Part 10

So far back as 1805 the Rev. George Barry, in his "History of the Orkney Islands," mentions a rodent that was known locally as the Vole Mouse, which he believed to be the same as the _agrestis_ of Linnæus. He says it "is very often found in marshy grounds that are covered with moss and short heath, in which it makes roads or tracks of about three inches in breadth, and sometimes miles in length, much worn by continual treading, and warped into a thousand different directions."

Towards the end of last century Mr. J. G. Millais obtained specimens, and on a critical examination found that they differed from the known forms in several details of skull structure and in the folds and angles of the teeth, sufficient in his opinion to constitute a new species, which he called _Microtus orcadensis_. It is larger than the Field Vole, with a longer and slightly broader head.

It was found subsequently that specimens from different islands in the Orkney group showed differences due to their segregation over a long period, and they have consequently been distinguished as five sub-species. These differences are minute, and it would be wearisome and out of place in a popular work such as the present to detail them. Generally speaking, they are much alike, and their habits are practically identical, so far as at present known.

The runs are a conspicuous feature of the islands, among the heather and the rough vegetation of the fields and hillsides, running along the surface and at intervals entering tunnels about two and a quarter inches in diameter--just sufficient to clear the spread of the Vole's whiskers. Their nesting places, like those of the Mole, are under small mounds connected with a network of runs. The nest itself is made of grass and roots in a rounded chamber, where at intervals during the spring and summer several litters, varying from three to six, are produced. Before they are three weeks old they are capable of independent existence, but for a time are still guarded by the mother.

The Orkney Vole appears to be specially fond of the roots of Heath Rush (_Juncus squarrosus_), but also feeds on grass and the crops in cultivated fields to which they can gain access. Mr. Millais found that in cold weather his captive Voles became inactive. It has many enemies to hold its increase in check, for every bird and beast large enough to capture it will eat it readily.

*Bank Vole* (_Evotomys glareolus_, Schreber).

There can be little doubt that in many places the Bank Vole has been mistaken for a bright variation of the Field Vole. Its habits are much the same, except that it haunts the hedgerow and wooded country rather than the open fields. As to the differences between the two species, the Bank Vole's head and body measurement is only three and three-quarter inches against four inches in the Field Vole, but its tail is actually (not merely proportionately) longer, being nearly half the length of head and body, and ends in a pencil of hairs. The ears and feet are proportionately larger, the former also being more oval than round. It further differs from the other Voles in the fact that the molar teeth become rooted in the jaws of the adults. The fur of the upper parts is a bright chestnut-red or Vandyke brown, excepting the hairy tail, which is black above. The under parts, including the lower side of the tail, are whitish varying to yellowish or even buff. The redder tint causes this species frequently to be styled the Red Vole. It has pink lips, and grey feet. Whiskers about an inch long. Black and albino varieties have been recorded.

It was considered formerly to be a rare British species, but more discriminating attention to the smaller mammals in recent years, and the wider adoption of trapping by naturalists, have tended to modify that view. It is probably more local, but it appears to be widely distributed, and to occur as far north certainly as Moray and Elgin; but it is not recorded from Ireland, Man, Hebrides, or Shetland. A local race is found in Skomer Island, and has been named _E. skomerensis_. When Yarrell detected the Bank Vole as a distinct species in 1832, it was considered to be of very restricted range in this country. The discovery was made in Essex, but it was soon reported from Herefordshire, Middlesex, Berks, and Cambridge, and more recently from Oxfordshire, Warwickshire, Shropshire, Worcestershire, the Lake District, Northumberland, Inverness, etc. It is restricted to Europe in its wider range. In this country it does not appear to occur at elevations of more than about 700 feet.

The Bank Vole is much more agile than the Field Vole, and not so much given to burrowing. It may be seen abroad in sunny situations at any time of the day, preferring warm, dry places, yet frequently to be found in wet places. It is a good swimmer and diver. It constructs shallow runs in the earth of a roadside bank or hedgebank. These have many entrances and exits above and below, as shown in our photograph; some of the passages connecting with the top of the bank, others enlarging into blind chambers. Its food includes herbage, roots, bulbs, fruits, and seeds; it appears to be particularly fond of turnips. In spring it has been observed climbing rose and hawthorn bushes in order to nibble the new leaves, and in autumn to obtain the hips and haws. It also seeks nuts, berries, the grain of wheat and barley, and the seeds of smaller grasses. Insects, snails, and even small birds are eaten by it, and the entrance to its burrows frequently gives evidence of the variety of its food. It has been known to eat the unpalatable Shrew that it has killed, and even to given way to cannibalism. In Scotland it is accused of eating the shoot-buds of young conifers, especially of larch, and gnawing the bark from branches.

