Cavity-Nesting Birds of North American Forests Agriculture Handbook 511
Part 3
Habitat: Saw-whet owls are small, nocturnal hunters of the deep north woods. They nest in the Rocky Mountains up to about 11,000 feet (Bailey and Niedrach 1965). This widely distributed owl nests in most of the forest types throughout the northern half of the United States, but only rarely do they nest as far south as central Missouri.
Nest: These small owls prefer to nest in old flicker or other woodpecker holes (Bent 1938). Nesting habitat may be improving in areas where Dutch Elm disease has infested many elms, and woodpeckers have drilled nest holes (Hamerstrom 1972). Saw-whets will use nesting boxes if sawdust or straw is provided. Nest boxes should be 6 × 6 × 9 inches with a 2.5-inch entrance hole (Hamerstrom 1972).
Food: Saw-whet owls consume mostly small mammals and insects. Specific food items include mice, shrews, young squirrels, chipmunks, bats, beetles, grasshoppers, and occasionally small birds (Scott and Patton 1975, Burton 1973, Hamerstrom 1972, Bent 1938).
Chimney swift _Chaetura pelagica_ L 5″ W 12½″
Habitat: Chimney swifts are found throughout the eastern half of the United States in wooded and open areas. They have adopted to man-made structures and are no longer dependent upon hollow trees for nesting and roosting.
Nest: Originally chimney swifts nested in hollow trees, especially sycamores. They now use chimneys, barn silos, cisterns, and wells (Pearson 1936). Their nests are made of twigs, which are glued to a vertical surface with saliva to form a “half-saucer” (Forbush and May 1939).
Food: Chimney swifts feed almost entirely on flying insects but will sometimes take small caterpillars hanging from tree branches or leaves (Forbush and May 1939).
Vaux’s swift _Chaetura vauxi_ L 4½″
Habitat: This small swift is most likely to be found in river valleys among dense Douglas-fir and redwood forests in the western United States.
Nest: Nests are usually located in tall hollow snags in burned or logged areas and are made from twigs (Peterson 1961, Robbins et al. 1966). Nests have been reported in unused chimneys and under building eaves (Bent 1940).
Food: Flying insects such as mosquitoes, gnats, flies, and small beetles captured in flight probably make up the entire diet (Bent 1940).
Coppery-tailed trogon _Trogon elegans_ L 10″
Habitat: Coppery-tailed trogons can be found along riparian streams and in pine-oak forests in Arizona, southwestern New Mexico, and southern Texas.
Nest: Nests are found 12 to 40 feet above the ground in deserted large woodpecker holes (Bent 1940). Cottonwood and sycamore snags are usually selected for nests. Of the 34 species in the family Trogonidae, this is the only one that breeds in the United States (Wetmore 1964).
Food: There is little information on the food of these birds, but apparently both animal and vegetable matter are included in the diet. Bent (1940) reported on stomach contents of two birds. One contained adult and larvae of moths and butterflies; the other contained 68 percent insects and 32 percent fruits. Insect food included grasshoppers, praying mantids, stink bugs, leaf beetles, and larvae of hawk moth, sawfly, and miscellaneous other insects. Vegetable food consisted of fruits of cissus and red pepper and undetermined plant fiber.
Common flicker _Colaptes auratus_ L 10½″
Habitat: Flickers are commonly found near large trees in open woodlands, fields, and meadows throughout North America. In winter, they occasionally seek shelter in coniferous woods or swamps. Previously three species were recognized: the yellow-shafted of the East, the red-shafted of the West, and the gilded of the southwestern desert. These are now considered a single species.
Nest: Flickers prefer to nest in open country or in lightly wooded suburban areas where park-like situations are plentiful (Bent 1939). Conner et al. (1975) reported that flickers usually nest in edge habitats and, in extensive forested areas, nest only in or around openings. Flickers excavate nest holes with a 2.75-inch entrance hole diameter in dead trees or dead limbs of many species of trees including aspen, cottonwood, oak, willow, sycamore, pine, and juniper. Nests are sometimes as high as 100 feet but usually between 10 and 30 feet (Scott and Patton 1975, Lawrence 1967).
Food: Sixty percent of common flicker food is animal matter. Of this, 75 percent is ants, more than taken by any other North American bird. Some flicker stomachs have contained over 2,000 ants. The rest of the insect material includes beetles, wasps, caterpillars, grubs, and crickets. The vegetable portion of the diet includes weed seeds, cultivated grain, and the fruit of wild shrubs and trees (Bent 1939, Forbush and May 1939).
