Part 16

_Juv._ (♂, 53,193, Truckee River, Nevada, July 24, 1867; R. Ridgway: first plumage). Above plumbeous-black, tail more slaty. Every feather broadly bordered terminally with dull cinnamon; these crescentic bars becoming gradually broader posteriorly, narrower and more obsolete on the head above. Tail distinctly tipped with pale cinnamon, the inner webs of feathers with obsolete transverse spots of the same, these touching neither the edge nor the shaft; scarcely apparent indications of corresponding spots on outer webs. Region round the eye, and broad “mustache” across the cheeks, pure black, the latter more conspicuous than in the older stages, being cut off posteriorly by the extension of the cream-color of the neck nearly to the eye. A broad stripe of pale ochraceous running from above the ear-coverts back to the occiput, where the two of opposite sides nearly meet. Lower parts purplish cream-color, or rosy ochraceous-white, deepest posteriorly; jugulum, breast, sides, flanks, and tibiæ with longitudinal stripes of plumbeous-black, these broadest on flanks and abdomen, and somewhat sagittate on the tibiæ; lower tail-coverts with distant transverse bars. Lining of the wing like the sides, but the markings more transverse; inner web of longest primary with nine transverse purplish-ochre spots. Wing-formula, 2–1, 3. Wing, 12.50; tail, 7.00. Length, 16.50; expanse, 39.25. Weight, 1½ lbs. Basal half of bill pale bluish-white, cere rather darker; terminal half (rather abruptly) slate-color, the tip deepening into black; iris very dark vivid vandyke-brown; naked orbital space pale bluish-white, with a slight greenish tint; tarsi and toes lemon-yellow, with a slight green cast; claws jet-black.

HAB. Entire continent of America, and neighboring islands.

Localities: Guatemala (SCL. Ibis I, 219); Veragua (SALV. P. Z. S. 1867, 158); Sta. Cruz (NEWTON, Ibis, I, 63); Trinidad (TAYLOR, Ibis, 1864, 80); Bahamas (BRYANT, Pr. Bost. Soc. 1859, VII); Cuba (CAB. Journ. II, lxxxiii); (GUNDL. Repert. 1865, 225); Jamaica, (GOSSE, B. Jam. 16; MARCH, Pr. Ac. N. S. 1863, 304, et Mus. S. I.); Tierra del Fuego (SHARPE, Ann. & Mag. N. H.; “_F. cassini_, SHARPE”).

The young plumage above described corresponds exactly with that of young _peregrinus_ from Europe, a comparison of the specimen above described with one of the same age from Germany (54,064, Schlüter Col.) showing no differences that can be expressed. Many American specimens in this plumage (as 19,397, Fort Simpson) show a wash of whitish over the forehead and anterior part of the crown; having before us but the one specimen, we cannot say whether or not this is ever seen in the European bird. Specimens more advanced in season—perhaps in second year—are colored as follows: The black above is more brownish, the feathers margined with pale brown,—these margins broader, and approaching to white, on the upper tail-coverts; the tail shows the ochraceous bars only on inner webs. The supraoral stripe of the youngest plumage is also quite apparent.

A still younger one from the same locality (No. 37,397) has the upper plumage similar to the last, the pale edges to the feathers, however, more distinct; tail with conspicuous spots. White beneath clearer, and invading the dusky of the head above as far back as the middle of the crown; the supraoral stripe is distinct, scarcely interrupted across the nape.

In the adult plumage the principal variation is in the extent and disposition of the bars beneath. In most individuals they are regularly transverse only laterally and posteriorly, those on the belly being somewhat broken into more irregular cordate spots, though always transverse; in no American specimen, however, are they as continuously transverse as in a male (No. 18,804) from Europe, which, however, in this respect, we think, forms an exception to most European examples, at least to those in the Smithsonian Collection. All variations in the form, thickness, and continuity of the markings below, and in the distinctness of the bars above, are individual.

Very old males (as 49,790, Fort Yukon; 27,188, Moose Factory (type of Elliott’s figure of _F. peregrinus_, in Birds of America); and 42,997, Spanishtown, Jamaica) lack almost entirely the reddish tinge beneath, and have the lateral and posterior portions strongly tinged with blue; the latter feature is especially noticeable in the specimen from Jamaica, in which also the bars are almost utterly wanting medially. Immature birds from this island also lack to a great degree the ochraceous tinge, leaving the whitish everywhere purer.