In this country it is occasionally captured in the act of robbing household stores, but in more northern regions, as in Norway and the Yukon, it is a constant inhabitant of houses. It is not one of the hibernating species, therefore as a rule it does not lay up stores; but Mr. Douglas English records the digging up of five Bank Voles with a store of ninety-three sound cob-nuts.

There are several litters of three to six naked and blind young during the year, produced in nests of grass, moss and wool, or feathers, usually placed above ground, sometimes in a bird's nest at some height above it. The males are very quarrelsome, and when fighting or pairing are very vocal, indulging in grunting squeaks.

* * * * *

Three geographical races or sub-species have been recognised by Barrett-Hamilton as distinct species under distinct names. These are Skomer Bank Vole (_Evotomys skomerensis_) from Skomer Island, off Pembroke; Alston's Bank Vole (_E. alstoni_) from the Isle of Mull; and the Raasay Bank Vole (_E. erica_) from Raasay Island, Skye. Barrett-Hamilton regards these as descendants of a former "Boreal" group of Voles, which have been supplanted on the British mainland by the competition of the Bank Vole.

*Rabbit* (_Oryctolagus cuniculus_, Linn.).

The Rabbits and the Hares being comparatively large and familiar members of our native fauna do not appear to stand in need of much space being devoted to them. Familiar as the two common species may be they require to be distinguished not only one from the other, but also from the two other and less familiar species, and in addition there may be a few facts of organisation and habit that are not well-known to all our readers. All members of the family Leporidæ, there are certain structural features in which they all agree in a general way. They belong to the section of Rodents known as Duplicidentata, because in the upper jaw there are always two pairs of incisors. All the other Rodents have only one pair, and they form the division Simplicidentata. The dentition of the Rabbits and Hares is therefore as follows: _i 2/1, c 0/0, p 3/2, m 3/3 = 28_.

The ears are remarkably long and out of all proportion to the size of the body when compared with other Mammals. If laid forward over the face they reach nearly to the tip of the nose. The eyes are large and prominent and placed well to the sides of the head. The hinder legs are longer than the forelegs, and so greatly developed as to be the main propelling power. Instead of pads on the soles to protect the foot and legs from the jars incidental to hard running, the Leporidæ have all the feet covered beneath with a thick coating of hair which gives a firm grip either on hard rock or slippery snow. The tail is very short and turned up. The fur is of triple formation: there is a dense, soft, woolly under-fur, through which push longer and stronger hairs and give the coat its colour, and a still longer but much less numerous set, scattered among the others. The two longer sorts of hair are more or less ringed. The coat becomes thicker in winter.

They are sexually mature at a very early age, and often begin to breed before they have attained to full size. The females are distinguished by the form of the head, which is longer and more delicately modelled than that of the male. The males (bucks), too, are restless and quarrelsome. They are promiscuous breeders, and the entire care of the family falls upon the mother (doe).

Litters of Rabbits succeed one another rapidly between February and September; less frequently in the autumn and winter months. The litters vary from two or three to eight, the higher numbers being those of the warmer months. Young Rabbits are but sparsely clothed and are blind and deaf, the ears being closed and having no power of movement until about the tenth day. The eyes open a day later. In a few days more they can run, and make short excursions from the underground nest. Before they are a month old they are capable of independent existence. Until then the mother will defend them against all-comers, including the Weasel and Stoat, using her powerful hind feet against her adversary, and to good purpose.

The Rabbit is a much smaller animal than the Hare, greyer in colour, with smaller ears and feet, and the black tips of the ears so noticeable in the Hare, are in the Rabbit much reduced or altogether wanting. Its average weight and measurements are: weight, 2-1/2 to 3 lbs.; length of head and body, 16-1/2 ins., tail, 3-3/4 ins., ear, 3 ins., hind foot with claws, 3-3/4 ins. It also differs from the Hare in the structure of its heavier skull, its smaller eyes, shorter ears, and lesser specialisation of the limbs for speed in running.

It is believed that originally the Rabbit was a native only of the western parts of the Mediterranean region--where it still teems--and to have spread northwards largely by human aid. It is known to have been introduced to Italy from Spain by the Romans, who are usually credited with having brought it to Britain. It is now thought, however, that we are indebted to the Normans for its presence. It was certainly here in the twelfth century. The name Rabbit is from the French, and originally indicated the suckling young; the adults being known as Conies.