Pileated Woodpecker _Dryocopus pileatus_ L 15″
Habitat: Forests of heavy timber and secondary growth consisting of mixed deciduous and coniferous trees are the preferred year-round habitat for pileated woodpeckers. These large woodpeckers have become less abundant over much of their former range where extensive agriculture or logging practices have eliminated large tracts of old growth forests. In the Ozarks, they are plentiful wherever extensive forests remain, preferring areas where past cutting practices (early 1900’s) have left scattered large cull trees throughout.
Nest: Pileated woodpecker nests have been found in beech, poplar, tulip-popular, birch, oak, hickory, maple, hemlock, pine, ash, elm, basswood, and aspen trees. Cavity heights range from 15 to 70 feet, with an entrance hole up to 4 inches in diameter (Hoyt 1941, 1957). Tall dead trees with smooth surfaces and few limbs are preferred. One tree may be used for several years, but rarely is a nest hole reused. This behavior provides cavities for other wildlife, including wood ducks, owls, and squirrels (Hoyt 1957). Timber stands with sawtimber of 15 to 18 inches dbh provide adequate habitat if there is a supply of dead and decaying trees (Conner et al. 1975).
Food: Insects make up more than 70 percent of the food of pileated woodpeckers. Ants (especially carpenter ants) and beetles are the major food items. In the fall, dogwood berries, wild cherries, acorns, and other wild fruit are included in the diet (Bent 1939).
Red-bellied woodpecker _Melanerpes carolinus_ L 8½″
Habitat: Red-bellied woodpeckers are common throughout southeastern forest types. This bird has habits similar to those of the red-headed woodpecker, except that the red-headed prefers open woodlands, farm yards, and field edges whereas the red-bellied prefers larger expanses of forest. Bailey and Niedrach (1965) reported that the red-bellied woodpecker is extending its range westward up the river valleys of the Great Plains.
Nest: These woodpeckers most commonly excavate nest holes in dead limbs of living trees. Excavations were found in a wide variety of tree species, and ranged from 33 to 72 feet above ground (Reller 1972). Cavities are usually located in mature timber stands. Between September and January, males and females roost in separate holes. Often one of the roost holes (usually that of the female) becomes the nest site (Kilham 1958).
Food: Although primarily insectivorous, red-bellied woodpeckers consume more vegetable matter than most woodpeckers. Insects that are eaten include ants, adult and larval beetles, and caterpillars. Vegetation eaten includes grain, berries, and fruits of holly, dogwood, and poison ivy. Acorns and berries are stored in crevices in the fall (Kilham 1963, Bent 1939).
Golden-fronted woodpecker _Melanerpes aurifrons_ L 8½″
Habitat: The golden-fronted woodpecker’s preferred habitat is mesquite and riparian woodlands in Texas and Oklahoma. Cooke (1888) listed this species as an abundant resident of the lower Rio Grande Valley, Texas, in 1884.
Nest: Nesting behavior of the golden-fronted is similar to that of the red-bellied woodpecker (Pearson 1936). Tall trees of pecan, oak, and mesquite are the major species used for nesting (Bent 1939). Occasionally fence posts, telephone poles, and bird boxes are used (Reed 1965).
Food: The diet of the golden-fronted woodpecker consists of both insects and vegetable matter. Grasshoppers make up more than half of the animal matter and other insects include beetles and ants (Pearson 1936, Bent 1939). Vegetable matter consumed consists of corn, acorns, wild fruits, and berries (Bent 1939).
Gila woodpecker _Melanerpes uropygialis_ L 8¼″
Habitat: This woodpecker is found on desert mesas in association with creosote bush, mesquite, and saguaro cactus from central Arizona to edges of adjacent states. It is also common in river bottoms and in foothill canyons among cottonwoods, willows, and sycamores.
Nest: The Gila woodpecker excavates holes in saguaro cacti for nests. Cottonwoods, willows, and mesquites are also used at higher elevations (Bent 1939, Ligon 1961).
Food: The diet of the Gila woodpecker consists of ants, beetles, grasshoppers, fruits from saguaro cactus, and mistletoe berries (Bent 1939). This woodpecker has been reported to remove eggs from the nests of various songbirds.
Red-headed woodpecker _Melanerpes erythrocephalus_ L 7½″
Habitat: Red-headed woodpeckers prefer to nest and roost in open areas. Farmyards, field edges, and timber stands that have been treated with herbicides or burned are preferred habitats. Redheads are attracted to areas with many dead snags and lush herbaceous ground cover, but not to woods with closed canopies. They are found throughout the East and along wooded streams of the prairie to eastern Colorado and Wyoming. Competition for nesting space is often intensive where starlings are abundant (Bailey and Niedrach 1965).
