A Practical Physiology: A Text-Book for Higher Schools

Chapter 4

Chapter 49,484 wordsPublic domain

The Bones.

27. The Skeleton. Most animals have some kind of framework to support and protect the soft and fleshy parts of their bodies. This framework consists chiefly of a large number of bones, and is called the skeleton. It is like the keel and ribs of a vessel or the frame of a house, the foundation upon which the bodies are securely built.

There are in the adult human body 200 distinct bones, of many sizes and shapes. This number does not, however, include several small bones found in the tendons of muscles and in the ear. The teeth are not usually reckoned as separate bones, being a part of the structure of the skin.

The number of distinct bones varies at different periods of life. It is greater in childhood than in adults, for many bones which are then separate, to allow growth, afterwards become gradually united. In early adult life, for instance, the skull contains 22 naturally separate bones, but in infancy the number is much greater, and in old age far less.

The bones of the body thus arranged give firmness, strength, and protection to the soft tissues and vital organs, and also form levers for the muscles to act upon.

28. Chemical Composition of Bone. The bones, thus forming the framework of the body, are hard, tough, and elastic. They are twice as strong as oak; one cubic inch of compact bone will support a weight of 5000 pounds. Bone is composed of earthy or mineral matter (chiefly in the form of lime salts), and of animal matter (principally gelatine), in the proportion of two-thirds of the former to one-third of the latter.

Illustration: Fig. 10.—The Skeleton.

The proportion of earthy to animal matter varies with age. In infancy the bones are composed almost wholly of animal matter. Hence, an infant’s bones are rarely broken, but its legs may soon become misshapen if walking is allowed too early. In childhood, the bones still contain a larger percentage of animal matter than in more advanced life, and are therefore more liable to bend than to break; while in old age, they contain a greater percentage of mineral matter, and are brittle and easily broken.

Experiment 3. _To show the mineral matter in bone_. Weigh a large soup bone; put it on a hot, clear fire until it is at a red heat. At first it becomes black from the carbon of its organic matter, but at last it turns white. Let it cool and weigh again. The animal matter has been burnt out, leaving the mineral or earthy part, a white, brittle substance of exactly the same shape, but weighing only about two-thirds as much as the bone originally weighed.

Experiment 4. _To show the animal matter in bone_. Add a teaspoonful of muriatic acid to a pint of water, and place the mixture in a shallow earthen dish. Scrape and clean a chicken’s leg bone, part of a sheep’s rib, or any other small, thin bone. Soak the bone in the acid mixture for a few days. The earthy or mineral matter is slowly dissolved, and the bone, although retaining its original form, loses its rigidity, and becomes pliable, and so soft as to be readily cut. If the experiment be carefully performed, a long, thin bone may even be tied into a knot.

Illustration: Fig. 11.—The fibula tied into a knot, after the hard mineral matter has been dissolved by acid.

29. Physical Properties of Bone. If we take a leg bone of a sheep, or a large end of beef shin bone, and saw it lengthwise in halves, we see two distinct structures. There is a hard and compact tissue, like ivory, forming the outside shell, and a spongy tissue inside having the appearance of a beautiful lattice work. Hence this is called cancellous tissue, and the gradual transition from one to the other is apparent.

It will also be seen that the shaft is a hollow cylinder, formed of compact tissue, enclosing a cavity called the medullary canal, which is filled with a pulpy, yellow fat called _marrow_. The marrow is richly supplied with blood-vessels, which enter the cavity through small openings in the compact tissue. In fact, all over the surface of bone are minute canals leading into the substance. One of these, especially constant and large in many bones, is called the _nutrient foramen_, and transmits an artery to nourish the bone.

At the ends of a long bone, where it expands, there is no medullary canal, and the bony tissue is spongy, with only a thin layer of dense bone around it. In flat bones we find two layers or plates of compact tissue at the surface, and a spongy tissue between. Short and irregular bones have no medullary canal, only a thin shell of dense bone filled with cancellous tissue.

Illustration: Fig. 12.—The Right femur sawed in two, lengthwise. (Showing arrangement of compact and cancellous tissue.)

Experiment 5. Obtain a part of a beef shin bone, or a portion of a sheep’s or calf’s leg, including if convenient the knee joint. Have the bone sawed in two, lengthwise, keeping the marrow in place. Boil, scrape, and carefully clean one half. Note the compact and spongy parts, shaft, etc.

Experiment 6. Trim off the flesh from the second half. Note the pinkish white appearance of the bone, the marrow, and the tiny specks of blood, etc. Knead a small piece of the marrow in the palm; note the oily appearance. Convert some marrow into a liquid by heating. Contrast this fresh bone with an old dry one, as found in the fields. Fresh bones should be kept in a cool place, carefully wrapped in a damp cloth, while waiting for class use.

A fresh or living bone is covered with a delicate, tough, fibrous membrane, called the periosteum. It adheres very closely to the bone, and covers every part except at the joints and where it is protected with cartilage. The periosteum is richly supplied with blood-vessels, and plays a chief part in the growth, formation, and repair of bone. If a portion of the periosteum be detached by injury or disease, there is risk that a layer of the subjacent bone will lose its vitality and be cast off.[5]

30. Microscopic Structure of Bone. If a very thin slice of bone be cut from the compact tissue and examined under a microscope, numerous minute openings are seen. Around these are arranged rings of bone, with little black bodies in them, from which radiate fine, dark lines. These openings are sections of canals called _Haversian canals_, after Havers, an English physician, who first discovered them. The black bodies are minute cavities called _lacunæ_, while the fine lines are very minute canals, _canaliculi_, which connect the lacunæ and the Haversian canals. These Haversian canals are supplied with tiny blood-vessels, while the lacunæ contain bone cells. Very fine branches from these cells pass into the canaliculi. The Haversian canals run lengthwise of the bone; hence if the bone be divided longitudinally these canals will be opened along their length (Fig. 13).