A female adult European bird differs from the average of North American examples in the conspicuous longitudinal streaks on the jugulum; but in a male these are hardly more distinct than in 13,077, ♀, Liberty Co., Georgia; 11,983, “United States”; 35,456, Peel’s River; 35,449, ♀, and 35,445, ♀, Fort Yukon, Alaska; 35,452, La Pierre’s Hous., H. B. Ter.; 35,459 ♂, Fort Anderson; and 28,099 ♀, Hartford, Conn. In none of these, however, are they so numerous and conspicuous as in a European female from the Schlüter Collection, which, however, differs in these respects only from North American specimens.

A somewhat melanistic individual (in second year? 32,735, Chicago, Ill.; Robert Kennicott) differs as follows: Above continuously pure black; upper tail-coverts and longer scapulars bordered terminally with rusty-whitish. Tail distinctly tipped with white; the inner webs of feathers with eight elliptical transverse bars of pale ochraceous, and indications of corresponding spots of the same on outer webs, forming as many inconspicuous bands. Beneath ochraceous-white; the neck, breast, and abdomen thickly marked with broad longitudinal stripes of clear black,—those on the jugulum cuneate, and on the breast and abdomen broadly sagittate; the tibiæ with numerous cordate spots, and sides marked more transversely; lower tail-coverts with narrow distant transverse bars. On the chin and throat only, the whitish is immaculate, on the other portions being somewhat exceeded in amount by the black. Inner web of longest primary with seven transverse elliptical bars of cream-color. Wing, 12.20; tail, 9.40.

Whether the North American and European Peregrine Falcons are or are not distinct has been a question undecided up to the present day; almost every ornithologist having his own peculiar views upon the relationship of the different forms which have been from time to time characterized. The most favorably received opinion, however, seems to be that there are two species on the American continent, and that one of these, the northern one, is identical with the European bird. Both these views I hold to be entirely erroneous; for after examining and comparing critically a series of more than one hundred specimens of these birds, from every portion of America (except eastern South America), including nearly all the West India Islands, as well as numbers of localities throughout continental North and South America, I find that, with the exception of the melanistic littoral race of the northwest coast (var. _pealei_), they all fall under one race, which, though itself exceedingly variable, yet possesses characters whereby it may always be distinguished from the Peregrine of all portions of the Old World.

There is such a great amount of variability, in size, colors, and markings, that the _F. nigriceps_, Cassin, must be entirely ignored as being based upon specimens not distinguishable in any respect from typical _anatum_. Judging from the characters assigned to the _F. cassini_ by its describer (who evidently had a very small series of American specimens at his command), the latter name must also most probably fall into the list of synonymes of _anatum_.

Slight as are the characters which separate the Peregrines of the New and Old World, i.e. the immaculate jugulum of the former and the streaked one of the latter, they are yet sufficiently constant to warrant their separation as geographical races of one species; along with which the _F. melanogenys_, Gould (Australia), _F. minor_, Bonap. (South Africa), _F. orientalis_, Gmel. (E. Asia), and _F. calidus_, Lath. (Southern India and East Indies), must also rank as simple geographical races of the same species. Whether the _F. calidus_ is tenable, I am unable to state, for I have not seen it; but the others appear to be all sufficiently differentiated. The _F. radama_, Verreaux (Gray’s Hand List, p. 19, No. 170), Mr. Gurney writes me, is the young female of var. _minor_. Whether the _F. peregrinator_, Sundevall (Gray’s Hand List, No. 169), is another of the regional forms of _F. communis_, or a distinct species, I am not able at present to say, not having specimens accessible to me for examination.