Nest: Red-headed woodpeckers most commonly excavate holes in the trunks of dead trees. Holes are excavated from 24 to 65 feet above the ground and the 1.8-inch diameter entrance hole often faces south or west (Reller 1972). These woodpeckers may excavate new holes each year, or use old nest sites.
Food: Red-headed woodpeckers consume about half animal matter (mostly insects) and half vegetable matter. Occasionally the eggs or the young of other birds are destroyed. Although a wide variety of vegetable matter is consumed, acorns from pin oak comprise a large portion of the winter diet. Nuts are stored whole or in pieces in cracks and crevices in bark, and in cavities which are sealed with bits of bark when full. These birds also store insects (especially grasshoppers) along with acorns in cavities and crevices (Kilham 1963, Bent 1939).
Acorn woodpecker _Melanerpes formicivorus_ L 8″
Habitat: The acorn woodpecker is a common resident of mixed oak-pine woodland and adjacent open grassland from Oregon along the Pacific Coast to the southwestern United States.
Nest: Acorn woodpeckers are communal nesters, and the young are fed by the entire group (Wetmore 1964). They usually excavate holes in ponderosa pine, but live and dead oaks of various species, sycamore, cottonwood, and willow are also used for nests. Their old holes are important for secondary cavity nesters such as small owls, purple martins, violet-green swallows, nuthatches, house wrens, and kestrels (Bent 1939).
Food: As the name implies, acorn woodpeckers feed mostly on acorns which are stored in holes drilled in communal trees. Sap from several species of oaks also is consumed from midwinter to summer (MacRoberts and MacRoberts 1972). About 25 percent of the diet is insects, including grasshoppers, ants, beetles, and flies (Bent 1939). Almonds, walnuts, and pecans are eaten when they are available.
Lewis’ woodpecker _Melanerpes lewis_ L 9″
Habitat: Open or parklike ponderosa pine forest is probably the major breeding habitat of the Lewis’ woodpecker. These woodpeckers also nest in burned over stands of Douglas-fir, mixed conifer, pinyon-juniper, riparian, and oak woodlands (Bock 1970).
Nest: The Lewis’ woodpecker generally excavates its own nest cavity, but will use natural cavities or holes excavated in previous years. Bock (1970) summarized the following nest data: height range 5 to 170 feet; 47 nests in dead stubs and 17 in live trees; 29 nests in conifers, 31 in cottonwood and sycamore, 6 in oaks, 2 in power poles, 1 in juniper, and 1 in catalpa. At Boca Reservoir, California, 10 of 11 nests were in dead ponderosa pines, and the other was in a hollow section of a living pine.
Food: Insects, including flies, ladybird beetle larvae, tent caterpillars, ants, and mayflies, were the primary food of Lewis’ woodpeckers during spring and summer (Bock 1970). Fruits and berries were the most frequently used food in late summer and fall, while winter food consisted mostly of acorns and almonds gathered and stored in crevices of dead trees, power poles, and oak bark. Hadow (1973) reported that, on snowy days when insects were inactive, Lewis’ woodpeckers in southeastern Colorado spent 99 percent of their feeding time feeding from caches of acorns and corn kernels.
Yellow-bellied sapsucker _Sphyrapicus varius_ L 7¾″
Habitat: The yellow-bellied sapsucker (sometimes called red-naped) is most abundant along streams in mixed hardwood-conifer forests. It is also found in ponderosa pine, aspen, mixed conifer, lodgepole pine, and in mixed stands of fir-larch-pine.
Nest: Yellow-bellied sapsuckers usually nest in cavities in snags or live trees with rotten heartwood. Aspen seems to be the preferred species (Howell 1952, Lawrence 1967, Kilham 1971), but nests have also been found in ponderosa pine, birch, elm, butternut, cottonwood, alder, willow, beech, maple, and fir (Bent 1939). Kilham (1971) noted that nest trees were often infected by the _Fomes_ fungus. Nest height varies from 5 to 70 feet above ground. The same nest tree is often used repeatedly, but a new cavity is excavated each year.