Thus bones are not dry, lifeless substances, but are the very type of activity and change. In life they are richly supplied with blood from the nutrient artery and from the periosteum, by an endless network of nourishing canals throughout their whole structure. Bone has, therefore, like all other living structures, a _self-formative_ power, and draws from the blood the materials for its own nutrition.

Illustration: Fig. 13.

A, longitudinal section of bone, by which the Haversian canals are seen branching and communicating with one another; B, cross section of a very thin slice of bone, magnified about 300 diameters—little openings (Haversian canals) are seen, and around them are ranged rings of bones with little black bodies (lacunæ), from which branch out fine dark lines (canaliculi); C, a bone cell, highly magnified, lying in lacuna.

The Bones of the Head.

31. The Head, or Skull. The bones of the skeleton, the bony framework of our bodies, may be divided into those of the head, the trunk, and the limbs.

The bones of the head are described in two parts,—those of the cranium, or brain-case, and those of the face. Taken together, they form the skull. The head is usually said to contain 22 bones, of which 8 belong to the cranium and 14 to the face. In early childhood, the bones of the head are separate to allow the brain to expand; but as we grow older they gradually unite, the better to protect the delicate brain tissue.

32. The Cranium. The cranium is a dome-like structure, made up in the adult of 8 distinct bones firmly locked together. These bones are:

One Frontal, Two Parietal, Two Temporal One Occipital, One Sphenoid, One Ethmoid.

The frontal bone forms the forehead and front of the head. It is united with the two parietal bones behind, and extends over the forehead to make the roofs of the sockets of the eyes. It is this bone which, in many races of man, gives a dignity of person and a beauty of form seen in no other animal.

The parietal bones form the sides and roof of the skull. They are bounded anteriorly by the frontal bone, posteriorly by the occipital, and laterally by the temporal and sphenoid bones. The two bones make a beautiful arch to aid in the protection of the brain.

The temporal bones, forming the temples on either side, are attached to the sphenoid bone in front, the parietals above, and the occipital behind. In each temporal bone is the cavity containing the organs of hearing. These bones are so called because the hair usually first turns gray over them.

The occipital bone forms the lower part of the base of the skull, as well as the back of the head. It is a broad, curved bone, and rests on the topmost vertebra (atlas) of the backbone; its lower part is pierced by a large oval opening called the _foramen magnum_, through which the spinal cord passes from the brain (Fig. 15).

The sphenoid bone is in front of the occipital, forming a part of the base of the skull. It is wedged between the bones of the face and those of the cranium, and locks together fourteen different bones. It bears a remarkable resemblance to a bat with extended wings, and forms a series of girders to the arches of the cranium.

The ethmoid bone is situated between the bones of the cranium and those of the face, just at the root of the nose. It forms a part of the floor of the cranium. It is a delicate, spongy bone, and is so called because it is perforated with numerous holes like a sieve, through which the nerves of smell pass from the brain to the nose.

Illustration: Fig. 14.—The Skull

33. The Face. The bones of the face serve, to a marked extent, in giving form and expression to the human countenance. Upon these bones depend, in a measure, the build of the forehead, the shape of the chin, the size of the eyes, the prominence of the cheeks, the contour of the nose, and other marks which are reflected in the beauty or ugliness of the face.

The face is made up of fourteen bones which, with the exception of the lower jaw, are, like those of the cranium, closely interlocked with each other. By this union these bones help form a number of cavities which contain most important and vital organs. The two deep, cup-like sockets, called the orbits, contain the organs of sight. In the cavities of the nose is located the sense of smell, while the buccal cavity, or mouth, is the site of the sense of taste, and plays besides an important part in the first act of digestion and in the function of speech.

The bones of the face are:

Two Superior Maxillary, Two Malar, Two Nasal, Two Lachrymal, Two Palate, Two Turbinated, One Vomer, One Lower Maxillary.

34. Bones of the Face. The superior maxillary or upper jawbones form a part of the roof of the mouth and the entire floor of the orbits. In them is fixed the upper set of teeth.

The malar or cheek bones are joined to the upper jawbones, and help form the sockets of the eyes. They send an arch backwards to join the temporal bones. These bones are remarkably thick and strong, and are specially adapted to resist the injury to which this part of the face is exposed.

The nasal or nose bones are two very small bones between the eye sockets, which form the bridge of the nose. Very near these bones are the two small lachrymal bones. These are placed in the inner angles of the orbit, and in them are grooves in which lie the ducts through which the tears flow from the eyes to the nose.

The palate bones are behind those of the upper jaw and with them form the bony part of the roof of the mouth. The inferior turbinated are spongy, scroll-like bones, which curve about within the nasal cavities so as to increase the surface of the air passages of the nose.

The vomer serves as a thin and delicate partition between the two cavities of the nose. It is so named from its resemblance to a ploughshare.

Illustration: Fig. 15.—The Base of the Skull.

A, palate process of upper jawbone; B, zygoma, forming zygomatic arch; C, condyle for forming articulation with atlas; D, foramen magnum; E, occipital bone.

The longest bone in the face is the inferior maxillary, or lower jaw. It has a horseshoe shape, and supports the lower set of teeth. It is the only movable bone of the head, having a vertical and lateral motion by means of a hinge joint with a part of the temporal bone.