Mr. Cassin’s type of “_nigriceps_” (13,856, ♂, July), from Chile, is before me, and upon comparison with adult males from Arctic America presents no tangible differences beyond its smaller size; the wing is a little more than half an inch, and the middle toe less than the eighth of an inch, shorter than in the smallest of the North American series,—a discrepancy slight indeed, and of little value as the sole specific character; the plumage being almost precisely similar to that of the specimen selected for the type of the description at the head of this article. In order to show the little consequence to be attached to the small size of the individual just mentioned, I would state that there is before me a young bird, received from the National Museum of Chile, and obtained in the vicinity of Santiago, which is precisely similar in plumage to the Nevada specimen described, and in size is even considerably larger, though it is but just to say that it is a female; the wing measures 13.25, instead of 12.50, and the middle toe, 2.00, instead of 1.85. No. 37,336, Tres Marias Islands, Western Mexico,—a young male in second year,—has the wing just the same length as in the smallest North American example, while in plumage it is precisely similar to 26,785, of the same age, from Jamaica. No. 4,367, from Puget’s Sound, Washington Territory,—also a young male,—has the wing of the same length as in the largest northern specimen, while the plumage is as usual.

Two adult females from Connecticut (Nos. 28,099 and 32,507, Talcott Mt.) are remarkable for their very deep colors, in which they differ from all other North American examples which I have seen, and answer in every particular to the description of _F. cassini_, Sharpe, above cited. The upper surface is plumbeous-black, becoming deep black anteriorly, the head without a single light feather in the black portions; the plumbeous bars are distinct only on the rump, upper tail-coverts, and tail, and are just perceptible on the secondaries. The lower parts are of a very deep reddish-ochraceous, deepest on the breast and abdomen, where it approaches a cinnamon tint,—the markings, however, as in other examples. They measure, wing, 14.75; tail, 7.50; culmen, 1.05–1.15; tarsus, 2.00; middle toe, 2.30. They were obtained from the nest, and kept in confinement three years, when they were sacrificed to science. The unusual size of the bill of these specimens (see measurements) is undoubtedly due to the influence of confinement, or the result of a modified mode of feeding. The specimens were presented by Dr. S. S. Moses, of Hartford.

An adult male (No. 8,501) from Shoal-water Bay, Washington Territory, is exactly of the size of the male described. In this specimen there is not the slightest creamy tinge beneath, while the blue tinge on the lower parts laterally and posteriorly is very strong. No. 52,818, an adult female from Mazatlan, Western Mexico, has the wing three quarters of an inch shorter than in the largest of four northern females, and of the same length as in the smallest; there is nothing unusual about its plumage, except that the bars beneath are sparse, and the ochraceous tinge quite deep. No. 27,057, Fort Good Hope, H. B. T., is, however, exactly similar, in these respects, and the wing is but half an inch longer. In No. 47,588, ♂, from the Farallones Islands, near San Francisco, California, the wing is the same length as in the average of northern and eastern specimens, while the streaks on the jugulum are nearly as conspicuous as in a male from Europe.

In conclusion, I would say that the sole distinguishing character between the Peregrines from America and those from Europe, that can be relied on, appears to be found in the markings on the breast in the adult plumage; in all the specimens and figures of var. _communis_ that I have seen, the breast has the longitudinal dashes very conspicuous; while, as a general rule, in _anatum_ these markings are entirely absent, though sometimes present, and occasionally nearly as distinct as in European examples. Therefore, if this conspicuous streaking of the breast is found in all European specimens, the American bird is entitled to separation as a variety; but if the breast is ever immaculate in European examples, then _anatum_ must sink into a pure synonyme of _communis_. The var. _melanogenys_ is distinguished from both _communis_ and _anatum_ by the black auriculars, or by a greater amount of black on the side of the neck, and by more numerous and narrower bars on the under surface. In the former feature examples of _anatum_ from the southern extremity of South America approach quite closely to the Australian form, as might be expected from the relative geographical position of the two regions. The var. _minor_ is merely the smaller intertropical race of the Old World, perhaps better characterized than the tropical American form named _F. nigriceps_ by Cassin, the characters of which are so unimportant, and withal so inconstant, as to forbid our recognizing it as a race of the same rank with the others.

LIST OF SPECIMENS EXAMINED.

National Museum, 45; Boston Society, 4; Philadelphia Academy, 22; Museum Comp. Zoöl. 5; New York Museum, 3; G. N. Lawrence, 6; R. Ridgway, 3. Total, 88.