Food: Sap is eaten throughout the year by the yellow-bellied sapsucker, but the amount taken and tree species used vary seasonally (Tate 1973, Lawrence 1967). The birds regularly tap one or two “favorite trees” in their area; Oliver (1970) found that these tend to be trees which have been wounded (by logging, porcupines, etc.). About 80 percent of the insect food taken consists of ants (McAtee 1911). Other insects in their diet include beetles and wasps, but none of the woodboring larvae. The fruits of dogwood, black alder, Virginia creeper, and blackberries are included in the small portion of vegetable matter eaten (Bent 1939).
Williamson’s sapsucker _Sphyrapicus thyroideus_ L 8¼″
Habitat: This sapsucker prefers mixed conifer-hardwood forests of the Rocky Mountain region but also inhabits the subalpine spruce-fir-lodgepole zone, and ponderosa pine, Douglas-fir, and aspen forests.
Nest: The choice of tree species for nesting seems to differ between regions. Bent (1939), Packard (1945), Bailey and Niedrach (1965), Burleigh (1972), and Jackman (1975) reported Williamson’s sapsuckers nesting primarily in conifers. Other authors (Rasmussen 1941, Hubbard 1965, Tatschl 1967, Ligon 1961, Crockett and Hadow 1975) found a preference for aspens. Of 57 nests in Colorado examined by Crockett and Hadow (1975), 49 were in aspens, especially aspens infected by the _Fomes_ fungus; where pines were used, there were no suitable aspen sites nearby. In Arizona, we found 17 nests in aspen snags, 3 in aspens with dead tops, and 1 nest in a live aspen.
Food: The diet of Williamson’s sapsuckers is made up of 87 percent animal and 13 percent vegetable material (Bent 1939). Most of the animal food taken is ants, and most of the vegetable material is cambium. Like the yellow-bellied sapsucker, the Williamson’s sapsucker feeds on sap, especially in spring, and picks out “favorite trees” which it taps regularly (Oliver 1970).
Hairy woodpecker _Picoides villosus_ L 7½″
Habitat: Hairy woodpeckers are residents of nearly all types of forest from central Canada south.
Nest: Live trees in open woodlands are preferred nesting sites of hairy woodpeckers. This species makes a nest entrance that exactly fits its head and body size (1.6 to 1.8 inches). Because this size also seems very convenient for starlings and flying squirrels, hairy woodpeckers are often troubled with invasions (Kilham 1968a, Lawrence 1967). Hairy woodpeckers will often excavate the entrance so it is camouflaged or hidden, such as on the underside of a limb. Nest heights vary from 15 to 45 feet but are commonly approximately 35 feet high. Hairies will often use the same hole year after year.
Food: Hairy woodpeckers prefer to feed on insects on dead and diseased trees (Bent 1939). Approximately 80 percent of the diet is animal matter; adult and larval beetles, ants, and caterpillars are the most frequently eaten items. The primarily insect diet is supplemented with fruit, corn, acorns, hazelnuts, and many other species (Beal 1911, Bent 1939). The males forage in trees away from the nest for large insects (usually borers) located deep in the wood. Females forage close to the nest on the surface of trees, shrubs, or on the ground for small prey (Kilham 1968a).
Downy woodpecker _Picoides pubescens_ L 5¾″
Habitat: Downy woodpeckers inhabit most of the wooded parts of North America. They are absent or rare in the arid deserts, and not common in the densely forested regions. Favorite habitat includes open woodland, hammocks, orchards, roadside hedges, farmyards, and urban areas (Bent 1939). Occasionally, these birds nest at elevations above 9,000 feet in the central Rockies (Bailey and Niedrach 1965). Most populations are considered nonmigratory; however, there is some movement from north to south and from high elevations to the plains during winter.
Nest: Downy woodpeckers resemble common flickers in many of their nesting habits. Both prefer to excavate near the tops of dead trees in fairly open timber stands. They generally excavate new cavities each year in the same tree, but do not usually use cavities of other birds or reuse old cavities (Lawrence 1967). In the fall, these birds excavate fresh holes to use as winter roosts (Kilham 1962). Nest holes are normally 8 to 50 feet above the ground with an entrance hole 1.2 to 1.4 inches in diameter (Bent 1939).
Food: The diet is about 75 percent animal and 25 percent vegetable material. Animal material consists mostly of economically harmful insects. Kilham (1970) found that beetles, mostly wood-boring larvae, made up 21.5 percent of the diet. Other materials included ants (21 percent), caterpillars (16.5 percent), weevils (3 percent), and fruit (6 percent). Like hairy woodpeckers, downy woodpeckers have been credited with reducing forest pests (MacLellan 1958, 1959, Olson 1953).