35. Sutures of the Skull. Before leaving the head we must notice the peculiar and admirable manner in which the edges of the bones of the outer shell of the skull are joined together. These edges of the bones resemble the teeth of a saw. In adult life these tooth-like edges fit into each other and grow together, suggesting the dovetailed joints used by the cabinet-maker. When united these serrated edges look almost as if sewed together; hence their name, sutures. This manner of union gives unity and strength to the skull.

In infants, the corners of the parietal bones do not yet meet, and the throbbing of the brain may be seen and felt under these “soft spots,” or _fontanelles_, as they are called. Hence a slight blow to a babe’s head may cause serious injury to the brain (Fig. 14).

The Bones of the Trunk.

36. The Trunk. The trunk is that central part of the body which supports the head and the upper pair of limbs. It divides itself into an upper cavity, the thorax, or chest; and a lower cavity, the abdomen. These two cavities are separated by a movable, muscular partition called the diaphragm, or midriff (Figs. 9 and 49).

The bones of the trunk are variously related to each other, and some of them become united during adult life into bony masses which at earlier periods are quite distinct. For example, the sacrum is in early life made up of five distinct bones which later unite into one.

The upper cavity, or chest, is a bony enclosure formed by the breastbone, the ribs, and the spine. It contains the heart and the lungs (Fig. 86).

The lower cavity, or abdomen, holds the stomach, liver, intestines, spleen, kidneys, and some other organs (Fig. 59).

The bones of the trunk may be subdivided into those of the spine, the ribs, and the hips.

The trunk includes 54 bones usually thus arranged:

Spinal Column, 26 bones:

7 Cervical Vertebræ. 12 Dorsal Vertebræ. 5 Lumbar Vertebræ. 1 Sacrum. 1 Coccyx.

Ribs, 24 bones:

14 True Ribs. 6 False Ribs. 4 Floating Ribs.

Sternum. IV. Two Hip Bones. V. Hyoid Bone.

37. The Spinal Column. The spinal column, or backbone, is a marvelous piece of mechanism, combining offices which nothing short of perfection in adaptation and arrangement could enable it to perform. It is the central structure to which all the other parts of the skeleton are adapted. It consists of numerous separate bones, called vertebræ. The seven upper ones belong to the neck, and are called cervical vertebræ. The next twelve are the dorsal vertebræ; these belong to the back and support the ribs. The remaining five belong to the loins, and are called lumbar vertebræ. On looking at the diagram of the backbone (Fig. 9) it will be seen that the vertebræ increase in size and strength downward, because of the greater burden they have to bear, thus clearly indicating that an erect position is the one natural to man.

Illustration: Fig. 16.—The Spinal Column.

This column supports the head, encloses and protects the spinal cord, and forms the basis for the attachment of many muscles, especially those which maintain the body in an erect position. Each vertebra has an opening through its center, and the separate bones so rest, one upon another, that these openings form a continuous canal from the head to the lower part of the spine. The great nerve, known as the spinal cord, extends from the cranium through the entire length of this canal. All along the spinal column, and between each two adjoining bones, are openings on each side, through which nerves pass out to be distributed to various parts of the body.

Between the vertebræ are pads or cushions of cartilage. These act as “buffers,” and serve to give the spine strength and elasticity and to prevent friction of one bone on another. Each vertebra consists of a body, the solid central portion, and a number of projections called processes. Those which spring from the posterior of each arch are the spinous processes. In the dorsal region they are plainly seen and felt in thin persons.

The bones of the spinal column are arranged in three slight and graceful curves. These curves not only give beauty and strength to the bony framework of the body, but also assist in the formation of cavities for important internal organs. This arrangement of elastic pads between the vertebræ supplies the spine with so many elastic springs, which serve to break the effect of shock to the brain and the spinal cord from any sudden jar or injury.

The spinal column rests on a strong three-sided bone called the sacrum, or sacred-bone, which is wedged in between the hip bones and forms the keystone of the pelvis. Joined to the lower end of the sacrum is the coccyx, or cuckoo-bone, a tapering series of little bones.

Experiment 7. Run the tips of the fingers briskly down the backbone, and the spines of the vertebræ will be tipped with red so that they can be readily counted. Have the model lean forward with the arms folded across the chest; this will make the spines of the vertebræ more prominent.

Experiment 8. _To illustrate the movement of torsion in the spine, or its rotation round its own axis_. Sit upright, with the back and shoulders well applied against the back of a chair. Note that the head and neck can be turned as far as 60° or 70°. Now bend forwards, so as to let the dorsal and lumbar vertebræ come into play, and the head can be turned 30° more.

Experiment 9. _To show how the spinal vertebræ make a firm but flexible column._ Take 24 hard rubber overcoat buttons, or the same number of two-cent pieces, and pile them on top of each other. A thin layer of soft putty may be put between the coins to represent the pads of cartilage between the vertebræ. The most striking features of the spinal column may be illustrated by this simple apparatus.

38. How the Head and Spine are Joined together. The head rests upon the spinal column in a manner worthy of special notice. This consists in the peculiar structure of the first two cervical vertebræ, known as the axis and atlas. The atlas is named after the fabled giant who supported the earth on his shoulders. This vertebra consists of a ring of bone, having two cup-like sockets into which fit two bony projections arising on either side of the great opening (_foramen magnum_) in the occipital bone. The hinge joint thus formed allows the head to nod forward, while ligaments prevent it from moving too far.

