek to peer into the darkness of this period, he will welcome the light proffered by physics and mathematics.
IV
YEAST
[1871]
It has been known, from time immemorial, that the sweet liquids which may be obtained by expressing the juices of the fruits and stems of various plants, or by steeping malted barley in hot water, or by mixing honey with water--are liable to undergo a series of very singular changes, if freely exposed to the air and left to themselves, in warm weather. However clear and pellucid the liquid may have been when first prepared, however carefully it may have been freed, by straining and filtration, from even the finest visible impurities, it will not remain clear. After a time it will become cloudy and turbid; little bubbles will be seen rising to the surface, and their abundance will increase until the liquid hisses as if it were simmering on the fire. By degrees, some of the solid particles which produce the turbidity of the liquid collect at its surface into a scum, which is blown up by the emerging air-bubbles into a thick, foamy froth. Another moiety sinks to the bottom, and accumulates as a muddy sediment, or "lees."
When this action has continued, with more or less violence, for a certain time, it gradually moderates. The evolution of bubbles slackens, and finally comes to an end; scum and lees alike settle at the bottom, and the fluid is once more clear and transparent. But it has acquired properties of which no trace existed in the original liquid. Instead of being a mere sweet fluid, mainly composed of sugar and water, the sugar has more or less completely disappeared; and it has acquired that peculiar smell and taste which we call "spirituous." Instead of being devoid of any obvious effect upon the animal economy, it has become possessed of a very wonderful influence on the nervous system; so that in small doses it exhilarates, while in larger it stupefies, and may even destroy life.
Moreover, if the original fluid is put into a still, and heated moderately, the first and last product of its distillation is simple water; while, when the altered fluid is subjected to the same process, the matter which is first condensed in the receiver is found to be a clear, volatile substance, which is lighter than water, has a pungent taste and smell, possesses the intoxicating powers of the fluid in an eminent degree, and takes fire the moment it is brought in contact with a flame. The Alchemists called this volatile liquid, which they obtained from wine, "spirits of wine," just as they called hydrochloric acid "spirits of salt," and as we, to this day, call refined turpentine "spirits of turpentine." As the "spiritus," or breath, of a man was thought to be the most refined and subtle part of him, the intelligent essence of man was also conceived as a sort of breath, or spirit; and, by analogy, the most refined essence of anything was called its "spirit." And thus it has come about that we use the same word for the soul of man and for a glass of gin.
At the present day, however, we even more commonly use another name for this peculiar liquid--namely, "alcohol," and its origin is not less singular. The Dutch physician, Van Helmont, lived in the latter part of the sixteenth and the beginning of the seventeenth century--in the transition period between alchemy and chemistry--and was rather more alchemist than chemist. Appended to his "Opera Omnia," published in 1707, there is a very needful "Clavis ad obscuriorum sensum referendum," in which the following passage occurs.--
"ALCOHOL.--Chymicis est liquor aut pulvis summé subtilisatus, vocabulo Orientalibus quoque, cum primis Habessinis, familiari, quibus _cohol_ speciatim pulverem impalpabilem ex antimonio pro oculis tingendis denotat ... Hodie autem, ob analogiam, quivis pulvis tenerior ut pulvis oculorum cancri summé subtilisatus _alcohol_ audit, haud aliter ac spiritus rectificatissimi _alcolisati_ dicuntur."
Similarly, Robert Boyle speaks of a fine powder as "alcohol"; and, so late as the middle of the last century, the English lexicographer, Nathan Bailey, defines "alcohol" as "the pure substance of anything separated from the more gross, a very fine and impalpable powder, or a very pure, well-rectified spirit." But, by the time of the publication of Lavoisier's "Traité Elémentaire de Chimie," in 1789, the term "alcohol," "alkohol," or "alkool" (for it is spelt in all three ways), which Van Helmont had applied primarily to a fine powder, and only secondarily to spirits of wine, had lost its primary meaning altogether; and, from the end of the last century until now, it has, I believe, been used exclusively as the denotation of spirits of wine, and bodies chemically allied to that substance.
The process which gives rise to alcohol in a saccharine fluid is known tones as "fermentation"; a term based upon the apparent boiling up or "effervescence" of the fermenting liquid, and of Latin origin.