_Measurements._

+----+-----------+-----------+---------+---------+-----------+----------+ |Sex.| Wing. | Tail. | Culmen. | Tarsus. |Middle Toe.|Specimens.| +----+-----------+-----------+---------+---------+-----------+----------+ | ♂ |11.30–13.00| 6.00–7.50| .75–0.80|1.60–1.90| 1.78–2.05| 29 | +----+-----------+-----------+---------+---------+-----------+----------+ | ♀ |13.00–14.75| 7.30–9.00| .85–1.00|1.95–2.10| 1.95–2.20| 28 | +----+-----------+-----------+---------+---------+-----------+----------+

Var. pealei, RIDGWAY.

BLACK PEREGRINE FALCON.

_? ? Accipiter falco niger_, BRISS. Orn. I, 337. _? ? Falco niger_, GMEL. S. N. 1789, 270. _Falco polyagrus_, CASS. B. Cal. & Tex. pl. xvi (dark figure).

SP. CHAR. In colors almost exactly similar to _F. gyrfalco_, var. _labradora_. Above continuous dark vandyke-brown, approaching brownish-black on the head, which is variegated only on the gular region, and inclining to grayish-brown on the tail; the whole surface entirely free from spots or markings of any kind. Beneath similar in color to the upper parts, but the feathers edged with whitish, this rather predominating on the throat; flanks and tibiæ with roundish white spots; lower tail-coverts with broad transverse bars of white. Lining of the wing with feathers narrowly tipped with white; inner webs of primaries with narrow, transverse elliptical spots of cream-color; inner webs of tail-feathers with badly defined, irregular, similar spots, or else with these wanting, the whole web being plain dusky-brown.

No. 12,022 (♀, Oregon; T. R. Peale). Wing, 15.00; tail, 8.50; culmen, .95; tarsus, 2.10; middle toe, 2.15. (Figured by Cassin as _F. polyagrus_, in Birds of California and Texas, pl. xvi.)

No. 45,814 (♀, Sitka, Alaska, May, 1866; F. Bischoff). Wing, 14.90; tail, 8.50; tarsus, 2.10; middle toe, 2.20. The two similar in color, but in the latter the white streaks on the lower parts a little broader, and the middle of the auriculars slightly streaked.

HAB. Northwest coast of North America, from Oregon to Sitka.

This curious race of _Falco communis_ is a good illustration of the climatic peculiarity of the northwest coast region, to which I have often referred before; the same melanistic tendency being apparent in birds of other species from the same region, as an example of which I may mention the Black Merlin (_Falco æsalon_, var. _suckleyi_), which is a perfect miniature of the present bird.

HABITS. The Great-footed Hawk of North America is very closely allied to the well-known Peregrine Falcon of Europe, and so closely resembles it that by many writers, even at the present day, it is regarded as identical with it. Without doubt, the habits of the two races are very nearly the same, though the peculiarities of the North American bird are not so well known as are those of the European. In its distribution it is somewhat erratic, for the most part confined to the rocky sea-coast, the river-banks, and the high ground of the northeastern parts of America. It is known to breed in a few isolated rocky crags in various parts of the country, even as far to the south as Pennsylvania, and it occurs probably both as migrant and resident in several of the West India Islands, in Central and in South America. A single specimen was taken by Dr. Woodhouse in the Creek country of the Indian Territory. Two individuals are reported by Von Pelzeln as having been taken in Brazil. The Newtons met with it in St. Croix. Mr. Gosse found it in Jamaica, and Dr. Gundlach gives it as a bird of Cuba. Jardine states it to be a bird of Bermuda, and also that it has been taken in the Straits of Magellan. A single specimen was taken at Dueñas, Guatemala, in February, by Mr. Salvin.

On the Pacific coast this Falcon has been traced as far south as the limit of the land. Dr. Cooper met with only two pairs, in March, 1854, frequenting a high wooded cliff at Shoal-water Bay. Dr. Suckley procured a single specimen from Steilacoom. Dr. Cooper states that the habits of these corresponded with those described for the _F. anatum_ and _F. peregrinus_, and that, like these Falcons, it is a terror to all land animals weaker than itself. It is said to breed on the rocky cliffs of the Pacific.