On the upper surface of the axis, the second vertebra, is a peg or process, called the _odontoid process_ from its resemblance to a tooth. This peg forms a pivot upon which the head with the atlas turns. It is held in its place against the front inner surface of the atlas by a band of strong ligaments, which also prevents it from pressing on the delicate spinal cord. Thus, when we turn the head to the right or left, the skull and the atlas move together, both rotating on the odontoid process of the axis.

39. The Ribs and Sternum. The barrel-shaped framework of the chest is in part composed of long, slender, curved bones called ribs. There are twelve ribs on each side, which enclose and strengthen the chest; they somewhat resemble the hoops of a barrel. They are connected in pairs with the dorsal vertebræ behind.

The first seven pairs, counting from the neck, are called the _true_ ribs, and are joined by their own special cartilages directly to the breastbone. The five lower pairs, called the _false_ ribs, are not directly joined to the breastbone, but are connected, with the exception of the last two, with each other and with the last true ribs by cartilages. These elastic cartilages enable the chest to bear great blows with impunity. A blow on the sternum is distributed over fourteen elastic arches. The lowest two pairs of false ribs, are not joined even by cartilages, but are quite free in front, and for this reason are called _floating_ ribs.

The ribs are not horizontal, but slope downwards from the backbone, so that when raised or depressed by the strong intercostal muscles, the size of the chest is alternately increased or diminished. This movement of the ribs is of the utmost importance in breathing (Fig. 91).

The sternum, or breastbone, is a long, flat, narrow bone forming the middle front wall of the chest. It is connected with the ribs and with the collar bones. In shape it somewhat resembles an ancient dagger.

40. The Hip Bones. Four immovable bones are joined together so as to form at the lower extremity of the trunk a basin-like cavity called the pelvis. These four bones are the sacrum and the coccyx, which have been described, and the two hip bones.

Illustration: Fig. 17.—Thorax. (Anterior view.)

The hip bones are large, irregularly shaped bones, very firm and strong, and are sometimes called the haunch bones or _ossa innominata_ (nameless bones). They are united to the sacrum behind and joined to each other in front. On the outer side of each hip bone is a deep cup, or socket, called the _acetabulum_, resembling an ancient vinegar cup, into which fits the rounded head of the thigh bone. The bones of the pelvis are supported like a bridge on the legs as pillars, and they in turn contain the internal organs in the lower part of the trunk.

41. The Hyoid Bone. Under the lower jaw is a little horseshoe shaped bone called the hyoid bone, because it is shaped like the Greek letter upsilon (Υ). The root of the tongue is fastened to its bend, and the larynx is hung from it as from a hook. When the neck is in its natural position this bone can be plainly felt on a level with the lower jaw and about one inch and a half behind it. It serves to keep open the top of the larynx and for the attachment of the muscles, which move the tongue. (See Fig. 46.) The hyoid bone, like the knee-pan, is not connected with any other bone.

The Bones of the Upper Limbs.

42. The Upper Limbs. Each of the upper limbs consist of the upper arm, the forearm, and the hand. These bones are classified as follows:

Upper Arm: Scapula, or shoulder-blade, Clavicle, or collar bone, Humerus, or arm bone,

Forearm: Ulna, Radius,

Hand: 8 Carpal or wrist bones, 5 Metacarpal bones, 14 Phalanges, or finger bones,

making 32 bones in all.

43. The Upper Arm. The two bones of the shoulder, the scapula and the clavicle, serve in man to attach the arm to the trunk. The scapula, or shoulder-blade, is a flat, triangular bone, placed point downwards, and lying on the upper and back part of the chest, over the ribs. It consists of a broad, flat portion and a prominent ridge or _spine_. At its outer angle it has a shallow cup known as the _glenoid cavity_. Into this socket fits the rounded head of the humerus. The shoulder-blade is attached to the trunk chiefly by muscles, and is capable of extensive motion.

The clavicle, or collar bone, is a slender bone with a double curve like an italic _f_, and extends from the outer angle of the shoulder-blade to the top of the breastbone. It thus serves like the keystone of an arch to hold the shoulder-blade firmly in its place, but its chief use is to keep the shoulders wide apart, that the arm may enjoy a freer range of motion. This bone is often broken by falls upon the shoulder or arm.

The humerus is the strongest bone of the upper extremity. As already mentioned, its rounded head fits into the socket of the shoulder-blade, forming a ball-and-socket joint, which permits great freedom of motion. The shoulder joint resembles what mechanics call a universal joint, for there is no part of the body which cannot be touched by the hand.

Illustration: Fig. 18.—Left Scapula, or Shoulder-Blade.

When the shoulder is dislocated the head of the humerus has been forced out of its socket. The lower end of the bone is grooved to help form a hinge joint at the elbow with the bones of the forearm (Fig. 27).

44. The Forearm. The forearm contains two long bones, the ulna and the radius. The ulna, so called because it forms the elbow, is the longer and larger bone of the forearm, and is on the same side as the little finger. It is connected with the humerus by a hinge joint at the elbow. It is prevented from moving too far back by a hook-like projection called the _olecranon process_, which makes the sharp point of the elbow.

The radius is the shorter of the two bones of the forearm, and is on the same side as the thumb. Its slender, upper end articulates with the ulna and humerus; its lower end is enlarged and gives attachment in part to the bones of the wrist. This bone radiates or turns on the ulna, carrying the hand with it.

Experiment 10. Rest the forearm on a table, with the palm up (an attitude called supination). The radius is on the outer side and parallel with the ulna If now, without moving the elbow, we turn the hand (pronation), as if to pick up something from the table, the radius may be seen and felt crossing over the ulna, while the latter has not moved.