Our Teutonic cousins call the same process "gähren," "gäsen," "göschen," and "gischen"; but, oddly enough, we do not seem to have retained their verb or their substantive denoting the action itself, though we do use names identical with, or plainly derived from, theirs for the scum and lees. These are called, in Low German, "gäscht" and "gischt"; in Anglo- Saxon, "gest," "gist," and "yst," whence our "yeast." Again, in Low German and in Anglo-Saxon there is another name for yeast, having the form "barm," or "beorm"; and, in the Midland Counties, "barm" is the name by which yeast is still best known. In High German, there is a third name for yeast, "hefe," which is not represented in English, so far as I know.
All these words are said by philologers to be derived from roots expressive of the intestine motion of a fermenting substance. Thus "hefe" is derived from "heben," to raise; "barm" from "beren" or "bären," to bear up; "yeast," "yst," and "gist," have all to do with seething and foam, with "yeasty" waves, and "gusty" breezes.
The same reference to the swelling up of the fermenting substance is seen in the Gallo-Latin terms "levure" and "leaven."
It is highly creditable to the ingenuity of our ancestors that the peculiar property of fermented liquids, in virtue of which they "make glad the heart of man," seems to have been known in the remotest periods of which we have any record. All savages take to alcoholic fluids as if they were to the manner born. Our Vedic forefathers intoxicated themselves with the juice of the "soma"; Noah, by a not unnatural reaction against a superfluity of water, appears to have taken the earliest practicable opportunity of qualifying that which he was obliged to drink; and the ghosts of the ancient Egyptians were solaced by pictures of banquets in which the wine-cup passes round, graven on the walls of their tombs. A knowledge of the process of fermentation, therefore, was in all probability possessed by the prehistoric populations of the globe; and it must have become a matter of great interest even to primaeval wine-bibbers to study the methods by which fermented liquids could be surely manufactured. No doubt it was soon discovered that the most certain, as well as the most expeditious, way of making a sweet juice ferment was to add to it a little of the scum, or lees, of another fermenting juice. And it can hardly be questioned that this singular excitation of fermentation in one fluid, by a sort of infection, or inoculation, of a little ferment taken from some other fluid, together with the strange swelling, foaming, and hissing of the fermented substance, must have always attracted attention from the more thoughtful. Nevertheless, the commencement of the scientific analysis of the phenomena dates from a period not earlier than the first half of the seventeenth century.
At this time, Van Helmont made a first step, by pointing out that the peculiar hissing and bubbling of a fermented liquid is due, not to the evolution of common air (which he, as the inventor of the term "gas," calls "gas ventosum"), but to that of a peculiar kind of air such as is occasionally met with in caves, mines, and wells, and which he calls "gas sylvestre."
But a century elapsed before the nature of this "gas sylvestre," or, as it was afterwards called, "fixed air," was clearly determined, and it was found to be identical with that deadly "choke-damp" by which the lives of those who descend into old wells, or mines, or brewers' vats, are sometimes suddenly ended; and with the poisonous aëriform fluid which is produced by the combustion of charcoal, and now goes by the name of carbonic acid gas.
During the same time it gradually became evident that the presence of sugar was essential to the production of alcohol and the evolution of carbonic acid gas, which are the two great and conspicuous products of fermentation. And finally, in 1787, the Italian chemist, Fabroni, made the capital discovery that the yeast ferment, the presence of which is necessary to fermentation, is what he termed a "vegeto-animal" substance; that is, a body which gives of ammoniacal salts when it is burned, and is, in other ways, similar to the gluten of plants and the albumen and casein of animals.
These discoveries prepared the way for the illustrious Frenchman, Lavoisier, who first approached the problem of fermentation with a complete conception of the nature of the work to be done. The words in which he expresses this conception, in the treatise on elementary chemistry to which reference has already been made, mark the year 1789 as the commencement of a revolution of not less moment in the world of science than that which simultaneously burst over the political world, and soon engulfed Lavoisier himself in one of its mad eddies.
"We may lay it down as an incontestable axiom that, in all the operations of art and nature, nothing is created; an equal quantity of matter exists both before, and after the experiment: the quality and quantity of the elements remain precisely the same, and nothing takes place beyond changes and modifications in the combinations of these elements. Upon this principle the whole art of performing chemical experiments depends; we must always suppose an exact equality between the elements of the body examined and those of the products of its analysis.