Although so famous as a digger of extensive underground dwellings, Nature does not appear to have specially built the Rabbit for this purpose; but where the soil is light the efforts of many generations of associated workers have resulted in a system of burrows both extensive and complicated, with bolt-runs as emergency exits and stop-runs for nursery use. Although it prefers the light sand of the dunes covered with Marram-grass, or a sandy heath overgrown with furze and heather, it will on occasion drive its tunnels into firm loam or dry clay; it has been known even to burrow deeply into a surface seam of coal. The forepaws are the principal burrowing tools, the loosened earth being thrown far back by the kicking of the hinder feet. Where stones come in the way that cannot be loosened by the paws, they have been known to be removed by the teeth. These tunnels are about six inches in diameter, increased locally to a foot to provide passing places. The residental quarters are always blind chambers leading from the main passages. The adult Rabbits do not indulge in bedding materials but rest on the bare soil. The does, however, make beds for their young by denuding their own under parts of fur. These tunnels are frequently made use of by other animals, if necessary, by enlarging the passage to admit their larger bodies. When Rabbit-earths are ferreted they sometimes yield more than Rabbits: a Fox, a Cat, a Stoat, with several Rabbits and Rats, have been driven out of the same earth.

Where the Rabbit finds the ground too hard or too wet, it contrives to do without tunnelling underground, making runs under the heather, furze, or matted herbage. Such exceptions are known to sportsmen as Stub-Rabbits or Bush-Rabbits, in the belief that they are a separate species. Occasionally, too, the doe will follow the example of the Hare, and make a nursery "form" in fallow land or among the growing turnips.

The Rabbit is almost exclusively a vegetarian, its chief food being grass and the tender shoots of furze; but in the vicinity of cultivated land they devastate the crops and inflict serious loss upon the farmer. The exception to a vegetable diet is found in its occasional indulgence in snails. Wherever there is sufficient food and his enemies are not too oppressive the Rabbit has extended his range to the most out-of-the-way corners of these islands. A century ago it was a scarce beast in Scotland, but it is now to be found in abundance up to the extreme north. It is found also all over Ireland. Its chief enemies, in addition to man, are all the members of the Weasel family, the Owls, and the Hawks.

Every one who has come across a party of Rabbits feeding must have noticed how conspicuous the white underside of the upturned tail makes them in flight. Wallace suggested that like the white patch on the hind parts of deer and antelope it served as "a signal flag of danger," a guide to the young and feeble to escape from danger by following the most vigorous seniors. This view has been strongly criticised, even ridiculed; but the critics have not offered a better explanation of the upturned Rabbit's "scut." It must, however, be admitted that any explanation ought to fit the case of the Hare which often carries its tail with the white underside exposed, but is a solitary animal with no companions to follow it. On the sand dunes the Rabbit's coat renders it invisible through harmony with the sand.

In the ordinary way of life the Rabbit is a silent animal, except that he gives vent to low growls and grunts to express anger or pleasure; but when terrorised by the imminence of attack by a Stoat the Rabbit finds its voice and gives utterance to a loud scream of agony. This has been referred to in the account of the Stoat (_ante_, p. 68).

*Brown Hare* (_Lepus europæus_, Pallas).

Although in general form and structure the Hare is similar to the closely related Rabbit, there are differences so great as to have induced recent systematists to put them into different genera; and, even superficially, they are sufficiently unlike to enable country folk to keep them distinct under different names. These differences are evident in the longer body, the great length of the hind limbs, the longer ears with their invariable black tips, and the tawny colour of the fur of the upper parts. To these distinctions they can add the patent facts that whilst the Rabbit is a sociable beast, associating in large communities, the Hare is as solitary and retired as a hermit.

There has never been any suggestion that the Hare's title to rank as a real native of Britain is open to doubt, for its name is Anglo-Saxon, and identical with that in use in Denmark and Sweden. It is widely distributed in England, Wales and Scotland up to about 2000 feet elevation; but in Ireland (which has a separate species of its own) the Brown Hare is not a native. It has been a favourite animal of the chase from the earliest times of which we have records; and our ancient sportsmen had age-names for it as for Deer. Thus, in its first year it was a Leveret, in the second year a Hare, and in the third a Great Hare. The male is distinguished as Jack-Hare, and the female as Doe.