Illustration: Fig. 19.—Left Clavicle, or Collar Bone. (Anterior surface.)

45. The Hand. The hand is the executive or essential part of the upper limb. Without it the arm would be almost useless. It consists of 27 separate bones, and is divided into three parts, the wrist, the palm, and the fingers.

Illustration: Fig. 20.—Left Humerus. Fig. 21.—Left Radius and Ulna.

The carpus, or wrist, includes 8 short bones, arranged in two rows of four each, so as to form a broad support for the hand. These bones are closely packed, and tightly bound with ligaments which admit of ample flexibility. Thus the wrist is much less liable to be broken than if it were to consist of a single bone, while the elasticity from having the eight bones movable on each other, neutralizes, to a great extent, a shock caused by falling on the hands. Although each of the wrist bones has a very limited mobility in relation to its neighbors, their combination gives the hand that freedom of action upon the wrist, which is manifest in countless examples of the most accurate and delicate manipulation.

The metacarpal bones are the five long bones of the back of the hand. They are attached to the wrist and to the finger bones, and may be easily felt by pressing the fingers of one hand over the back of the other. The metacarpal bones of the fingers have little freedom of movement, while the thumb, unlike the others, is freely movable. We are thus enabled to bring the thumb in opposition to each of the fingers, a matter of the highest importance in manipulation. For this reason the loss of the thumb disables the hand far more than the loss of either of the fingers. This very significant opposition of the thumb to the fingers, furnishing the complete grasp by the hand, is characteristic of the human race, and is wanting in the hand of the ape, chimpanzee, and ourang-outang.

The phalanges, or finger bones, are the fourteen small bones arranged in three rows to form the fingers. Each finger has three bones; each thumb, two.

The large number of bones in the hand not only affords every variety of movement, but offers great resistance to blows or shocks. These bones are united by strong but flexible ligaments. The hand is thus given strength and flexibility, and enabled to accomplish the countless movements so necessary to our well-being.

In brief, the hand is a marvel of precise and adapted mechanism, capable not only of performing every variety of work and of expressing many emotions of the mind, but of executing its orders with inconceivable rapidity.

The Bones of the Lower Limbs.

46. The Lower Limbs. The general structure and number of the bones of the lower limbs bear a striking similarity to those of the upper limbs. Thus the leg, like the arm, is arranged in three parts, the thigh, the lower leg, and the foot. The thigh bone corresponds to the humerus; the tibia and fibula to the ulna and radius; the ankle to the wrist; and the metatarsus and the phalanges of the foot, to the metacarpus and the phalanges of the hand.

The bones of the lower limbs may be thus arranged:

Thigh: Femur, or thigh bone,

Lower Leg: Patella, or knee cap, Tibia, or shin bone, Fibula, or splint bone,

Foot: 7 Tarsal or ankle bones, 5 Metatarsal or instep bones, 14 Phalanges, or toes bones,

making 30 bones in all.

Illustration: Fig. 22.—Right Femur, or Thigh Bone.

47. The Thigh. The longest and strongest of all the bones is the femur, or thigh bone. Its upper end has a rounded head which fits into the _acetabulum_, or the deep cup-like cavity of the hip bone, forming a perfect ball-and-socket joint. When covered with cartilage, the ball fits so accurately into its socket that it may be retained by atmospheric pressure alone (sec. 50).

The shaft of the femur is strong, and is ridged and roughened in places for the attachment of the muscles. Its lower end is broad and irregularly shaped, having two prominences called _condyles_, separated by a groove, the whole fitted for forming a hinge joint with the bones of the lower leg and the knee-cap.

48. The Lower Leg. The lower leg, like the forearm, consists of two bones. The tibia, or shin bone, is the long three-sided bone forming the front of the leg. The sharp edge of the bone is easily felt just under the skin. It articulates with the lower end of the thigh bone, forming with it a hinge joint.

The fibula, the companion bone of the tibia, is the long, slender bone on the outer side of the leg. It is firmly fixed to the tibia at each end, and is commonly spoken of as the small bone of the leg. Its lower end forms the outer projection of the ankle. In front of the knee joint, embedded in a thick, strong tendon, is an irregularly disk-shaped bone, the patella, or knee-cap. It increases the leverage of important muscles, and protects the front of the knee joint, which is, from its position, much exposed to injury.

Illustration: Fig. 23.—Patella, or Knee-Cap.

49. The Foot. The bones of the foot, 26 in number, consist of the tarsal bones, the metatarsal, and the phalanges. The tarsal bones are the seven small, irregular bones which make up the ankle. These bones, like those of the wrist, are compactly arranged, and are held firmly in place by ligaments which allow a considerable amount of motion.

One of the ankle bones, the _os calcis_, projects prominently backwards, forming the heel. An extensive surface is thus afforded for the attachment of the strong tendon of the calf of the leg, called the tendon of Achilles. The large bone above the heel bone, the _astragalus_, articulates with the tibia, forming a hinge joint, and receives the weight of the body.

The metatarsal bones, corresponding to the metacarpals of the hand, are five in number, and form the lower instep.

The phalanges are the fourteen bones of the toes,—three in each except the great toe, which, like the thumb, has two. They resemble in number and plan the corresponding bones in the hand. The bones of the foot form a double arch,—an arch from before backwards, and an arch from side to side. The former is supported behind by the os calcis, and in front by the ends of the metatarsal bones. The weight of the body falls perpendicularly on the astragalus, which is the key-bone or crown of the arch. The bones of the foot are kept in place by powerful ligaments, combining great strength with elasticity.