An individual of this bird was taken by Colonel Grayson at the Tres Marias Islands. When shot, it was endeavoring to capture a Sparrow-hawk, indicating its indifference as to the game it pursues. He adds that this bird attacks with vigor everything it sees, from the size of a Mallard Duck down, and is the terror of all small birds. Its range must be very great, as it often ventures far out to sea. On his passage from Mazatlan to San Francisco, in 1858, on the bark Carlota, one of these Falcons came on board more than a hundred miles off the coast of Lower California, and took up its quarters on the main-top yard, where it remained two days, during which time it captured several Dusky Petrels. It would dart headlong upon these unsuspecting birds, seldom missing its aim. It would then return to its resting-place and partly devour its prize. At other times it dropped its victims into the sea in wanton sport. Finally, as if tired of this kind of game, it made several wide circles around the ship, ascended to a considerable height, and departed in the direction of the Mexican shore.

This Falcon is found along the Atlantic coast from Maine to the extreme northern portion, breeding on the high rocky cliffs of Grand Menan and in various favorable situations thence northward. A few breed on Mount Tom, near the Connecticut River in Massachusetts, on Talcott Mountain in Connecticut, in Pennsylvania, and near Harper’s Ferry, in Maryland.

Mr. Boardman has several times taken their eggs from the cliffs of Grand Menan, where they breed in April, or early in May. In one instance he found the nest in close proximity to that of a pair of Ravens, the two families being apparently on terms of amity or mutual tolerance.

For several years two or more pairs of these birds have been known to breed regularly on Mount Tom, near Northampton. The nests were placed on the edges of precipitous rocks very early in the spring, the young having been fully grown by the last of June. Their young and their eggs have been taken year after year, yet at the last accounts they still continued to nest in that locality. Dr. W. Wood has also found this species breeding on Talcott Mountain, near Hartford. Four young were found, nearly fledged, June 1. In one instance four eggs were taken from a nest on Mount Tom, by Mr. C. W. Bennett, as early as April 19. This was in 1864. Several times since he has taken their eggs from the same eyrie, though the Hawks have at times deserted it and sought other retreats. In one year a pair was twice robbed, and, as is supposed, made a third nest, and had unfledged young as late as August. Mr. Allen states that these Hawks repair to Mount Tom very early in the spring, and carefully watch and defend their eyrie, manifesting even more alarm at this early period, when it is approached, than they evince later, when it contains eggs or young. Mr. Bennett speaks of the nest as a mere apology for one.

This Hawk formerly nested on a high cliff near the house of Professor S. S. Haldeman, Columbia, Penn., who several times procured young birds which had fallen from the nest. The birds remained about this cliff ten or eleven months of the year, only disappearing during the coldest weather, and returning with the first favorable change. They bred early in spring, the young leaving the nest perhaps in May. Professor Haldeman was of the opinion that but a single pair remained, the young disappearing in the course of the season.

Sir John Richardson, in his Arctic expedition in 1845, while descending the Mackenzie River, latitude 65°, noticed what he presumed to be a nest of this species, placed on the cliff of a sandstone rock. This Falcon was rare on that river.

Mr. MacFarlane found this species not uncommon on the banks of Lockhart and Anderson Rivers, in the Arctic regions. In one instance he mentions finding a nest on a cliff thirty feet from the ground. There were four eggs lying on a ledge of the shale of which the cliff was composed. Both parents were present, and kept up a continued screaming, though at too great a distance for him to shoot either. He adds that this bird is by no means scarce on Lockhart River, and he was informed that it also nests along the ramparts and other steep banks of the Upper Anderson, though he has not been able to learn that it has been found north of Fort Anderson. In another instance the nest was on a ledge of clayey mud,—the eggs, in fact, lying on the bare ground, and nothing resembling a nest to be seen. A third nest was found on a ledge of crumbling shale, along the banks of the Anderson River, near the outlet of the Lockhart. This Hawk, he remarks, so far as he was able to observe, constructs no nest whatever. At least, on the Anderson River, where he found it tolerably abundant, it was found to invariably lay its eggs on a ledge of rock or shale, without making use of any accessory lining or protection, always availing itself of the most inaccessible ledges. He was of the opinion that they do not breed to the northward of the 68th parallel. They were also to be found nesting in occasional pairs along the lime and sandstone banks of the Mackenzie, where early in August, for several successive years, he noticed the young of the season fully fledged, though still attended by the parent birds.

In subsequent notes, Mr. MacFarlane repeats his observations that this species constructs no nest, merely laying its eggs on a ledge of shale or other rock. Both parents were invariably seen about the spot. In some instances the eggs found were much larger than in others.