The total length of the Hare is about twenty-four inches, to which the tail contributes three inches and two-thirds, and the head nearly four inches. The ears fall short of five inches. The weight averages about eight pounds. The tawny fur of the upper side is harsher than that of the Rabbit, which is due to a predominance of the strong hairs of medium length described under Rabbit. The shoulders, neck, and flanks are of a ruddier hue than the back, and a ruddy band crosses the loins. The sides of the face, and the outer surfaces of the limbs, incline to a yellow tint. The underside is pure white except at the breast and loins where the ruddy tint is continued from above. There is a profusion of black and white whiskers, of which the white are the longer and as much as three and a half inches in length. The tail, which is carried curved up over the back or straight behind, is black above and white on the sides and below. The large, prominent eyes have a horizontal pupil. As it is almost impossible to come upon a Hare asleep, it was formerly believed that they have no eyelids and are compelled, therefore, to sleep with their eyes open. This, of course, was an "inexactitude" comparable to the belief in the Mole's lack of eyes and ears. The prominence of the dark eyes of the Hare, and their situation well to the sides of the head give him a wide field of vision. As regards sexual distinctions, the Jack-Hare has a smaller body, shorter head and redder shoulders than the Doe.

The Hare is not a burrowing animal, and does not seek refuge underground from his enemies, unless hard pressed, when he may enter a Rabbit-burrow temporarily. He relies upon his russet coat harmonising generally with his surroundings; and content with a slight depression among the grass known as a "form," he sits all day and surveys the landscape, ever ready to use his powerful limbs when his keen senses tell him there is danger near. At dusk he goes abroad to feed, and returns to the form at dawn. To break the continuity of scent, when he is leaving his form, and again when returning to it, he will suddenly turn at right angles to his former course and make a prodigious leap--fifteen feet or more--to the top of a bank, then take another long bound, perhaps into marshy ground where the scent will not lie, and repairing to the feeding-ground feel safe from being tracked by Fox or Polecat. He always adopts this leaping trick, also the plan of doubling on his track, which has been the admiration and vexation of the hunter from old times. Shakespeare has told at some length

"How he outruns the winds, and with what care He cranks and crosses with a thousand doubles: The many musets through the which he goes Are like a labyrinth to amaze his foes."

As he courses across the fields you get the impression that he is longer than the measurement given above; the impression is due to the length of the hind legs extended in running, and from which he especially gets the advantage over pursuers when the course lies uphill. He is a good swimmer, and often crosses rivers in order to reach a better feeding-ground, to avoid pursuit, or to seek a mate. Hares have been known to cross the Trent in numbers, where it was two hundred yards wide, in order to reach a field of carrots on the further side; and Yarrel saw one cross an arm of the sea a mile broad.

The "form" is made in rank grass among thickets of gorse and briar, or in the open field where the ground is dry beneath it. It takes and retains the shape of the animal's body, and may be used for a long period. Here the doe brings forth her litter of two, three, or four young--occasionally more. There is much variation in this respect. These are born with their eyes open, and a short furry coat, which however lacks the ruddiness of the adult. They are capable of using their limbs, and are so well advanced in development before birth, that soon each makes its own little form beside the mother's, and when a month old they are quite independent. When left alone on the form, whilst the mother goes off to feed, and anything alarms them, they cry "leek, leek." The adults pair promiscuously; and there appear to be three or four litters a year.

The Hare appears to moult twice a year--in early autumn and early spring; the former being the principal. Like the Rabbit, it is exclusively vegetarian in its feeding, including bark, grain, and roots as well as herbaceous plants in its bill of fare. It is very destructive to young trees in plantations, and the farmer and market-gardener suffer severely from its depredations among the crops of carrot, lettuce, turnip, etc. In the open country it prefers grasses of the genera _Poa_, _Festuca_, and _Molinia_, clover, sow-thistle, and chicory. When it gets into gardens it shows distinct preference for dahlias, carnations, pinks, nasturtiums, parsley, and thyme. In shrubberies it is very destructive to bark and boughs, especially of coniferous trees.

The proverbial expression, "Mad as a March Hare," has reference to the insane antics of the Jack-Hare during the rutting season. He grunts and kicks, bucks like a broncho, and has stand-up boxing-matches with his rivals. In bucking he leaps over his opponent and kicks him vigorously with the hind feet. Though usually harmless, these encounters have been known to have fatal terminations. Though regarded generally as a mute animal, this is not the fact. The Hare has a low but clear cry, which has been described as "don't," "[=o]nt" or "aunt," with varying inflections denoting different moods. When wounded or badly frightened it utters a scream like that of a child in pain, and sportsmen have declared that the pitifulness of it caused them to give up shooting Hares. They have also a warning sound made by grinding the teeth, and it is passed on from Hare to Hare, having the same result as the stamping of feet by the Rabbit. The amorous notes of buck and doe are different, and their imitation by poachers and gamekeepers is known as Hare-sucking.

The doe is a model mother for a time, and will fight desperately in defence of her young; but as soon as they are capable of looking after themselves she casts them off or deserts them.

*Alpine Hare* (_Lepus timidus_, Linn.).