Illustration: Fig. 24.—Right Tibia and Fibula (Anterior surface.)

Illustration: Fig. 25.—Bones of Right Foot. (Dorsal surface.)

The Joints.

50. Formation of Joints. The various bones of the skeleton are connected together at different parts of their surfaces by joints, or articulations. Many different kinds of joints have been described, but the same general plan obtains for nearly all. They vary according to the kind and the amount of motion.

The principal structures which unite in the formation of a joint are: bone, cartilage, synovial membrane, and ligaments. Bones make the chief element of all the joints, and their adjoining surfaces are shaped to meet the special demands of each joint (Fig. 27). The joint-end of bones is coated with a thin layer of tough, elastic cartilage. This is also used at the edge of joint-cavities, forming a ring to deepen them. The rounded heads of bones which move in them are thus more securely held in their sockets.

Besides these structures, the muscles also help to maintain the joint-surfaces in proper relation. Another essential to the action of the joints is the pressure of the outside air. This may be sufficient to keep the articular surfaces in contact even after all the muscles are removed. Thus the hip joint is so completely surrounded by ligaments as to be air-tight; and the union is very strong. But if the ligaments be pierced and air allowed to enter the joint, the union at once becomes much less close, and the head of the thigh bone falls away as far as the ligaments will allow it.

51. Synovial Membrane. A very delicate connective tissue, called the synovial membrane, lines the capsules of the joints, and covers the ligaments connected with them. It secretes the _synovia_, or joint oil, a thick and glairy fluid, like the white of a raw egg, which thoroughly lubricates the inner surfaces of the joints. Thus the friction and heat developed by movement are reduced, and every part of a joint is enabled to act smoothly.

52. Ligaments. The bones are fastened together, held in place, and their movements controlled, to a certain extent, by bands of various forms, called ligaments. These are composed mainly of bundles of white fibrous tissue placed parallel to, or closely interlaced with, one another, and present a shining, silvery aspect. They extend from one of the articulating bones to another, strongly supporting the joint, which they sometimes completely envelope with a kind of cap (Fig. 28). This prevents the bones from being easily dislocated. It is difficult, for instance, to separate the two bones in a shoulder or leg of mutton, they are so firmly held together by tough ligaments.

While ligaments are pliable and flexible, permitting free movement, they are also wonderfully strong and inextensible. A bone may be broken, or its end torn off, before its ligaments can be ruptured. The wrist end of the radius, for instance, is often torn off by force exerted on its ligaments without their rupture.

The ligaments are so numerous and various and are in some parts so interwoven with each other, that space does not allow even mention of those that are important. At the knee joint, for instance, there are no less than fifteen distinct ligaments.

53. Imperfect Joints. It is only perfect joints that are fully equipped with the structures just mentioned. Some joints lack one or more, and are therefore called imperfect joints. Such joints allow little or no motion and have no smooth cartilages at their edges. Thus, the bones of the skull are dovetailed by joints called sutures, which are immovable. The union between the vertebræ affords a good example of imperfect joints which are partially movable.

Illustration: Fig. 26.—Elastic Tissue from the Ligaments about Joints. (Highly magnified.)

54. Perfect Joints. There are various forms of perfect joints, according to the nature and amount of movement permitted. They an divided into hinge joints, ball-and-socket joints and pivot joints.

The hinge joints allow forward and backward movements like a hinge. These joints are the most numerous in the body, as the elbow, the ankle, and the knee joints.

In the ball-and-socket joints—a beautiful contrivance—the rounded head of one bone fits into a socket in the other, as the hip joint and shoulder joint. These joints permit free motion in almost every direction.

In the pivot joint a kind of peg in one bone fits into a notch in another. The best example of this is the joint between the first and second vertebræ (see sec. 38). The radius moves around on the ulna by means of a pivot joint. The radius, as well as the bones of the wrist and hand, turns around, thus enabling us to turn the palm of the hand upwards and downwards. In many joints the extent of motion amounts to only a slight gliding between the ends of the bones.

55. Uses of the Bones. The bones serve many important and useful purposes. The skeleton, a general framework, affords protection, support, and leverage to the bodily tissues. Thus, the bones of the skull and of the chest protect the brain, the lungs, and the heart; the bones of the legs support the weight of the body; and the long bones of the limbs are levers to which muscles are attached.

Owing to the various duties they have to perform, the bones are constructed in many different shapes. Some are broad and flat; others, long and cylindrical; and a large number very irregular in form. Each bone is not only different from all the others, but is also curiously adapted to its particular place and use.

Illustration: Fig. 27.—Showing how the Ends of the Bones are shaped to form the Elbow Joint. (The cut ends of a few ligaments are seen.)

Nothing could be more admirable than the mechanism by which each one of the bones is enabled to fulfill the manifold purposes for which it was designed. We have seen how the bones of the cranium are united by sutures in a manner the better to allow the delicate brain to grow, and to afford it protection from violence. The arched arrangement of the bones of the foot has several mechanical advantages, the most important being that it gives firmness and elasticity to the foot, which thus serves as a support for the weight of the body, and as the chief instrument of locomotion.

The complicated organ of hearing is protected by a winding series of minute apartments, in the rock-like portion of the temporal bone. The socket for the eye has a jutting ridge of bone all around it, to guard the organ of vision against injury. Grooves and canals, formed in hard bone, lodge and protect minute nerves and tiny blood-vessels. The surfaces of bones are often provided with grooves, sharp edges, and rough projections, for the origin and insertion of muscles.

Illustration: Fig. 28.—External Ligaments of the Knee.