Ladder-backed woodpecker _Picoides scalaris_ L 7″
Habitat: Ladder-backed woodpeckers are commonly found in mesquite and deciduous woodland along streams in desert regions of the Southwest.
Nest: Ladder-backed woodpecker nests are located in a variety of trees such as mesquite, screw bean, palo verde, hackberry, china tree, willow, cottonwood, walnut and oak, usually from 2 to 30 feet above ground. Saguaro cactus, yucca stalks, and branches are sometimes used for nests, as are telephone poles and fence posts (Bent 1939, Phillips et al. 1964).
Food: Insects, especially larvae of wood-boring beetles, caterpillars, and ants, are major food items. The ladder-backed woodpecker also has been reported to eat the ripe fruit of saguaro cactus (Bent 1939).
Nuttall’s woodpecker _Picoides nuttallii_ L 6¾″
Habitat: This western woodpecker is an inhabitant of oak woodlands, riparian woods, and chapparal west of the Sierras in California.
Nest: From a literature survey and personal observations, Miller and Bock (1972) summarized the following nest-tree data for 57 nests: 23 percent in oak, 19 percent in willow, 18 percent in sycamore, 16 percent in cottonwood, and 12 percent in alder. Cavities were excavated in dead limbs and trunks of trees, from 3 to 45 feet above ground.
Food: About 80 percent of the diet of Nuttall’s woodpecker is insects, including 28 percent beetles, 15 percent hemipterans, 14 percent lepidopteran larvae, and 8 percent ants (Beal 1911). Most of the insects are gleaned from trunk and limb surfaces or captured on the wing (Short 1971). Wild fruits, poison oak seeds, and occasional acorns make up the vegetable portion of the diet. Nuttall’s woodpeckers in California have been known to take almonds, occasionally robbing the caches of Lewis’ woodpeckers (Emlen 1937, Bock 1970).
Arizona woodpecker _Picoides arizonae_ L 7¼″
Habitat: Arizona woodpeckers are found in live oak and oak-pine forests and canyons from 4,000 to 7,500 feet in Arizona and New Mexico.
Nest: The Arizona woodpecker excavates holes in dead branches of living trees, primarily walnuts, oaks, maples, and sycamores. One nest was reportedly located in a mescal stalk (Bent 1939).
Food: This woodpecker’s diet probably consists largely of the adult and larval stages of insects, with some fruit and acorns, but few details of food items have been reported (Bent 1939).
Red-cockaded woodpecker _Picoides borealis_ 7¼″
Habitat: Red-cockaded woodpeckers need open, mature (at least 60 year old) pine forest with a high fire occurrence (Bent 1939, Jackson 1971, Hopkins and Lynn 1971). Pine species used during breeding season include: longleaf (Crosby 1971), slash (Lowry 1960), loblolly (Sprunt and Chamberlain 1949), and shortleaf (Sutton 1967). Red-cockaded woodpeckers are on the national “Endangered species” list.
Nest: These woodpeckers prefer living pines infected with red heart rot for nesting. These trees have a soft, easily excavated interior with a living exterior, leaving the tree less susceptible to destruction by fire than a dead tree. Cavities can often be reused for at least 20 years and for several years by the same pair (Ligon 1971). The height of cavity is influenced by the location of red heart infection and the height and density of undergrowth (Crosby 1971). The majority of cavities face west, and, when found in leaning trees, are generally on the low side (Beckett 1971, Baker 1971).
Food: Insects make up the major portion of the diet of red-cockaded woodpeckers. Beal (1911) and Beal et al. (1916) examined 99 stomachs and found 86 percent insects and 14 percent vegetable matter, mostly mast. Beetle larvae (16 percent) and ants made up an important part of the year-round diet. The corn earworm can be a major food source during several weeks where conditions are suitable (Ward 1930). Plant material recorded being eaten includes wax myrtle, magnolia, poison ivy, wild grape, pokeberry, blueberry, wild cherry, black gum, and pecan (Beal 1911, Baker 1971, Ligon 1971).
White-headed woodpecker _Picoides albolarvatus_ L 7¾″
Habitat: Open ponderosa pine forest from Washington to central California is the primary habitat of the white-headed woodpecker, but it also occurs in sugar pine, Jeffrey pine, and red and white fir forests (Grinnell and Miller 1944).
Nest: This woodpecker seems to prefer dead pines, but nests have also been found in live and dead fir, oak, and aspen. White-headed woodpeckers usually excavate a new nest cavity every year and often excavate several holes before selecting one to nest in (Bent 1939). Average nest height is 8 feet above ground.