56. The Bones in Infancy and Childhood. The bones of the infant, consisting almost wholly of cartilage, are not stiff and hard as in after life, but flexible and elastic. As the child grows, the bones become more solid and firmer from a gradually increased deposit of lime salts. In time they become capable of supporting the body and sustaining the action of the muscles. The reason is that well-developed bones would be of no use to a child that had not muscular strength to support its body. Again, the numerous falls and tumbles that the child sustains before it is able to walk, would result in broken bones almost every day of its life. As it is, young children meet with a great variety of falls without serious injury.

But this condition of things has its dangers. The fact that a child’s bones bend easily, also renders them liable to permanent change of shape. Thus, children often become bow-legged when allowed to walk too early. Moderate exercise, however, even in infancy, promotes the health of the bones as well as of the other tissues. Hence a child may be kept too long in its cradle, or wheeled about too much in a carriage, when the full use of its limbs would furnish proper exercise and enable it to walk earlier.

57. Positions at School. Great care must be exercised by teachers that children do not form the habit of taking injurious positions at school. The desks should not be too low, causing a forward stoop; or too high, throwing one shoulder up and giving a twist to the spine. If the seats are too low there will result an undue strain on the shoulder and the backbone; if too high, the feet have no proper support, the thighs may be bent by the weight of the feet and legs, and there is a prolonged strain on the hips and back. Curvature of the spine and round shoulders often result from long-continued positions at school in seats and at desks which are not adapted to the physical build of the occupant.

Illustration: Fig. 29.—Section of the Knee Joint. (Showing its internal structure)

A, tendon of the semi-membranosus muscle cut across; B, F, tendon of same muscle; C, internal condyle of femur; D, posterior crucial ligament; E, internal interarticular fibro cartilage; G, bursa under knee-cap; H, ligament of knee-cap; K, fatty mass under knee-cap; L, anterior crucial ligament cut across; P, patella, or knee-cap

A few simple rules should guide teachers and school officials in providing proper furniture for pupils. Seats should be regulated according to the size and age of the pupils, and frequent changes of seats should be made. At least three sizes of desks should be used in every schoolroom, and more in ungraded schools. The feet of each pupil should rest firmly on the floor, and the edge of the desk should be about one inch higher than the level of the elbows. A line dropped from the edge of the desk should strike the front edge of the seat. Sliding down into the seat, bending too much over the desk while writing and studying, sitting on one foot or resting on the small of the back, are all ungraceful and unhealthful positions, and are often taken by pupils old enough to know better. This topic is well worth the vigilance of every thoughtful teacher, especially of one in the lower grades.

58. The Bones in After Life. Popular impression attributes a less share of life, or a lower grade of vitality, to the bones than to any other part of the body. But really they have their own circulation and nutrition, and even nervous relations. Thus, bones are the seat of active vital processes, not only during childhood, but also in adult life, and in fact throughout life, except perhaps in extreme old age. The final knitting together of the ends of some of the bones with their shafts does not occur until somewhat late in life. For example, the upper end of the tibia and its shaft do not unite until the twenty-first year. The separate bones of the sacrum do not fully knit into one solid bone until the twenty-fifth year. Hence, the risk of subjecting the bones of young persons to undue violence from injudicious physical exercise as in rowing, baseball, football, and bicycle-riding.

The bones during life are constantly going through the process of absorption and reconstruction. They are easily modified in their growth. Thus the continued pressure of some morbid deposit, as a tumor or cancer, or an enlargement of an artery, may cause the absorption or distortion of bones as readily as of one of the softer tissues. The distortion resulting from tight lacing is a familiar illustration of the facility with which the bones may be modified by prolonged pressure.

Some savage races, not content with the natural shape of the head, take special methods to mould it by continued artificial pressure, so that it may conform in its distortion to the fashion of their tribe or race. This custom is one of the most ancient and widespread with which we are acquainted. In some cases the skull is flattened, as seen in certain Indian tribes on our Pacific coast, while with other tribes on the same coast it is compressed into a sort of conical appearance. In such cases the brain is compelled, of course, to accommodate itself to the change in the shape of the head; and this is done, it is said, without any serious result.

59. Sprains and Dislocations. A twist or strain of the ligaments and soft parts about a joint is known as a sprain, and may result from a great variety of accidents. When a person falls, the foot is frequently caught under him, and the twist comes upon the ligaments and tissues of the ankle. The ligaments cannot stretch, and so have to endure the wrench upon the joint. The result is a sprained ankle. Next to the ankle, a sprain of the wrist is most common. A person tries, by throwing out his hand, to save himself from a fall, and the weight of the body brings the strain upon the firmly fixed wrist. As a result of a sprain, the ligaments may be wrenched or torn, and even a piece of an adjacent bone may be torn off; the soft parts about the injured joint are bruised, and the neighboring muscles put to a severe stretch. A sprain may be a slight affair, needing only a brief rest, or it may be severe and painful enough to call for the most skillful treatment by a surgeon. Lack of proper care in severe sprains often results in permanent lameness.

A fall or a blow may bring such a sudden wrench or twist upon the ligaments as to force a bone out of place. This displacement is known as a dislocation. A child may trip or fall during play and put his elbow out of joint. A fall from horseback, a carriage, or a bicycle may result in a dislocation of the shoulder joint. In playing baseball a swift ball often knocks a finger out of joint. A dislocation must be reduced at once. Any delay or carelessness may make a serious and painful affair of it, as the torn and bruised parts rapidly swell and become extremely sensitive.

60. Broken Bones. The bones, especially those of the upper limbs, are often fractured or broken. The _simple_ fracture is the most common form, the bone being broken in a single place with no opening through the skin. When properly adjusted, the bone heals rapidly. Sometimes bones are crushed into a number of fragments; this is a _comminuted_ fracture. When, besides the break, there is an opening through the soft parts and surface of the body, we have a _compound_ fracture. This is a serious injury, and calls for the best surgical treatment.

A bone may be bent, or only partly broken, or split. This is called “a green-stick fracture,” from its resemblance to a half-broken green stick. This fracture is more common in the bones of children.

Fractures may be caused by direct violence, as when a bone is broken at a certain point by some powerful force, as a blow from a baseball bat or a fall from a horse. Again, a bone may be broken by indirect violence, as when a person being about to fall, throws out his hand to save himself. The force of the fall on the hand often breaks the wrist, by which is meant the fracture of the lower end of the radius, often known as the “silver-fork fracture.” This accident is common in winter from a fall or slip on the ice.

Sometimes bones are broken at a distance from the point of injury, as in a fracture of the ribs by violent compression of the chest; or fracture may occur from the vibration of a blow, as when a fall or blow upon the top of the head produces fracture of the bones at the base of the brain.[6]

61. Treatment for Broken Bones. When a bone is broken a surgeon is needed to set it, that is, to bring the broken parts into their natural position, and retain them by proper appliances. Nature throws out between and around the broken ends of bones a supply of repair material known as plastic lymph, which is changed to fibrous tissue, then to cartilage, and finally to bone. This material serves as a sort of cement to hold the fractured parts together. The excess of this at the point of union can be felt under the skin for some time after the bone is healed.

With old people a broken bone is often a serious matter, and may cripple them for life or prove fatal. A trifling fall, for instance, may cause a broken hip (popularly so called, though really a fracture of the neck of the femur), from the shock of which, and the subsequent pain and exhaustion, an aged person may die in a few weeks. In young people, however, the parts of a broken bone will knit together in three or four weeks after the fracture is reduced; while in adults, six or even more may be required for firm union. After a broken bone is strong enough to be used, it is fragile for some time; and great care must be taken, especially with children, that the injured parts may not be broken again before perfect union takes place.[7]

62. The Effect of Alcohol upon the Bones. While the growth of the bones occurs, of course, mainly during the earlier years of life, yet they do not attain their full maturity until about the twenty-fifth year; and it is stated that in persons devoted to intellectual pursuits, the skull grows even after that age. It is plainly necessary that during this period of bone growth the nutrition of the body should be of the best, that the bones may be built up from pure blood, and supplied with all the materials for a large and durable framework. Else the body will be feeble and stunted, and so through life fall short of its purpose.

If this bony foundation be then laid wrong, the defect can never be remedied. This condition is seen in young persons who have been underfed and overworked. But the use of alcoholic liquors produces a similar effect, hindering bone cell-growth and preventing full development.[8] The appetite is diminished, nutrition perverted and impaired, the stature stunted, and both bodily and mental powers are enfeebled.

63. Effect of Tobacco upon the Bones. Another narcotic, the destructive influence of which is wide and serious, is tobacco. Its pernicious influence, like that of alcohol, is peculiarly hurtful to the young, as the cell development during the years of growth is easily disturbed by noxious agents. The bone growth is by cells, and a powerful narcotic like tobacco retards cell-growth, and thus hinders the building up of the bodily frame. The formation of healthy bone demands good, nutritious blood, but if instead of this, the material furnished for the production of blood is poor in quality or loaded with poisonous narcotics, the body thus defrauded of its proper building material becomes undergrown and enfeebled.

Two unfavorable facts accompany this serious drawback: one is, that owing to the insidious nature of the smoky poison[9] (cigarettes are its worst form) the cause may often be unsuspected, and so go on, unchecked; and the other, that the progress of growth once interrupted, the gap can never be fully made up. Nature does her best to repair damages and to restore defects, but never goes backwards to remedy neglects.

Additional Experiments.

Experiment 11. Take a portion of the decalcified bone obtained from Experiment 4, and wash it thoroughly in water: in this it is insoluble. Place it in a solution of carbonate of soda and wash it again. Boil it in water, and from it gelatine will be obtained.

Experiment 12. Dissolve in hydrochloric acid a small piece of the powdered bone-ash obtained from Experiment 3. Bubbles of carbon dioxid are given off, indicating the presence of a carbonate. Dilute the solution; add an excess of ammonia, and we find a white precipitate of the phosphate of lime and of magnesia.

Experiment 13. Filter the solution in the preceding experiment, and to the filtrate add oxalate of ammonia. The result is a white precipitate of the oxalate of lime, showing there is lime present, but not as a phosphate.

Experiment 14. To the solution of mineral matters obtained from Experiment 3, add acetate of soda until free acetic acid is present, recognized by the smell (like dilute vinegar); then add oxalate of ammonia. The result will be a copious white precipitate of lime salts.

Experiment 15. _To show how the cancellous structure of bone is able to support a great deal of weight_. Have the market-man saw out a cubic inch from the cancellous tissue of a fresh beef bone and place it on a table with its principal layers upright. Balance a heavy book upon it, and then gradually place upon it various articles and note how many pounds it will support before giving way.

Experiment 16. Repeat the last experiment, using a cube of the decalcified bone obtained from Experiment 4.

Note. As the succeeding chapters are studied, additional experiments on bones and their relation to other parts of the body, will readily suggest themselves to the ingenious instructor or the thoughtful student. Such experiments may be utilized for review or other exercises.

Review Analysis: The Skeleton (206 bones).